Coenypha ditissima ( Nicolet, 1849 )

Machado, Miguel, Previato, Thales, Grismado, Cristian J. & Teixeira, Renato, 2023, Taxonomic review of the Andean crab spiders genus Coenypha Simon, 1895 (Thomisidae: Stephanopinae), Zootaxa 5306 (3), pp. 301-330 : 312-317

Coenypha ditissima ( Nicolet, 1849)

Figs 6A–F , 7A–F , 14B

Thomisus ditissimus Nicolet, 1849: 394 , pl. 3, fig. 9. Holotype female from Valdivia, Chile, deposited in MNHN (4176), examined. Ramírez 1989: 11.

Thomisus spissus Nicolet, 1849: 395 . Holotype female from Chile, deposited in MNHN (4178), examined. Ramírez 1989: 11. New Synonymy.

Thomisus variabilis Nicolet, 1849: 396 . Holotype male (together two immatures) from Chile, deposited in MNHN (4180), examined. Roewer 1955 (declared nomen dubium, rejected here); Ramírez 1989: 11. New Synonymy.

Stephanopis ditissima (Nicolet) : Keyserling 1880: 175, pl. 3, fig. 96; Simon 1887: E8, pl. 2, fig. 1; Mello-Leit„o 1951: 332, fig. 7; Ramírez 2014: 223, figs 151E–F, 172E; Machado et al. 2017: 454, fig. S12C; Machado and Teixeira 2021: 296 (transferred to Coenypha ).

Stephanopis spissa (Nicolet) : Simon 1895: 1054.

Stephanopis maulliniana Mello-Leit „o, 1951: 334. Holotype female from Maullín , Llanquihue, Chile, deposited in MNRJ, examined. New Synonymy.

Misumenoides nicoleti Roewer, 1951: 448 . Replacement name for Thomisus cinereus Nicolet, 1849 due to homonymy with Thomisus cinereus C. L. Koch, 1837 . Synonymized by Lehtinen & Marusik (2008: 186) with Misumenops temibilis ( Holmberg, 1876) , here rejected. New Synonymy.

Note: In the case of T. ditissimus , there was a single spider in the vial MNHN 4176, therefore we can consider this specimen the type by monotypy as stated in article 73.1.2 of the ICZN (2022).Two species described by Nicolet in 1849 ( T. cinereus and T. variabilis ) have been part of a confusing taxonomic history that involved several species of the distantly related genera Misumenoides F. O. Pickard-Cambridge, 1900 and Misumenops F. O. Pickard-Cambridge, 1900 , originated from taxonomic decisions made without the examination of the type specimens (see Mello-Leit„o 1939; Roewer 1955 and especially, the discussion of Lehtinen & Marusik 2008). As a result of this, the Stephanopine Thomisus cinereus Nicolet, 1849 appeared listed among the synonyms of Misumenops temibilis ( Holmberg, 1876) , a species belonging in the tribe Misumenini . Although the type specimens of these species have been found (the vial MNHN 4182 is T. cinereus Nicolet, 1849 , and MNHN 4180 is T. variabilis ), as reported by Ramírez (1989), those were unfortunately overlooked by Lehtinen and Marusik (2008). Considering the abovementioned and according to article 75.8 of the ICZN (2022), we do not recognize the neotype of T. cinereus (= T. nicoleti ) proposed by Lehtinen & Marusik (2008) and therefore, their synonymy with Misumenoides temibilis . Moreover, we examined all Nicolet’s types listed above in synonymy of Coenypha ditissima and conclude that there are no significant differences between them, hence they should be considered synonyms. Finally, the priority of T. ditissimus over T. cinereus (= T. nicoleti ), T. spissus , and T. variabilis (ICZN 2.4.2.1 and 2.4.2.2.) is given once Nicolet (1849) mention this species prior the other three in the literature.

