Monteiroconus hoernesi ( Doderlein, 1863 )

Harzhauser, Mathias & Landau, Bernard, 2016, A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea, Zootaxa 4210 (1), pp. 1-178 : 109-110

Monteiroconus hoernesi ( Doderlein, 1863)

Figs 17 S, 26A1–A3, 26B1–B3, 26C1–C3

Conus Aldrovandi Brocc. —Hörnes 1851: 18, pl. 1, figs 2a–b [non Monteiroconus aldrovandi ( Brocchi, 1814) ]. [ Conus ] Hornesi [sic] nob.— Doderlein 1863: 25 [nov. nom. pro Conus aldrovandi in Hörnes 1851 ]. [ Lithoconus ] [ Conus ] Karreri n. f.— Hoernes 1878a: 195 (nomen nudum).

[ Lithoconus ] [ Conus ] Fuchsi n. f.— Hoernes 1878a: 195 (nomen nudum).

C. [onus] Karreri — Hoernes 1878b: 205 (nomen nudum).

Conus (Lithoconus) Karreri nov. form.— Hoernes & Auinger 1879: 26, pl. 4, fig. 7.

Conus (Lithoconus) Fuchsi nov. form.— Hoernes & Auinger 1879: 26, pl. 4, figs 4–5.

Lithoconus parvicaudatus (Sacco, 1893) — Kovács & Vicián 2013: 74, figs 77–78 [non Conus parvicaudatus Sacco, 1893 ].

non Conus (Dendroconus) aldrovandi Brocchi—Atanacković 1969: 213 , pl. 13, figs 1–1b [non Monteiroconus aldrovandi ( Brocchi, 1814) ] [= Kalloconus tietzei ( Hoernes & Auinger, 1879) ].

non Conus (Lithoconus) karreri Hoernes & Auinger, 1879 — Atanacković 1985: 180, pl. 40, figs 5–6 [= Kalloconus tietzei ( Hoernes & Auinger, 1879) ].

Type material. Lectotype (designated herein), NHMW 1851 /0002/0004, Grund ( Austria), specimen illustrated in Hörnes (1851, pl. 1, fig. 2); middle Miocene , Badenian ( Langhian ). The type locality of Conus karreri Hoernes & Auinger, 1879 , considered to be junior synonym of M. hoernesi , are the middle Miocene ( Badenian ) localities Bad Vöslau and Gainfarn in Austria . The type locality of Conus fuchsi Hoernes & Auinger, 1879 , considered to be junior synonym of M. hoernesi , is the middle Miocene (Badenian) locality Lăpugiu de Sus in Romania .

Studied material. Lectotype and 2 spec. NHMW 1868 /0001/0203, Grund ( Austria) ; 1 spec. NHMW 1874 / 0024/0058 Ritzing ( Austria) ; 1 spec. NHMW 1853 /0002/0045, Bad Vöslau ( Austria); 1 spec . NHMW 1849 /0023/ 0 0 0 4, Bad Vöslau ( Austria), syntype of Conus fuchsi illustrated in Hoernes & Auinger (1879, pl. 4, fig. 5); 1 spec . NHMW 1856 /0050/0110, Gainfarn ( Austria), syntype of Conus fuchsi illustrated in Hoernes & Auinger (1879, pl. 4, fig. 4); 1 spec . NHMW 1870 /0033/0001, Lăpugiu de Sus ( Romania), syntype of Conus karreri illustrated in Hoernes & Auinger (1879, pl. 4, fig. 7).

Illustrated material. Figs 26A 1 –A3: lectotype, Grund ( Austria): SL: 76.0 mm, MD: 49.1 mm, NHMW 1851/ 0002/0004, illustrated in Hörnes (1851, pl. 1, fig. 2); Figs 26 B1–B3: Lăpugiu de Sus ( Romania): SL: 78.2 mm, MD: 51.2 mm, NHMW 1870/0033/0001, illustrated in Hoernes & Auinger (1879, pl. 4, fig. 7); Figs 26 C1–C3, 17S: Gainfarn ( Austria): SL: 75.5 mm, MD: 47.5 mm, NHMW 1856/0050/0110, illustrated in Hoernes & Auinger (1879, pl. 4, fig. 4).

Revised description. Large robust shells; spire low and conical with weakly convex, blunt, smooth to faintly striate whorls; rarely a shallow concavity may be developed (“ fuchsi ”-morph). Late spire whorls slightly gradate; suture impressed. Subsutural flexure of medium depth, moderately curved, moderately asymmetrical. Prominent, rounded shoulder, with position of maximum diameter slightly below shoulder, whorl profile straight to slightly constricted below, resulting in a depressed weakly club-shaped profile. Moderately narrow aperture with shallow, narrow posterior canal; anteriorly widening, terminating in wide, very short, straight canal. Fasciole weakly swollen, indistinct, demarcated from twisted inner lip by deep groove; base somewhat constricted below siphonal fasciole. Faint spiral cords on last whorl, visible only in grazing light. No colour pattern preserved.

Shell measurements and ratios. n = 8: largest specimen: SL: 81.1 mm, MD: 53.9 mm, mean SL: 74.8 mm (σ = 2.1), mean MD: 48.7 mm (σ = 1.6), spire angle: µ = 123.6° (σ = 7.4°), last whorl angle: µ = 43.5° (σ = 2.8°), LW: µ = 1.5 (σ = 0.1), RD: µ = 0.75 (σ = 0.03), PMD: µ = 0.84 (σ = 0.01), RSH: µ = 0.13 (σ = 0.06).

