Ventrilia imbricata (Hörnes, 1854)

Harzhauser, Mathias & Landau, Bernard, 2012, A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472, Zootaxa 3472, pp. 1-71 : 61

Ventrilia imbricata (Hörnes, 1854)

Figs 10I 1–I 3

Cancellaria imbricata Hörnes, 1854: 327 , pl. 35, figs 16a–c.

Cancellaria imbricata M. Hörnes. — Hilber 1879: 30.

Cancellaria (g. Trigonostoma ) imbricata M. Hoernes — Hoernes & Auinger 1890: 276.

Cancellaria imbricata Hörnes —Pereira da Costa 1867: 208, pl. 25, figs 6–7.

Cancellaria (Trigonostoma) imbricata Hörnes — Sieber 1936: 83.

Cancellaria (Trigonostoma) exgeslini Sacco — Csepreghy-Meznerics 1950: 58, pl. 3, fig. 16 [non Trigonostoma exgeslini ( Sacco, 1894) ].

Trigonostoma (Trigonostoma) cf. imbricatum (Hörnes, 1856 [sic])— Davoli 1982: 32, pl. 3, figs 1–3.

Trigonostoma (Trigonostoma) imbricatum (Hörnes, 1856) [sic]— Harzhauser 2002: 111, pl. 9, figs 3–5.

Trigonostoma imbricatum (Hörnes, 1856) — Bałuk 2006: 214, pl. 16, figs 3–4.

Trigonostoma (Ventrilia) imbricatum (Hörnes, 1854) — Landau et al. 2006: 68 (partim, figs. exclus).

Trigonostoma imbricatum (Hörnes, 1856) — Brunetti et al. 2008: 54, figs 2A–E.

Ventrilia imbricata (Hörnes, 1854) — Landau et al. 2011: 30, pl. 15, fig. 13.

non Trigonostoma aff. imbricatum (Hörnes, 1854) — Brunetti 2000: 29, figs 7–9 [= Scalptia etrusca Brunetti, Della Bella, Forli

& Vecchi, 2008]. non Trigonostoma (Ventrilia) imbricatum (Hörnes, 1854) — Landau et al. 2006: 68 (partim), pl. 4, figs 7–8 [= Scalptia etrusca

Brunetti, Della Bella, Forli & Vecchi, 2008].

Type material. NHM Vienna ( Austria); see below for details .

Studied material. 3 spec. Grund ( Austria) NHMW 1855 /0002/0017; 14 spec. Enzesfeld ( Austria) NHMW 1846 /0037/0287; 3 spec. Pöls ( Austria) NHMW 1861 /0001/0263; 23 spec. Steinebrunn ( Austria) NHMW A385 .

Illustrated material. Figs 10I 1–I 3: syntype illustrated in Hörnes (1854, pl. 35, figs 16a–c), Enzesfeld ( Austria), height: 29.3 mm, width: 19.4 mm, NHMW 1846 /0037/0287 [note that the dimensions given in Hörnes 1854 are incorrect] .

Discussion. This species was discussed in detail by Landau et al. (2011).

Distribution. This species appears during the late Early Miocene in the Paratethys Sea where it becomes a widespread species during the Middle Miocene. During the Late Serravallian–Early Tortonian it occurs also in the Proto –Mediterranean Sea, documented from the Turkish Karaman Basin (Landau in prep.) and is known from several Tortonian localities in Italy ( Sacco, 1894; Davoli, 1982; Brunetti et al., 2008). At that time it spreads even to the North East Atlantic (Pereira da Costa, 1867) where it persists up the Early Pliocene ( Landau et al., 2011).

Paratethys —Karpatian: Korneuburg Basin ( Austria: Kleinebersdorf); Badenian: North Alpine Foreland Basin ( Austria: Grund); Vienna Basin ( Czech Republic: Mikulov; Austria: Enzesfeld, Steinebrunn, Grinzing, Gainfarn); Eisenstadt–Sopron Basin ( Austria: Forchtenau); Styrian Vasin ( Austria: Pöls); Transylvanian Basin: Romania (Lăpugiu de Sus). North East Atlantic —Tortonian: Portugal (Cacela Basin); Early Pliocene: Spain (Guadalquivir Basin). Proto –Mediterranean Sea —Late Serravallian–Early Tortonian: Turkey (Karaman Basin); Tortonian: Italy (Po Basin).