Other material examined. ARGENTINA: Neuquén: 1♁ and 1♀, Lanín , 40°6’19.00”S, 71°40’11.00”W, January 1985, M. Ramírez ( MACN-Ar 18651 ) ; 1j, Lago Lácar , 40°10’37.79”S, 71°29’38.80”W, January 1954, N. Kormilev ( MACN-Ar 4164 ) ; 1j, Parque Nacional Nahuel Huapi , 40°53’27.87”S, 71°34’34.11”W, 30 January 1985, M. Ramírez ( MACN-Ar 18663 ) ; 1♁, Nahuel Huapi , 40°55’36.13”S, 71°29’34.65”W, 1945, D. Hiebermann ( MACN-Ar 18669 ) ; 1♀, Nahuel Huapi ( Puerto Blest ), 41°1’59.98”S, 71°49’0.05”W, 07–20 January 2000, L. Lopardo & A. Quaglino ( MACN-Ar 18650 ) . Río Negro: 2j, Bariloche , 41°8’44.93”S, 71°17’56.85”W, February 1954, M.E. Galliano ( MACN-Ar 5404 ) . Chubut: 1♀, Lago Puelo , 42°5’49.20”S, 71°36’44.20”W, 10–11 December 2021, M. Pacheco, L. Piacentini & E. Soto ( MACN-Ar 42752 ) ; 1♀, same locality, date and collectors of the previous vial ( MACN-Ar 42751 ) ; 1j, Parque Nacional Los Alerces , 42°58’26.85”S, 71°38’37.90”W, February 1985, M. Ramírez ( MACN-Ar 18657 ) . Tierra del Fuego: 1♀, Isla de losEstados, 54°47’59.46”S, 64°25’36.47”W, April 1935, J.B. Daguerre-Cancelles ( MACN-Ar 18652 ) ; 2♀ and 7j, Ushuaia , 54°48’19.44”S, 68°19’27.14”W, 16 February 1949, B. Aguirre ( MACN-Ar 2796 ) ; 1♀, same locality of the previous vial, 23 January 1960, A. Bachmann ( MACN-Ar 18658 ) ; 1♀, Parque Nacional Tierra del Fuego, margen E del Lago Roca , 15 km W deUshuaia, 54°49’32.92”S, 68°33’46.80”W, 04 December 2012, M. Ramírez, C. Grismado, A. Ojanguren & E.M. Soto ( MACN 29783 ) ; 1j, same locality of the previous vial, February 1963, Maury ( MACN-Ar 18666 ) . CHILE: Coquimbo: 3j, Choapa , 31°48’14.70”S, 71°0’8.03”W, 13 November 1987, Maurj ( MACN-Ar 18668 ) . Valparaíso: 1♀ and 2j, Zapallar , 32°33’13.70”S, 71°27’29.29”W, 27 November, Ross & Michelbacher ( CAS 9071265 ) ; 2♁ and 2j, same locality, date and collectors of the previous vial ( CAS 9071264 ) ; 2j, Quintero , 32°46’20.32”S, 71°31’59.88”W, 12 December 1980, L. Peña ( AMNH) ; 1♀, Caracoles , 32°47’30.18”S, 70° 3’38.57”W, August 1943, Rosa ( CAS 9072308 ) ; 1♀, Quillota ( Parque Nacional La Campana ), 32°57’40.40”S, 71° 3’34.00”W, 18 February 2005, M. Ramírez & F. Labarque ( MACN-Ar 10848 ) ; 6j, El Melón , 33° 3’55.11”S, 71°35’25.08”W, 03 November 1981, L. Peña ( AMNH) ; 1j, Tunquén , 33°16’25.45”S, 71°38’49.10”W, 14 October 1982, M. Pino ( MHNS 630 ) . Santiago: 6♀ and 4j, Quilicura , 33°21’27.63”S, 70°43’45.38”W, May 1979, L. Peña ( AMNH) ; 1♀, Las Condes , 33°23’56.33”S, 70°33’26.32”W, January 1983, no collector given ( MHNS 739 ) ; 5♀ and 3j, Cordoba Coast , 33°26’56.00”S, 70°40’9.35”W, 15–20 February 1979, L. Peña ( AMNH) ; 1♀, Malleco , 33°30’37.01”S, 70°40’16.68”W, November 1979, L. Peña ( AMNH) ; 15j, Cuesta La Dormida , 33°31’16.74”S, 70°47’42.40”W, 13–18 November 1982, L. Peña ( AMNH) ; 2♀,15j, El Canelo , 33°34’34.36”S, 70°27’0.00”W, 1980, L. Peña ( AMNH) ; 1♀, Guayacán , 33°35’60.00”S, 70°22’0.12”W, January 1984, P. Goloboff ( MACN-Ar 18704 ) ; 1♀, Pirque , 33°40’45.64”S, 70°34’59.19”W, 05 October 1982, L. Peña ( AMNH) ; 1♀ and 1j, Rio Maipo Región , 33°42’6.78”S, 