Saprininae, Blanchard, 1845

Penati, Fabio & Vienna, Pierpaolo, 2006, An updated catalogue of the Histeridae (Insecta: Coleoptera) of the Arabian Peninsula, with biogeographical remarks, Zootaxa 1157 (1), pp. 1-74 : 34-40

Subfam. Saprininae

28. Saprinus (Saprinus) aegyptiacus aegyptiacus Marseul ( Fig. 12 )

Distribution: Egypt, Tunisia, Algeria, Morocco, Mauritania, Iran, Turkmenistan ( Gomy 1996a), Saudi Arabia ( Kryzhanovskij 1979).

Chorology: Saharo­Turano­Sindian.

Literature. Saudi Arabia: Wadi Tumair [= Wadi Tumayr, 23°52’00”N 46°58’60”E], 20.II.1976, 1 ex. ( Kryzhanovskij 1979) .

29. Saprinus (Saprinus) bicolor (Olivier) ( Fig. 12 )

Distribution: Senegal, St. Helen Is., Mauritania, Guinea­Bissau, Ethiopia, Somalia, Angola, Republic of South Africa, Saudi Arabia, Yemen ( Gomy 2004).

Chorology: Afrotropical (widespread).

Literature. Saudi Arabia: Asir Mts., Wadi Shuqub [20°43’00”N 41°19’60”E] , 7.IV.1983, 4 ex. ( Mazur 1994); Fifa near Gizan [= Faifa, 17°08’38”N 43°07’09”E] , 27–31.III.1983, 2 ex. ( Mazur 1994); Riyadh [= Riyad, 24°38’27”N 46°46’22”E] , 6.II.1980, 2 ex. ( Mazur 1994). Yemen: Sana’a [15°21’17”N 44°12’24”E], ca. 2400 m , XI.1939, in house, 1 ex. ( Müller 1954); Wadi Sharez [a few km east of Hajjah, 15°41’36”N 43°36’11”E] , II.1985, cow dung, 1 ♂ ( Gomy 2004).

Material. Yemen: Sana’a NW env., Wadi Dharhan , 15°26’43”N 44°67’34”E, 2240 m, 15–17.XI.2003, 1 ex. ( CHTL) .

30. Saprinus (Saprinus) caerulescens caerulescens (Hoffmann) ( Fig. 12 )

Synonym: Saprinus semipunctatus (Fabricius) : Alonso­Zarazaga & Yélamos 1994: 179.

Distribution: “Barbaria”, Mediterranean Subregion, Azores, Cape Verde, Caucasus, Kyrgyzstan, Turkmenistan, Afghanistan, Iran, Saudi Arabia, introduced to Peru.

Chorology: Turano­Mediterranean.

Literature. Saudi Arabia: Jeddah [= Jiddah, 21°31’01”N 39°13’09”E], 23.II.1976, 1 ex. ( Kryzhanovskij 1979, as S. semipunctatus ); Fifa near Gizan [= Faifa, 17°08’38”N 43°07’09”E], 23–31.III.1983, 32 ex. ( Mazur 1994, as S. semipunctatus ); Asir Mts., Wadi Shuqub [20°43’00”N 41°19’60”E], 7.IV.1983, 4 ex. ( Mazur 1994, as S. semipunctatus ); Usfan [21°49’41”N 39°18’05”E], 29.IX.1983, 1 ex. ( Mazur 1994, as S. semipunctatus ); Khulais valley [= Khulays, 22°09’14”N 39°19’06”E], 80 km NE of Jeddah, 1997–1998 ( Al­Ghamdi 1999, as S. semipunctatus ).

31. Saprinus (Saprinus) calatravensis Fuente ( Fig. 13 )

Distribution: Spain, Mediterranean Subregion, Central Asia, Afghanistan, Saudi Arabia, Oman.

Chorology: Centralasiatic­Mediterranean.

