Phoenoteleia Kieffer, 1916

Lahey, Zachary, Musetti, Luciana, Masner, Lubomir & Johnson, Norman F., 2021, Revision of Phoenoteleia Kieffer (Hymenoptera, Scelionidae, Scelioninae), Journal of Hymenoptera Research 87, pp. 575-611 : 575

Phoenoteleia Kieffer

Phoenoteleia Kieffer, 1916: 62 (original description. Type: Phoenoteleia rufa Kieffer, by monotypy and original designation); Kieffer, 1926: 265, 550 (description, keyed); Dodd, 1929: 35 (description); Muesebeck and Walkley, 1956: 384 (citation of type species); Baltazar, 1966: 185 (cataloged, catalog of species of the Philippines); Masner, 1976: 31, 32 (description, key to males of Anteromorpha Dodd and Phoenoteleia Kieffer); Galloway and Austin, 1984: 7, 9, 20 (diagnosis, list of species described from Australia, keyed); Johnson, 1992: 460 (cataloged, catalog of world species); Austin and Field, 1997: 22, 68 (structure of ovipositor system, discussion of phylogenetic relationships).

Plagioscelio Kieffer, 1916: 185 (original description. Type: Plagioscelio rufescens Kieffer, by original designation. Synonymized by Masner (1976)); Kieffer, 1926: 266, 356 (description, keyed, key to species); Muesebeck and Walkley, 1956: 384 (citation of type species); Baltazar, 1966: 176 (cataloged, catalog of species of the Philippines); Masner, 1976: 32 (junior synonym of Phoenoteleia Kieffer).

Diagnosis.

Phoenoteleia can be distinguished from other scelionines by the following combination of characters: epitorular carinae present on frons; T1 of female always produced into an elongate horn which fits into a deep channel bisecting the metascutellum, mesoscutellum, and at least the posterior portion of the mesoscutum; venter of horn membranous; T1 with armilla; T2 clearly longer than T3; central portion of propodeum triangular, in same plane as mesoscutum, mesoscutellum, and metascutellum (male only); submarginal vein strongly curving towards costal margin of fore wing apically; marginal vein distinctly shorter than stigmal vein; postmarginal vein at least twice as long as stigmal vein; basitarsus on hind leg at least twice as long as combined length of remaining tarsomeres, distinctly incrassate in males.

Description.

Length 2.21-3.95 mm; body elongate, gracile to robust.

Head. Head shape in dorsal view: transverse. Hyperoccipital carina: absent. Occipital carina: present, complete. Length of OOL: lateral ocellus <1 OD from inner margin of compound eye. Shape of upper frons: convex. Antennal scrobe: undifferentiated from surrounding surface sculpture. Submedian carina: absent. Orbital carina: absent. Course of inner orbits: diverging ventrally. IOS/EH: IOS shorter than EH. Central keel: absent. Antennal foramen: oriented laterally on interantennal process. Facial striae: present. Malar striae: present. Malar sulcus: present. Setation of compound eye: present, short. Gena: convex, distinctly produced behind compound eye. Shape of clypeus: convex, trapezoidal, lateral corners not produced. Ventral margin of clypeus: rounded. Anteclypeus: absent. Postclypeus: absent. Labrum: transverse, visible anteriorly. Number of mandibular teeth: 3. Arrangement of mandibular teeth: transverse. Number of maxillary palpomeres: 3. Shape of maxillary palpomeres: cylindrical. Number of labial palpomeres: 2. Sculpture of occiput: granulate.

Antenna. Number of antennomeres in female: 12. Number of antennomeres in male: 12. Insertion of radicle into A1: parallel to longitudinal axis of A1. Shape of A1: more or less cylindrical, not flattened. Length of A3 of female: longer than A2, approximately as long or slightly longer than radicle. Claval formula: 1-2-2-2-1; 1-2-2-2-2; 1-2-2-2-2-1. Number of clavomeres: 5; 6. Arrangement of papillary sensilla: longitudinal. Antennomeres bearing tyloids in male antenna: A5. Shape of male flagellum: filiform.

Mesosoma. Transverse pronotal carina: present. Posterior apex of pronotum in dorsal view: articulate with tegula. Epomial carina: present. Anterior face of pronotum: visible dorsally, short. Netrion: present. Shape of netrion: wide, closed dorsally, open ventrally. Netrion sulcus: present. Anterior portion of mesoscutum: vertical, flexed ventrally to meet pronotum. Shape of mesoscutum: pentagonal, excavate at base of wings. Skaphion: absent. Parapsidial lines: absent; present. Antero-admedian lines: absent. Transscutal articulation: developed, narrow. Shape of mesoscutellum: transverse, present lateral to horn of T1 in females, complete in males. Transaxillar carina: present. Axillular carina: present. Lateral mesoscutellar spine: absent. Median mesoscutellar spine: absent. Axillular spine: absent. Surface of mesoscutellum: in same plane as mesoscutum. Median longitudinal furrow on mesoscutellum: absent. Metascutellum: transverse, lateral to horn of T1 in females, complete in males. Setation of metascutellum: absent. Lateral propodeal projection: absent. Medial portion of metascutellum in males: plate-like triangular, elevated relative to lateral portions. Median propodeal projection: absent. Subacropleural sulcus: present, indicated by a dorsoventral line of setae. Mesopleural carina: present. Mesepimeral sulcus: present. Posterior mesepimeral area: present. Sculpture of posterior mesepimeral area: smooth. Mesal course of acetabular carina: not separating fore coxae. Mesopleural pit: present. Metapleural sulcus: present. Paracoxal sulcus: present.

Legs. Number of mesotibial spurs: 1. Number of metatibial spurs: 1. Relative length of metabasitarsus: at least twice the length of remaining tarsomeres. Dorsal surface of metacoxa: smooth. Shape of metacoxa: cylindrical, ecarinate. Trochantellus: indicated by transverse sulcus on femur.

Wings. Wing development of female: macropterous. Wing development of male: macropterous. Tubular veins in fore wing: present. Setation of R vein: present, sparse, long. Bulla of fore wing R vein: absent. Length of marginal vein of fore wing: at least 2 times shorter than stigmal vein. Origin of stigmal vein (r-rs) in fore wing: arising from marginal vein along costal margin. Development of R vein in hind wing: complete, reaching frenal hooks.

Metasoma. Number of external metasomal tergites in female: 6. Number of external metasomal sternites in female: 6. Number of external metasomal tergites in male: 8. Number of external metasomal sternites in male: 7. Shape of metasoma: fusiform, narrowly constricted between T1 and T2 in females. Laterotergites: present. Laterosternites: present. T1 of female: produced into an elongate horn that bisects the metascutellum, mesoscutellum, and at least the posterior margin of the mesoscutum. Armilla: present. Relative size of metasomal terga: T2 longest (excluding horn of T1 in females), T3 longer than T4. Transverse sulcus on T2: present. Metasomal terga with basal crenulae: T2 in females, T1 and T2 in males. Sublateral carinae on metasomal terga: present. Median longitudinal carina on metasomal terga: absent. Shape of T6 in female: widest anteriorly. Anterior margin of S1: straight, not produced anteriorly. Felt fields on S2: present. Felt fields on S3: absent. Ovipositor: Scelio -type ( Austin and Field 1997).

Generic transfer of Phoenoteleia fusca (Kieffer)