Phasmoconus fuscocingulatus (Hörnes, 1851),

Harzhauser, Mathias & Landau, Bernard, 2016, A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea, Zootaxa 4210 (1), pp. 1-178: 123-125

publication ID

http://doi.org/10.11646/zootaxa.4210.1.1

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scientific name

Phasmoconus fuscocingulatus (Hörnes, 1851)
status

 

Phasmoconus fuscocingulatus (Hörnes, 1851) 

Figs 17 W, 17X, 29A1–A3, 29B1–B3, 29C1–C3, 29D1–D3, 29E1–E2, 29F1–F3

Conus  fusco-cingulatus Bronn—Hörnes 1851: 21 (partim), pl. 1, figs 5a –c [non figs 4a –c, = Kalloconus moravicus ( Hoernes & Auinger, 1879)  ].

Conus (Chelyconus) fuscocingulatus Bronn—Hoernes & Auinger 1879: 47  , pl. 1, figs 10–13.

[ Conus  ( Dendroconus  )] ochreocingulata Sacc.  — Sacco 1893a: 12 [nov. nom. pro Conus fuscocingulatus in Hoernes & Auinger 1879  , pl. 1, figs 10–11].

[ Conus  ( Dendroconus  )] pötzleinsdorfensis Sacc.— Sacco 1893a: 12 [nov. nom. pro Conus fuscocingulatus in Hoernes & Auinger 1879  , pl. 1, fig. 13].

Conus fuscocingulatus Bronn—Friedberg 1911: 54  , pl. 2, figs 17–18.

Conus (Lithoconus) fuscocingulatus Bronn, 1848  — Moisescu 1955a: 163, pl. 15, figs 1–4.

Conus (Lithoconus) friedbergi  nomen nov.— Moisescu 1955a: 165, pl. 14, figs 5–6.

Conus fuscocingulatus Bronn.  — Eremija 1959: 187, pl. 1, figs 7–7a.

Conus (Dendroconus) fuscocingulatus Bronn 1848  —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 214, pl. 51, fig. 2.

Conus (Dendroconus) fuscocingulatus Bronn—Florei 1961: 689  , pl. 9, fig. 67.

? Conus (Chelyconus) fuscocingulatus Bronn, 1848  — Hinculov 1968: 149, pl. 37, figs 11–13 [non fig. 10].

Conus (Chelyconus) friedbergi Moisescu, 1955  — Hinculov 1968: 149, pl. 37, figs 14a –b.

Conus (Dendroconus) berghausi exfuscocingulatus (Sacco, 1893)  — Hinculov 1968: 152, pl. 38, fig. 9 [non Conus exfuscocingulatus Sacco, 1893  ].

Conus  (Dendroconu s) fuscocingulatus Bronn—Atanacković 1963: 78  , pl. 15, figs 5–5a.

Conus (Chelyconus) fuscocingulatus Bronn—Nicorici 1972: 70  , pl. 17, figs 5–6.

Conus (Chelyconus) fuscocingulatus Bronn in Hörnes, 1856  — Nicorici & Sagatovici 1973: 176, pl. 27, figs 4–5.

Conus (Chelyconus) fuscocingulatus M. Hoernes—Švagrovský 1982: 403  , pl. 5, fig. 3.

Conus  (Dendroconu s) fuscocingulatus Bronn, 1848  — Atanacković 1985: 181, pl. 40, figs 11–12.

Conus fuscocingulatus Bronn—Ionesi & Nicorici 1994: 62  , pl. 5, figs 1–4.

? Conus (Chelyconus) fuscocingulatus Bronn—Tiţă 2007: 554  , fig. 6/c.

non Conus (Chelyconus) fuscocingulatus Bronn in Hörnes, 1856  — Strausz 1966: 459, pl. 68, figs 8–11. non Conus (Chelyconus) fuscocingulatus Bronn—Stancu et al. 1971: 126  , pl. 8, fig. 8.

non Conus fuscocingulatus  Bronn—Csepreghy-Meznerics 1972: 34, pl. 17, fig. 20.

non Conus (Chelyconus) fuscocingulatus Hörnes, 1856  — Švagrovský 1981: 152, pl. 48, fig. 10. non Dendroconus fuscocingulatus (Bronn in Hörnes, 1856)  — Kovács & Vicián 2013: 66, figs 43–46.