Literature. Saudi Arabia: Wadi Ilyab, 20°07’N 40°57’E, 160 m, 11.XI.1984, 2 ex. ( Mazur 1994); Bani Musaygirah [= Bani Musayjirah, 20°21’00”N 44°31’60”N], 850 m, 16: IX.1981, 1 ex. ( Mazur 1994). Oman: SE of Muscat, Wadi Mayh, 23°27’N

58°35’E, 200 m, 11–12.IV.1985, 2 ex. ( Mazur 1994).

32. Saprinus (Saprinus) chalcites (Illiger) ( Fig. 13 )

Distribution: Portugal, Mediterranean Subregion, Africa, Arabian Peninsula, Central Asia, India, Burma, Australia.

Chorology: Subcosmopolitan.

Literature. Saudi Arabia: Hawi [Hawi saddle, 20°34’00”N 42°37’60”E], N of Prisha [=Qal`at Bishah] and ca. 260 km NE of Al Qunfidha, 25.V.1936, 3 ex. ( Müller 1954); Mahdatha [21°52’23”N 40°41’44”E], 96 km NE of Mecca [= Makkah], I.1945, 1 ex. ( Müller 1954); 9 km Gizan/Taif road [= Jizan surroundings, 16°53’31”N 42°32’59”E], 29.XII.1978, 1 ex. ( Mazur 1994); Asir Mts., Wadi Shuqub [20°43’00”N 41°19’60”E], 7.IV.1983, 1 ex. ( Mazur 1994); Dawasir [20°38’60”N 45°08’60”E], 24.III.1983, 5 ex. ( Mazur 1994); Sharoura [= Sharurah, 17°34’31”N 47°09’42”E], 29.III.1979, 1 ex. ( Mazur 1994); Gizan on the Red Sea [= Jizan, 16°53’31”N 42°32’59”E], 25– 26.III.1983, 2 ex. ( Mazur 1994); Shoiba [= Shu`aybah, 20°41’60”N 39°25’60”E], 22.XI.1983, 3 ex. ( Mazur 1994); Wadi Ilyab, 20°07’N 40°57’E, 160 m, 11.XI.1984, 3 ex. ( Mazur 1994); Khulais valley [= Khulays, 22°09’14”N 39°19’06”E], 80 km NE of Jeddah, 1997–1998 ( Al­Ghamdi 1999); Riyadh surroundings [= Riyad, 24°38’27”N 46°46’22”E], 12.X.1975, 1 ♀ ( Kryzhanovskij 1979, as S.? chalcites ). Kuwait: Umm al Heman, 28°57’N 48°12’E, 12 m, 9.III.1988, 1 ex. ( Mazur 1994). Yemen: Sana’a [15°21’17”N 44°12’24”E], within walls of Bir­el­Azab, ca. 2400 m, I.1938, 1 ex. ( Müller 1954); ibidem, 10–15.X.1937, 1 ex. ( Müller 1954); Mukeiras [= Mukayris, 13°56’38”N 45°40’25”N], 137 km NE of Aden, 2130 m, 29.XII.1939 – 6.I.1940, 2 ex. ( Müller 1954); Wadi Dhabab [= Wadi Dhabah, 13°55’00”N 44°15’00”E], II.1984, 1 ♂ ( Gomy 2004); Marib [15°25’16”N 45°20’01”E], III.1985, 7 ex. ( Gomy 2004); Wadi Sharez [a few km east of Hajjah, 15°41’36”N 43°36’11”E], 24.III.1985, cow dung, 4 ♂ and 3 ♀ ( Gomy 2004). Oman: SE of Muscat, Wadi Mayh, 23°27’N 58°35’E, 200 m, 11–12.IV.1985, 16 ex. ( Mazur 1994); Mintirib, 22°26’N 58°48’E, 268 m, without date, 1 ex. ( Mazur 1994).