70°20’8.44”W, November 1984, G. Arriagada ( MHNS 892 ) ; 3♀, Melipilla (La Viluma), 33°46’54.04”S, 71°4’44.55”W, 14 May 1980, L. Peña ( AMNH) . Maule: 1♁, Talca (Provincia de Linares) , 35°50’47.07”S, 71°35’58.61”W, 11–20 November 1964, L. Peña ( MCZ) . Bío-Bío: 1♁, Ñuble ( Las Trancas ), 36°37’9.12”S, 72° 4’55.24”W, 17 February 1983, L. Peña ( AMNH) ; 1♀, Ñuble ( Fundo Los Robles ), 36°42’16.20”S, 71°36’9.60”W, 15 February 2005, M. Ramírez & F. Labarque ( MACN-Ar 10847 ) ; 1♀, Concepción ( Chome ), 36°42’52.73”S, 73° 8’21.39”W, 07 December 1995, T. Cekalovic ( AMNH) ; 1♀ and 1j, 50 Km east of San Carlos , 36°43’21.27”S, 71°45’44.09”W, 26 December 1950, Ross & Michelbacher ( CAS 9072307 ) ; 1j, Concepción ( Bosque Ramuntcho ), 36°45’10.41”S, 73°11’8.79”W, 14–16 October 1961, A.F. Archer ( AMNH) ; 1♀, Concepción ( Hualpén ), 36°47’12.03”S, 73°6’35.83”W, 11 January 1989, M. Ramírez ( MACN-Ar 18655 ) ; 2♁, Concepción ( Periquillo ), 36°49’12.49”S, 73°2’39.80”W, 21 December 1996, T. Cekalovic ( AMNH) ; 1♀ and 1♁, same locality and collector of the previous vial, 16 March 1997 ( AMNH) ; 1♀, same locality and collector of the previous vial, 20 January 1996 ( AMNH) ; 1♀, same locality and collector of the previous vial, 22 March 1997 ( AMNH) ; 4♁ and 2j, same locality and collector of the previous vial, 29 December 1996 ( AMNH) 2j, Camino Chome Ramuntcho , 36°49’8.18”S, 73°3’1.15”W, 08 November 1996, T. Cekalovic ( AMNH) ; 1♀, Ñuble ( Las Comadres ), 36°53’14.22”S, 71°34’12.75”W, 05–09 February 1983, L. Peña ( AMNH) ; 1j, El Manzano , 36°54’58.31”S, 72°54’49.50”W, 23 December 1996, T. Cekalovic ( AMNH) ; 1♀, El Abanico , 37°20’27.52”S, 71°31’57.53”W, 30 December 1950, Ross & Michelbacher ( CAS 9072309 ) ; 1♁ and 1j, Ancon , 37°48’21.86”S, 72°42’13.70”W, 28 November 1950, E. Zapudo ( CAS 9071267 ) . Araucanía: 1♀, Parque Nacional Nahuelbuta , 37°49’24.60”S, 72°58’57.00”W, 07–25 December 2002, M. Thayer & A. Newton ( FMNH 2857856 ) ; 2j, Parquenco , 38°9’48.82”S, 72°31’13.74”W, 06 January 1951, Ross & Michelbacher ( CAS 9071271 ) ; 11j, Malalcahuello , 38°28’19.37”S, 71°36’21.67”W, 13–31 December 1982, A. Newton & M. Thayer ( AMNH) ; 9♁, 9♀ and 6j, Cautín ( Chacamo ), 38°34’56.21”S, 73°3’11.75”W, 17–23 February 1981, L. Peña ( AMNH) ; 1♁, Cautín ( Temuco ), 38°44’9.25”S, 72°35’25.35”W, January 1989, M. Ramírez ( MACN-Ar 18656 ) ; 1♀, Villarica , 39°16’55.24”S, 72°13’50.81”W, 03 March 1965, H. Levi ( MCZ 133890 ) ; 2♁, 1♀ and 1j, same locality, date and collector of the previous vial ( MCZ 133891 ) ; 1♀, Northeast of Villarica , 39°16’55.25”S, 72°13’50.81”W, 16-31 December 1964, L. Peña ( MCZ) . Los Ríos: 2♁ and 3♀, Valdivia (Sender Los Tapuales as Renoval Canelos ), 39°49’10.51”S, 73°14’42.76”W, 09January2002, T.Cekalovic ( FMNH71617 ) ; 4♁and3j, Las Lajas , 39°49’10.51”S, 73°14’42.76”W, 09–13 January 1990, L. Peña ( AMNH) ; 1♀ and 1j, Valdivia , 39°49’6.53”S, 73°14’37.94”W, 13 January 1951, Ross & Michelbacher ( CAS 9071266 ) ; 4♀ and 11j, same locality of the previous vial, December 1982, E. Krahmer ( MHNS 695 ) ; 1♁, 3♀ and 3j, same