Type material. Syntype NHMW 1846View Materials /0037/0055, Pötzleinsdorf ( Austria), illustrated in Hörnes (1851, pl. 1, fig. 5); middle Miocene , Badenian (Serravallian). 

Studied material. Syntypes and 6 spec. NHMW 1861View Materials /0033/0001, Bujtur ( Romania), 7 spec  . NHMW 1861View Materials / 0033/0002, Bujtur ( Romania) including specimen illustrated in Hoernes & Auinger (1879, pl. 1, figs 10–11), including syntypes of Conus ochreocingulatus Sacco, 1893  ; 32 spec  . NHMW 1846View Materials /0037/0054a, Pötzleinsdorf ( Austria); 8 spec  . NHMW A1608, Bujtur ( Romania), 41 spec  . NHMW 1836View Materials /0012/0108, Bujtur ( Romania). 

Illustrated material. Figs 29AView FIGURE 29 1View FIGURE 1 –A3: Bujtur (Romania), SL: 36.5 mm, MD: 18.2 mm, NHMW 1861View Materials /0033/ 0 0 0 1, Bujtur ( Romania)  ; Figs 29View FIGURE 29 B1–B3: Bujtur (Romania), SL: 34.1 mm, MD: 17.5 mm, NHMW 1861View Materials /0033/0001, Bujtur ( Romania)  ; Figs 29View FIGURE 29 C1–C3: Bujtur ( Romania), SL: 31.7 mm, MD: 18.7 mm, NHMW A1608 (poetzleinsdorfensis- morph); Figs 29View FIGURE 29 D1–D3, 17W: Bujtur ( Romania), SL: 34.4 mm, MD: 17.6 mm  , NHMW 1861View Materials / 0033/0002, specimen in Hoernes & Auinger (1879, pl. 1, fig. 11); Figs 29View FIGURE 29 E1–E2: Bujtur ( Romania), SL: 30.8 mm, MD: 15.6 mm  , NHMW 1836View Materials /0012/0108a; Figs 29View FIGURE 29 F1–F 3, 17X: Bujtur ( Romania), SL: 24.7 mm, MD: 14.7 mm  , NHMW 1836/0012/0108b (poetzleinsdorfensis- morph).

Revised description. Moderately small to medium-sized shells; paucispiral protoconch; high conical initial spire; later spire low, with beaded keel just above suture; beads move towards suture within 3rd –5th spire whorls and fade out as indistinct swellings within 6th whorl. Later spire of variable height, usually elevated and slightly coeloconoid; spire whorl tops weakly convex, faintly striate (only visible in high magnification); impressed suture. Subsutural flexure variable, very shallow, weakly curved, moderately asymmetrical. Last spire whorl slightly broadening with indistinct shoulder. Last whorl moderately elongate, weakly ventricose, slightly constricted; aperture regularly widening towards long and reflected anterior canal. Fasciole swollen, long, twisted, often with weak axial threads; inner lip narrow, straight. Deep irregularly spaced spiral grooves with broad cords in lower third of last whorl. Colour pattern often preserved as brown lines and very prominent in UV light, consisting of about 12–15 rather regularly spaced, well defined, continuous spirals on last whorl (rarely discontinuous spirals occur), with broad interspaces; broadest interspace usually two spirals below shoulder. Spire with axially elongate flammulae and blotches especially on shoulder and last whorl; some specimens display a broad dark band on spire whorl tops (in UV) and a light spiral at suture.

Shell measurements and ratios. n = 14: largest specimen: SL: 39.7 mm, MD: 21.9 mm, mean SL: 34.4 mm (σ = 2.3), mean MD: 18.5 (σ = 1.6), spire angle: µ = 104.6° (σ = 11.2°), last whorl angle: µ = 35.2° (σ = 2.1°), LW: µ = 1.87 (σ = 0.14), RD: µ = 0.63 (σ = 0.04), PMD: µ = 0.89 (σ = 0.03), RSH: µ = 0.14 (σ = 0.02).

