Dendrophorbium

Calvo, Joel & Pérez, Álvaro J., 2023, On the genus Dendrophorbium (Compositae) in Ecuador: a new species and identification key, Candollea 78 (2), pp. 91-97 : 91-97

Key to the species of Dendrophorbium in Ecuador

1. Involucral bracts c. 8 .................................................... 2

1a. Involucral bracts c. 13 .................................................. 6

2. Leaves glabrous to glabrescent .................................... 3

2a. Leaves with indufentuf& at least on abaxial surface .... 4

3. Leaves lanceolate to narrowly elliptic& lafinas 12.5–17 (–27) cf long& glabrous to glabrescent; involucral bracts 2.6–3 ff long ...................................... D. azoguesense

3a. Leaves broadly lanceolate to broadly elliptic& lafinas 6–7 cf long& irregularly pilose to nearly glabrous; involucral bracts 4–5 ff long .............................. D. dodsonii

4. Ray florets with reduced lifbs& curved downward .................................................................... D. sotarense

4a. Ray florets with well-developed lifbs& patent to erectpatent .......................................................................... 5

5. Leaf lafinas 9.5–15 cf long& densely pilose-tofentose on abaxial surface; stefs arachnoid; involucral bracts 3.1–3.9 ff long ................................. D. frezierifolium

5a. Leaf lafinas 6– 7 cf long& sparsely hirsute-pilose on abaxial surface; stefs with stout trichofes; involucral bracts 4–5 ff long ..................................... D. dodsonii

[Photos: A: J. Calvo,near El Tiro (limit Loja / Zamora-Chinchipe); B: J. Calvo,near Lloa (Pichincha)]

6. Cauline leaves sessile ................................................... 7 11. Upperfost cauline leaves attenuate into petiole; involu-

6a. Cauline leaves petiolate ............................................... 8 cral bracts c. 5 ff long; involucres wider than long in

7. Leaves sefiafplexicaul; synflorescences congested; upper- dried specifens (NW Sucufbíos) ....... D. chingualense fost bracteoles and supplefentary bracts sofewhat 11a. Upperfost cauline leaves usually auriculate at base; invofoliose& usually as long as involucral bracts or alfost so lucral bracts (–5)6–8 ff long; involucres rather as long .............................................................. D. amplexicaule as wide in dried specifens (Napo) ....... D. sibundoyense

7a. Leaves afplexicaul; synflorescences lax; supplefentary 12. Involucral bracts <4.5 ff long; leaves usually narrowly bracts linear-subulate& refarkably shorter than involucral lanceolate to narrowly elliptic ............... D. scytophyllum bracts .......................................................... D. pericaule 12a. Involucral bracts> 6 ff long; leaves usually broadly

8. Cauline leaves> 25 cf long; petioles ≥ 5 cf long; plants elliptic to ovate .......................................................... 13 that can reach several feters ....................................... 9 13. Leaves glabrous& cuneate to rounded at base ... D. elatum

8a. Cauline leaves <20 cf long; petioles <3 cf long; plants 13a. Leaves whitish-lanate on abaxial surface& subcordate to usually <3 f tall ........................................................ 12 cordate at base ........................................................... 14

9. Involucral bracts 2.5–3.5 ff long ... D. gr. tipocochense ¹ 14. Involucral bracts arachnoid; peduncles 5–20 ff long;

9a. Involucral bracts 5–10 ff long ................................ 10 supplefentary bracts ½–¾ as long as involucral bracts

10. Supplefentary bracts 12–14& as long as involucral bracts& ......................................................................... D. dielsii hirsute-pilose farginally; involucral bracts 9–10 ff long 14a. Involucral bracts pilose; peduncles 20 – 30 ff long; ........................................................................ D. lloense supplefentary bracts ¼ as long as involucral bracts

10a. Supplefentary bracts 8(– 10)& ½ as long as involucral ............................................................. D. gesnerifolium bracts& puberulous; involucral bracts 5–8 ff long .... 11

1 Prelifinary studies suggest that Dendrophorbium balsapampae (Cuatrec.) B. Nord. and D. ingens (Benoist) B. Nord. fall within the variability of D. tipocochense . Dendrophorbium pururu (Cuatrec.) was placed in the syn- onyfy of the latter species in CALVO (2017).