Discussion. Doderlein (1863) introduced Conus Hornesi as new name for Conus Aldrovandi sensu Hörnes, 1851 (non Brocchi , 1814) including a very brief description. He obviously referred to the Viennese palaeontologist Moritz Hörnes and “ hornesi ” is a type setting error for “ hörnesi ”, which should be corrected to “ hoernesi ” according to ICZN Article 32.5. Therefore, we propose to emend the name to Conus hoernesi . Doderlein (1863) used the name for Tortonian specimens from Montegibbio and S. Agata in Italy but also as replacement name for the specimen of Hörnes (1851). Therefore, the Badenian shell from Grund in Austria is a syntype . For the sake of nomenclatural stability we select the specimen NHMW 1851 /0002/0004 from Grund ( Austria) as lectotype .

Hoernes & Auinger (1879) stated that the specimen of Hörnes (1851, pl. 1, fig. 2) is conspecific with their Conus karreri , which is confirmed herein, but obviously they overlooked the paper by Doderlein (1863). Therefore, Conus karreri is a subjective junior synonym of C. hoernesi . Conus fuchsi was separated by Hoernes & Auinger (1879) based on the slight mid-whorl constriction of the last whorl and the higher position of the fasciole. We consider these differences to reflect only intraspecific variability and treat the illustrated syntypes of Conus fuchsi as subadult specimens of Monteiroconus hoernesi . The seemingly concave and incised siphonal canal of the specimen illustrated by Hoernes & Auinger (1879, pl. 4, fig. 7) is an artefact due to fragmentation. All other specimens show a wide, straight and convex margin of the siphonal canal.

Monteiroconus hoernesi ( Doderlein, 1863) is separated from M. daciae and M. girondicus by its smaller maximum size and its less depressed spire. It is closely similar to M. conicomaculatus , from which it differs in being larger shelled, in having less strongly striate spire whorls and not having a weakly ventricose last whorl, as in M. conicomaculatus .

When describing Conus hungaricus Hoernes & Auinger, 1879 included also some specimens of Monteiroconus hoernesi as can be seen from their handwritten labels. Kalloconus hungaricus , however, differs in its much wider last spire whorl.

The specimens described by Kovács & Vicián (2013) as Lithoconus parvicaudatus (Sacco, 1893) might rather represent a low spired M. hoernesi , representing the “ fuchsi ”-morph with concave last whorl.

Paleoenvironment. Probably shallow marine coastal habitats based on the co-occurring mollusc assemblages (e.g. Letkés, Kovács & Vicián 2013, Ritzing, Harzhauser et al. 2014).

Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Bad Vöslau ( Austria); Alpine Carpathian Foredeep: Grund ( Austria) ( Hoernes & Auinger 1879); Oberpullendorf Basin: Ritzing ( Austria) (own data); Pannonian Basin: Letkés ( Hungary) ( Kovács & Vicián 2013); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus ( Romania) ( Boettger 1902, 1906).

Figures

FIGURE 1. Map of central and south – eastern Europe showing the geographic position of selected localities with rich Conidae diversities (modified from Kováč et al. 2007). 1: Korytnica (Poland), 2: Podhorce, Zukowce, Biłka, Tarnopol (Ukraine), 3: Grund, Guntersdorf, Immendorf (Austria), 4: Steinebrunn (Austria), Mikulov (Czech Republic), 5: Gainfarn, Enzesfeld, Baden, Bad Vöslau, Baden-Sooß (Austria), 6: Ritzing (Austria), 7: Weitendorf, Pöls (Austria), 8: Letkés, Szob (Hungary), 9: Borsodbóta (Hungary), 10: Várpalota (Hungary), 11: Hidas (Hungary), 12: Coşteiu de Sus, Lăpugiu de Sus (Romania).

FIGURE 26 A 1 – A 3. Monteiroconus hoernesi (Doderlein, 1863), Grund (Austria), NHMW 1851 / 0002 / 0004, lectotype. 26 B 1 – B 3. Monteiroconus hoernesi (Doderlein, 1863), Lăpugiu de Sus (Romania), NHMW 1870 / 0033 / 0001. 26 C 1 – C 3. Monteiroconus hoernesi (Doderlein, 1863), Gainfarn (Austria), NHMW 1856 / 0050 / 0110. 26 D 1 – D 3. Monteiroconus neugeboreni (Hoernes & Auinger, 1879), Lăpugiu de Sus (Romania), NHMW 1949 / 0005 / 0001. 26 E 1 – E 2. Monteiroconus pervindobonensis (Sacco, 1893), Bad Vöslau (Austria), NHMW 1869 / 0001 / 0020. 26 F 1 – F 3. Monteiroconus supracompressus (Sacco, 1893), Mikulov (Czech Republic), NHMW 1851 / 0010 / 0004, holotype. 26 G 1 – G 3. Monteiroconus supracompressus (Sacco, 1893), Mikulov (Czech Republic), NHMW 2016 / 0034 / 0001. 26 H. Monteiroconus supracompressus (Sacco, 1893), Mikulov (Czech Republic), NHMW 2016 / 0034 / 0001. 26 I. Monteiroconus mojsvari (Hoernes & Auinger, 1879), Pötzleinsdorf (Austria), NHMW 1847 / 0037 / 0024. 26 J. Monteiroconus mojsvari (Hoernes & Auinger, 1879), Gainfarn (Austria), NHMW 1853 / 0010 / 0001, syntype.