locality and collector of the previous vial, 1984 ( MHNS 841 ) ; 1♁, 11♀ and 1j, same locality and collector of the previous vial, 1983 ( MHNS 804 ) ; 1♁ and 1♀, Corral , 39°53’18.87”S, 73°25’53.76”W, 16 January 1989, M. Ramírez ( MACN-Ar 18654 ) . Los Lagos: 1♀, Osorno ( Pucatrihue ), 40°32’6.87”S, 73°42’31.82”W, February 1967, L. Peña ( MCZ 133410 ) ; 1♀, same locality and collector of the previous vial, 25–31 January 1978 ( AMNH) ; 3♁, 1♀ and 9j, Osorno ( Puyehue ), 40°34’34.28”S, 73°6’53.81”W, 24 January 1951, Ross & Michelbacher ( CAS 9072306 ) ; 3♁, 3♀ and 4j, same locality and collector of the previous vial, 25 January 1951 ( CAS 9071270 ) ; 2♀ and 2j, Osorno (La Picada), 40°34’48.75”S, 73°10’34.90”W, 15-20 January 1980, L. Peña ( AMNH) ; 1♁ and 1♀, Osorno ( Petrohue ), 40°34’52.18”S, 73°9’19.58”W, 13 January 1980 L. Peña ( AMNH) ; 1♁, 2♀ and 2j, Osorno ( Salto Pilmaiquén ), 40°39’11.36”S, 72°39’36.24”W, 26 January 1951, Ross & Michelbacher ( CAS 9071268 ) ; 1j, Osorno ( Anticura ), 40°39’12.71”S, 72°15’44.90”W, November 1982, no collector ( MHNS 709 ) ; 6j, Aguas Calientes ( Parque Nacional Puyehue ), 40°43’16.56”S, 72°19’3.87”W, 19-26 December 1982, A. Newton & M. Thayer ( AMNH) ; 4j, Aguas Calientes ( Parque Nacional Puyehue ), 40°43’43.88”S, 72°18’43.16”W, 13–17 December 1998, M. Ramírez, L. Compagnucci, C. Grismado & L. Lopardo ( MACN-Ar 18662 ) ; 1♀, Aguas Calientes ( Parque Nacional Puyehue ), 40°43’43.88”S, 72°18’43.17”W, 02–05 January 1982, L. Peña ( AMNH) ; 1♀, Llanquihue (Entre Lagos), 40°58’36.61”S, 72°52’55.06”W, 20 January 1969, L. Peña ( MCZ 133667 ) ; 1♀, Llanquihue ( Ensenada ), 41°12’25.50”S, 72°32’19.60”W, 18 March 1965, H. Levi ( MCZ 133893 ) ; 1♁, Llanquihue ( Correntoso ), 41°15’38.27”S, 73°0’28.23”W, January 1969 L. Peña ( MCZ) ; 1♁, 1♀ and 3j, Puerto Varas , 41°19’0.12”S, 72°58’59.88”W, 16 January 1951, Ross & Michelbacher ( CAS 9072305 ) ; 1♁, Llanquihue ( Chamisa ), 41°21’38.30”S, 72°31’13.74”W, 13 December 1968, L. Peña ( MCZ 133413 ) ; 2♁, 3♀ and 6j, Llanquihue ( Los Muermos ), 41°23’57.87”S, 73°27’53.96”W, 19 January 1951, E.I. Schlinger & E.S. Ross ( CAS 9071269 ) ; 5♁, 13♀ and 10j, same locality and date of the previous vial, Ross & Michelbacher ( CAS 9071272–9071273 ) ; 3♁ and 1j, Llanquihue ( Correntoso ), 41°25’13.82”S, 72°40’11.31”W, December 1968, L. Peña ( MCZ 133407 ) ; 1♁ and 2j, same locality ad collector of the previous vial, 20–25 January 1980 ( AMNH) ; 1♁, Llanquihue ( Puerto Montt ), 41°28’8.10”S, 72°56’28.09”W, February 1983, G. Arriagada ( MHNS 715 ) ; 1♁, Llanquihue (X Región), 41°34’59.99”S, 72°40’60.00”W, 13–15 January 1990, L. Peña ( AMNH) ; 1♁, Lepihué , 41°35’17.85”S, 73°33’52.64”W, 21 January 1951, Ross & Michelbacher ( CAS 9071263 ) ; 1♁ and 1♀, Isla Grande de Chiloé ( Guabún ), 41°49’34.75”S, 74° 1’57.21”W, 13–15 January 1980, L. Peña ( AMNH) ; 1♀, Isla Grande do Chiloé (Puente Rio Pudeto ), 41°52’40.12”S, 73°50’9.60”W, 17 February 1996, T. Cekalovic ( AMNH) ; 1♀, Isla Grande de Chiloé ( Duhatao ), 41°58’50.74”S, 74° 2’25.97”W, 16–21 January 1981, L. Peña ( AMNH) ; 