Material. Saudi Arabia: “Saudi Arabia”, sine data ( CHTY) ; Daharan, Al Jubail [= Dhahran, al Jubayl, 27°02’50”N 49°39’41”E], 4.IV.2001, 5 ex. ( MSNTC) . Kuwait: Kuwait City , 29°20’N 48°00’E, 11–20.IV.1996, 1 ♀ ( CMNH) . Yemen: near Shiakh Othman ( Wadi ) [=Ash Shaykh`Uthman, 12°52’53”N 44°59’42”E], IV.1985, 12 ♀ and 3 ♂ ( MNHUB) ; ibidem, V.1985, 3 ♂ ( MNHUB) ; ibidem, VI.1985, 2 ♀ and 2 ♂ ( MNHUB) ; ibidem, VII.1985, 1 ♀ ( MNHUB) ; ibidem, III.1987, 2 ♀ and 5 ♂ ( MNHUB) ; ibidem, VI.1987, 6 ♀ and 5 ♂ ( MNHUB) ; ibidem, VII.1985, 1 ♀ ( MNHUB) ; Sana’a NW env., Wadi Dharhan , 15°26’43”N 44°57’34”E, 2240 m, 15– 17.XI.2003, 9 ex. ( CHTL) . Oman: Dhofar, Qeiroon Heiritti env. [17°17’N 54°05’E], 5 km from the village, 850 m, 24.VIII.2000, 2 ex. ( MSNTC) ; Dhofar, Marbat , 26.VIII.2000, 1 ex. ( MSNTC) ; Dhofar, Rd. Taqah to Madnat al Haq , 2 km after jct., 5.IX.2002, 2 ex. ( MSNTC) ; Dhofar, Rd. 45– Rakhyut env., 16°45’07”N 53°24’59”E, 30 m, 30.IX.2001, 2 ex. ( MSNTC) ; Dhofar, Rakhyut env. N, 16°45’00”N 53°24’67”E,, 12.IX.2002, 1 ex. ( MSNTC); Dhofar, Rd. 47 Arift env., 16°50’31”N 53°22’85”E, 1040 m, 28.VIII.2000, 2 ex. ( MSNTC) ; Dhofar, Al Mughsayl env., 16°53’01”N 53°46’47’E, 30 m, 2.X.2001, 1 ex. ( MSNTC) ; Dhofar, Rd. 47– Al Mughsayl env., 16°53’71”N 53°49’01”E, 30 m, 2.X.2001, 1 ♂ ( MSNTC); Dhofar, Wadi Ashawq, 16°53’83”N 53°46’31”E, 40 m, 23.IX.2001, 2 ex. ( MSNTC) ; Dhofar, Ad Dahariz env., Robat Palace , 17°01’56”N 54°08’99”E, 20 m, 3.X.2001, 1 ex. ( MSNTC); Dhofar, Khor Rori, 17°02’27”N 54°26’16”E, 40 m, 24.VIII.2000, 20 ex. ( MSNTC); ibidem, 4.IX.2000, 1 ex. ( MSNTC); ibidem, 12.IX.2000, 1 ex. ( MSNTC); ibidem, 1.IX.2002, 1 ex. ( MSNTC); Dhofar, Rd. Marbat to Sadh, 17°03’06”N 54°50’97”E, 135 m, 7.IX.2000, 10 ex. ( MSNTC) ; Dhofar, Rd. Salalah to Darbat , 17°03’14”N 54°25’58”E, 20 m, 9.IX.2000, 2 ex. ( MSNTC) ; Dhofar, Wadi Darbat , 17°04’50”N 54°25’80”E, 24.VIII.2000, 7 ex. ( MSNTC); Dhofar, Rd. Marbat to Sadh, km 30.1 E Marbat, 17°04’67”N 