1♁, Puntra , 42°7’7.39”S, 73°48’39.88”W, 21–23 December 1981, L. Peña ( AMNH) ; 1♁, Isla Grande de Chiloé ( Butalcura ), 42°13’59.88”S, 73°44’60.00”W, 21 February 1997, T. Cekalovic ( AMNH) ; 14♁, 15♀ and 10j, Isla Grande de Chiloé ( Dalcahue ), 42°22’29.25”S, 73°39’6.35”W, 01 February 1981, L. Peña ( AMNH) ; 2♀, Isla Grande de Chiloé ( Dalcahue ), 42°22’39.47”S, 73°39’6.78”W, same collector of the previous vial, 01–04 April 1941 ( MCZ 133412 ) ; 1♀, Isla Grande de Chiloé ( Cole Cole ), 42°25’22.39”S, 74°4’58.62”W, 08–11 February 1991, M. Ramírez ( MACN-Ar 18648 ) ; 2♀ and 3j, Isla Grande de Chiloé ( Gicao ), 42°27’22.52”S, 73°52’47.37”W, February 1981, L. Peña ( AMNH) ; 1♀ and 1j, Isla Grande de Chiloé ( Cerros de Cucao ), 42°28’60.00”S, 74°3’46.00”W, 01 March 1981, L. Peña ( AMNH) ; 2♀, Isla Quinchao ( Hullar Alto ), 42°28’9.00”S, 73°30’53.00”W, 15 January 2002, T. Cekalovic ( FMNH 71620 ) ; 4♁, 3♀ and 1j, same locality and collector of the previous vial, 19 February 1997 ( AMNH) ; 2♀, same locality and collector of the previous vial, 16 February 1996 ( AMNH) ; 2♁, Isla Grande de Chiloé ( Chepu ), 42°31’24.34”S, 73°52’47.36”W, 30 January 1981, L. Peña ( AMNH) ; 1♀ and 1j, Isla Grande de Chiloé ( Puente La Caldera ), 42°37’26.28”S, 73°55’35.66”W, 18 February 1997, T. Cekalovic ( AMNH) ; 2♀, same locality and collector of the previous vial, 15 February 1996 ( AMNH) ; 1♁ and 1♀, Isla Grande de Chiloé , 42°37’26.29”S, 73°55’35.66”W, 04 February 2001, T. Cekalovic ( FMNH 71619 ) , 1♀, 10 January 2002 ( FMNH 71630 ) ; 11♀, 19 February 1996 ( AMNH) , 1♁ and 2♀, 14 February 1996 ( AMNH) , 2♁ and 4♀, 17 February 1997 ( AMNH) , 1♀, 19 February 1997 ( AMNH) , 1♁ and 1♀, 22 February 1997 ( AMNH) , 2♀, 15-19 March 1981, L. Peña ( AMNH) , 1♀ and 2j, 22 February 1997, T. Cekalovic ( AMNH) ; 1♁, Isla Grande de Chiloé ( Piruquihue ), 42°37’26.29”S, 73°55’35.65”W, 10 February 1993, T. Cekalovic ( AMNH) ; 1♀, Isla Grande de Chiloé ( Cucao ), 42°37’59.88”S, 74°7’0.12”W, 01 February 1991, M. Ramírez ( MACN-Ar 18649 ) ; 1♀ and 1j, Isla Lemuy ( Puerto Haro ), 42°37’9.00”S, 73°38’57.00”W, 20February1996, T.Cekalovic ( AMNH) ; 1♀, Isla Grande de Chiloé ( Puente Trainel ), 42°40’30.57”S, 73°47’57.64”W, 18 February 1997, T. Cekalovic ( AMNH) ; 3♁ and 3j, Isla Grande de Chiloé ( Terao ), 42°42’40.07”S, 73°39’16.15”W, 18–21 February 1990, L. Peña ( AMNH) ; 4♀, Isla Grande de Chiloé ( San Antonio de Chadmo ), 42°54’15.73”S, 73°38’17.02”W, 08 February 2001, T. Cekalovic ( FMNH 61793 ) ; 2j, Palena ( Chaitén ), 42°54’41.51”S, 72°42’56.30”W, 04 December 1981, N. Platnick & R.T. Schuh ( AMNH) . Aysén: 2♀, 46°23’54.41”S, 72° 0’1.76”W, 17 February 1991, M. Ramírez ( MACN-Ar 18647 ) ; 2♀ and 10j, Aisén del General Carlos Ibanez del Campo, 46°24’43.55”S, 72°14’33.98”W, 27 January 1990, L. Peña ( AMNH) ; 1♀, Parque Nacional Los Glaciares , 50°19’50.00”S, 73°14’3.00”W, 17 January 1972, E. Hernandez ( MACN-Ar 18665 ) ; 2♀, Magallanes (Puerto Natales, Torres del Paine ), 51°43’40.76”S, 72°30’5.84”W, 12 December 2009, L. Almeida, H. Wood & C. Griswold ( IBSP 155000 ) .

Diagnosis. Females of C. ditissima resemble those of C. foliacea sp. nov. by the short prosoma covered by hyaline setae ( Fig. 6B ) and opisthosoma posteriorly enlarged, bearing two pairs of blunt and short projections disposed dorsolaterally, almost like dorsal bumps on the opisthosoma ( Fig. 6A ) (discrete and flattened in C. foliacea sp. nov., Fig. 10A ). Moreover, C. ditissima can be distinguished from C. foliacea sp. nov. by the presence of stout femoral apophysis on legs I and II (like in C. edwardsi ), epigynal plate with anteriorly narrow MSept and long copulatory duct ( Figs 6C, 6D ) coiling on itself before reaching the spermatecha ( Figs 6C, D, E, F ) (instead of going straight except for a short anterior curve in C. foliacea sp. nov., Figs 10C, D, E, F ). Males can be distinguished from its congeneric species by the presence of a guanine white-spot on the thoracic portion of prosoma ( Fig. 7A ) and by the palp presenting a long ribbon-shaped embolus that rests coiled on the tegular ridge instead of encircling the tegulum ( Fig. 7C ).

Description. Female (from Los Lagos,AMNH): Anterior eye row recurved and posterior row slightly recurved; ALE have almost twice the diameter of the AME ( Figs 6A, B ). Prosoma reddish-brown on the cephalic portion and yellowish-brown on the thoracic area, being entirely covered by needle-shaped and filiform setae ( Figs 6A, B ). Legs I and II predominantly yellowish-brown, with darker patellae; legs III and IV predominantly reddish-brown with metatarsi, tarsi and the proximal portion of the femora lighter. Opisthosoma with an accentuated concavity on the anterior margin, yellowish-brown on the dorsum and median-posterior area; reddish-brown on the sides and projections ( Fig. 6A ). Copulatory openings are long and narrowed ( Figs 6C, E ), leading to long and membranous copulatory ducts; spermathecae black, walnut-shaped and sclerotized ( Fig. 6D, F ). Measurements: eyes diameters and eyes interdistances: AME 0.09, ALE 0.15, PME 0.15, PLE 0.17, AME-AME 0.19, AME-ALE 0.09, PME-PME 0.17, PME-PLE 0.15. MOQ length 0.41, MOQ posterior width 0.51, MOQ anterior width 0.38; leg formula: 1243: leg I—femur 2.52/ patella 1.23/ tibia 1.69/ metatarsus 1.56/ tarsus 0.93/ total 7.93; II—2.28/ 1.06/ 1.47/ 1.29/ 0.80/ 6.90; III—1.28/ 0.81/ 0.79/ 0.63/ 0.52/ 4.03; IV—1.72/ 0.80/ 1.11/ 0.75/ 0.55/ 4.93. Prosoma length 2.50, width 2.62, opisthosoma length 3.56, total body length 6.06; clypeus height 0.35, sternum length 1.27, width 1.14, endites length 0.68, width 0.34, labium length 0.39, width 0.46.