54°53’39”E, 170 m, 6.X.2001, 6 ex. ( MSNTC) ; Dhofar, Darbat Pool , 17°05’70”N 54°26’80”E, 220 m, 22.IX.2001, 1 ex. ( MSNTC); Dhofar, Rd. to Jabal Samhan, Tawi Atayr env., 17°06’81”N 54°38’21”E, 950 m, 25.IX.2001, 4 ex. ( MSNTC) ; Dhofar, Ain Sahanawt env., 17°07’40”N 54°09’44”E, 24.IX.2001, 1 ex. ( MSNTC) ; Dhofar, Teetaam env., 17°07’50”N 53°55’64”E, 1000 m, 2.IX.2000, 1 ex. ( MSNTC); Dhofar, Ain Razat 17°07’75”N 54°14’24”E, 110 m, 21.IX.2001, 6 ex. ( MSNTC) ; ibidem, 22.IX.2001, 1 ex. ( MSNTC) ; ibidem, 2.X.2001, 2 ex. ( MSNTC) ; Dhofar, Rd. Razat to Madnat, As San , 17°08’91”N 54°13’21”E, 420 m, 21.IX.2001, 1 ex. ( MSNTC) ; Dhofar, Teetaam env., 17°13’37”N 53°54’22”E, 600– 780 m, 27.VIII.2000, 2 ex. ( MSNTC) ; Dhofar, Jibjat env. E, 17°14’15”N 54°31’90”E, 980 m, 1.IX.2000, 1 ex. ( MSNTC); Dhofar, Hajahif env., 17°14’53”N 54°01’67”E, 900 m, 1.X.2001, 2 ex. ( MSNTC) ; Dhofar, Rd. to Uyun, Uyun env., sink, 17°14’71”N 53°57’43”E, 800 m, 26.IX.2001, 6 ex. ( MSNTC); Dhofar, Hajaif env., 17°14’72”N 53°57’38”E, 770 m, 2.IX.2000, 2 ex. ( MSNTC) ; Dhofar, Jibjat env., 17°15’61”N 54°29’84”E, 950 m, 29.VIII.2000, 1 ex. ( MSNTC) ; Dhofar, Rd. 31 to Thumrayt , 17°17’58”N 54°05’21”E, 760 m, 29.VIII.2000, 1 ex. ( MSNTC) ; Dhofar, Rd. 31, Queiroon env. N, 17°17’59”N 54°05’19”E, 750 m, 1.X.2001, 4 ex. ( MSNTC) ; ibidem, 7.X.2001, 1 ex. ( MSNTC) ; Dhofar, Wadi Dokha , 17°19’70”N 54°04’80”E, 650 m, 8.IX.2002, 3 ex. ( MSNTC) ; Dhofar, Al Mughsayl env., 16°53’01”N 53°46’47”E, 30 m, 1.III.2004, 2 ex. ( MSNTC) ; ibidem, 16°52’53”N 53°43’51”E, 8.III.2004, 1 ex. ( MSNTC) ; Dhofar, Wadi Darbat , 17°05’45”N 54°26’59”E, 200 m, 3.III.2004, 1 ex. ( MSNTC) ; Dhofar, Wadi Dowkha , 17°20’32”N 54°04’16”E, 5–9.III.2004, UV light trap, 6 ex. ( MSNTC) ; Dhofar, Ain Sahanawt env., 17°08’N 54°10’E, 6.III.2004, 1 ex. ( MSNTC) ; Dhofar, Ain Garziz env., 17°06’29”N 54°04’35”E, 110 m, 14.III.2004, 1 ♂ ( MSNTC) ; Dhofar, Ain Razat env., 17°07’N 54°14’E, 16.III.2004, 1 ex. ( MSNTC) .