Male (FMNH 71619): Eyes as in the female.Prosoma reddish-brown,lighter in the median portion and presenting a triangular guanine spot on the thoracic area ( Fig. 7A ). Anterior legs (I and II) light-brown and posterior (III and IV) predominantly yellow, darker on patellae. Cymbium presents a retrolateral concavity/arching to accommodate the embolus ( Figs 7D, F ). Tegulum discoid, rough-surfaced ( Figs 7C, E ); RTA short and acute. Measurements: eye diameters and interdistances: AME 0.08, ALE 0.12, PME 0.13, PLE 0.11, AME-AME 0.14, AME-ALE 0.09, PME-PME 0.16, PME-PLE 0.15. MOQ length 0.32, MOQ posterior width 0.41, MOQ anterior width 0.30; leg formula: 1243: leg I—femur 2.36/ patella 0.94/ tibia 1.86/ metatarsus 1.64/ tarsus 1.01/ total 7.81; II—1.96/ 0.80/ 1.40/ 1.27/ 0.78/ 6.21; III—1.02/ 0.57/ 0.70/ 0.55/ 0.46/ 3.30; IV—1.28/ 0.56/ 0.90/ 0.74/ 0.49/ 3.97. Prosoma length 2.07, width 1.90, opisthosoma length 2.02, total body length 4.09; clypeus height 0.30, sternum length 0.89, width 0.95, endites length 0.51, width 0.27, labium length 0.30, width 0.37.

Distribution. ARGENTINA: Neuquén, Río Negro, Chubut and Tierra del Fuego; CHILE: Coquimbo, Valparaíso, Santiago, Maule, Bío-Bío, Araucanía, Los Ríos, Los Lagos and Aysén ( Fig. 14B ).

Figures

FIGURE 6. Coenypha ditissima (Nicolet, 1849), female (AMNH). A–B habitus (A dorsal, B frontal); C, E epigyne ventral; D, F vulva dorsal.

FIGURE 7. Coenypha ditissima (Nicolet, 1849), male (FMNH 71619). A–B habitus (A dorsal, B frontal); C–F left palp (C, E ventral, D, F retrolateral).

FIGURE 10. Coenypha foliacea sp. nov., female (MHNS 8375). A–B habitus (A dorsal, B frontal); C, E epigyne ventral; D, F vulva dorsal.

FIGURE 14. Distribution records of Coenypha species. A C. antennata (triangles), C. edwardsi (circles); B C. ditissima (squares), C. nodosa (diamonds); C black triangles are C. trapezium sp. nov. (black triangles), C. foliacea sp. nov. (white triangles).