33. Saprinus (Saprinus) confalonierii G. Müller ( Fig. 14 )

Distribution: Tunisia, Libya, Egypt, Algeria, Mauritania ( Gomy 1996a), Saudi Arabia, Oman ( Mazur 1997).

Chorology: Saharo­Sahelo­Arabian.

Literature. Saudi Arabia: Nejd, Mudhnib district [25°45’00”N 44°15’00”E], 2.IV.1946, probably predacious on egg­masses of locusts, 4 ex. ( Müller 1954, as S. gilvicornis confalonierii Müller ; one specimen is deposited in Collection Müller); Hail [27°31’25”N 41°42’18”E], IV–VII.1944, 2 ex. ( Müller 1954, as S. gilvicornis confalonierii Müller ); Salim [?], 20.IX.1976, 1 ex. ( Mazur 1994); al Hunavy [= Al Hunayy, 24°57’50”N 48°45’23”E], 22.X.1978, 1 ex. ( Mazur 1994); Sharoura [= Sharurah, 17°34’31”N 47°09’42”E], 27.III.1979, 2 ex. ( Mazur 1994); Qusuriyah [23°45’00”N 44°34’60”E], 13–14.V.1980, 1 ex. ( Mazur 1994); Sah al Rimth [= Saja Um Al­Rimth Protected Area, about 23°00’N 43°00’E], 750 m, 24–25.IV.1981, 2 ex. ( Mazur 1994); Adama [19°41’N 42°04’E], 22.IX.1978, 1 ex. ( Mazur 1994); Ghazwar [26°43’60”N 41°21’00”E], 1010 m, 29.IV.1981, 1 ex. ( Mazur 1994); al Uraiq [?], 750 m, 2–3.XI.1986, 1 ex. ( Mazur 1994; reported coordinates “ 29°54’N 28°17’E ” are wrong!). Oman: Wadi Twasinat [= Wadi Tusinat], 17°53’N 52°57’E, 416 m, 12– 16.V.1982, 13 ex. ( Mazur 1994); Yalooni, 19°57’N 57°06’E, 2–4.X.1984, 1 ex. ( Mazur 1994); Jiddat al Harasis, Kuria Muria, al Qibuyah, 17°30’N 56°20’E, 10–190 m, 3–7.VII.1983, 1 ex. ( Mazur 1994); Qarhat Mu’ammar, Camp, 21°31’N 59°18’E, 130 m, 24.I.1986, 1 ex. ( Mazur 1994).

Material. Saudi Arabia: Daharan , Al Jubail [= Dhahran, al Jubayl, 27°02’50”N 49°39’41”E], 12.IX.1999, 3 ex. ( CHFP) ; ibidem, 29.III.2001, 1 ex. ( MSNTC) . Oman: 15 km NNE Fasab (N), 18°46’20”N 53°08’90”E, 290 m, 29– 30.01.1998, on lower slopes of high dunes in empty quarter, 1 ♂ and 1 ex. ( CHPK) ; idem, 4 ex. ( CHRP) ; 15 km NW Shigag , 19°37’N 54°04’E, 190 m, 30.XI–1.XII.1997, dune with some bushes, to light, 2 ex. ( CHFP) .

34. Saprinus (Saprinus) cruciatus (Fabricius) subsp. flavipennis Péringuey ( Fig. 14 )

Distribution: Republic of South Africa: Capland, South Africa, Zambia, Malawi, Kenia, Yemen ( Gomy 2005).

Chorology: Afrotropical (spread in E and S Africa).

Literature. Yemen: Al Bara [= Al Barah, 14°18’28”N 44°08’17”E], 29.III.1976, 1 ♀ ( Gomy 2005) .

35. Saprinus (Saprinus) cupreus Erichson ( Fig. 13 )

Distribution: Republic of South Africa: Capland, Cape Verde, Togo, Cameroon, Sudan, Ethiopia, Somalia, Rwanda, Namibia, Yemen (Aden), Burma, Vietnam, St. Helen Is., Ascension, Australia ( Gomy 2001).

Chorology: Subcosmopolitan.

Literature. Yemen: Aden [=`Adan, 12°46’46”N 45°02’12”E] ( Mazur 1976) .

Figures

FIGURE 12. Distribution of the Histeridae in the Arabian Peninsula. Saprinus (s. str.) aegyptiacus, S. (s. str.) bicolor, S. (s. str.)?buqueti, S. (s. str.) caerulescens.

FIGURE 13. Distribution of the Histeridae in the Arabian Peninsula. Saprinus (s. str.) calatravensis, S. (s. str.) chalcites, S. (s. str.) cupreus.

FIGURE 14. Distribution of the Histeridae in the Arabian Peninsula. Saprinus (s. str.) confalonierii, S. (s. str.) cruciatus flavipennis, S. (s. str.) flexuosofasciatus, S. (s. str.) gilvicornis.