Discussion. The authorship is often attributed to Bronn, who used the name in his correspondence and listed it in Bronn (1848) as nomen nudum (considering it as junior synonym of Conus raristriatus Bellardi & Michelotti, 1840  ). Hörnes (1851) made the name available and based his description on specimens from numerous localities of the Austro-Hungarian Empire, clearly mixing several different species. Consequently, the status of this species is highly confused. Hörnes (1851) illustrated specimens as Conus fuscocingulatus  , which represent two different species. He illustrated a specimen from Mikulov ( Czech Republic) as figure 4 on plate 1 and a second one as variety from Pötzleinsdorf ( Austria) as figure 5. Later, Hoernes & Auinger (1879) separated the specimen from Mikulov as Conus moravicus  and kept the “variety” of Hörnes (1851) in the synonymy of Conus fuscocingulatus  . The confusion was completed by Hoernes & Auinger (1879) by illustrating additional specimens as C. fuscocingulatus  , which represent at least three species. The specimens illustrated by Hoernes & Auinger as fig. 12 is a stout shell with very low spire and distinct shoulder; under UV light it displays an irregular pattern of densely spaced, spirally arranged dashes and differs completely from Phasmoconus fuscocingulatus  [the illustration of the colour pattern in Hoernes & Auinger (1879) is incorrect]. It is most probably a Kalloconus moravicus ( Hoernes & Auinger, 1879)  . Their fig. 13 is also a very stout shell with nearly flat spire. For this specimen Sacco (1893a) introduced Conus poetzleinsdorfensis  as new name. Unfortunately, the specimens of Hörnes (1851, pl. 1, fig. 5) and Hoernes & Auinger (1879, pl. 1, fig. 10) are rather untypical for Phasmoconus fuscocingulatus  due to the broad last whorl, whereas all other specimens are more slender. Consequently, Sacco (1893a) proposed Conus ochreocingulata  as new name for the shells o Hoernes & Auinger (1879, pl. 1, figs 10–11), which are treated as typical P. fuscocingulatus  herein. This confusion is reflected in a very broad range of morphologies and lumped species identified in Paratethyan literature as this species. A Principal Component Analysis of the available specimens did not allow any separation of the “ poetzleinsdorfensis  ”-morphs from Phasmoconus fuscocingulatus  (Fig. 12); moreover, both morphs co-occur at Pötzleinsdorf and Bujtur. Therefore, we treat Conus poetzleinsdorfensis Sacco, 1893  and Conus ochreocingulata Sacco, 1893  as subjective junior synonyms of P. fuscocingulatus  .

Moisescu (1955) introduced Conus friedbergi  for specimens from Bujtur (Romania), which do not differ from P. fuscocingulatus  . In her discussion she referred to the specimen from Białogon (Poland) illustrated by Friedberg (1911, pl. 2, fig. 19) as Conus cf. avellana Lamarck. This  specimen is lost and the illustration does not allow a clear identification. As Moisescu (1955) did not clearly state whether she considered the Polish or her Romanian specimen as type specimen, the status of Conus friedbergi  remains vague. It is either a nomen dubium (or species inquirenda) or a subjective junior synonym of P. fuscocingulatus  .

Paleoenvironment. Most probably shallow marine environments based on the co-occurring mollusc assemblage and the geological setting at the very margin of the Vienna Basin (Sieber 1954).

Distribution in Paratethys. Badenian (middle Miocene): widespread in all Paratethyan basins.

NHMW

Naturhistorisches Museum, Wien

Kingdom

Animalia

Phylum

Mollusca

Class

Gastropoda

Order

Neogastropoda

Family

Conidae

Genus

Phasmoconus

Loc

Phasmoconus fuscocingulatus (Hörnes, 1851)

Harzhauser, Mathias & Landau, Bernard 2016
2016
Loc

Conus

Atanackovic 1985: 181
1985
Loc

Conus (Chelyconus) fuscocingulatus Hörnes, 1856

Kovacs 2013: 66
Svagrovsky 1981: 152
1981
Loc

Conus (Chelyconus) fuscocingulatus Bronn in Hörnes, 1856

Nicorici 1973: 176
1973
Loc

Conus (Chelyconus) fuscocingulatus

Hinculov 1968: 149
1968
Loc

Conus (Chelyconus) friedbergi

Hinculov 1968: 149
1968
Loc

Conus (Dendroconus) berghausi exfuscocingulatus

Hinculov 1968: 152
1968
Loc

Conus (Chelyconus) fuscocingulatus Bronn in Hörnes, 1856

Strausz 1966: 459
1966
Loc

Conus (Dendroconus) fuscocingulatus

Kojumdgieva 1960: 214
1960
Loc

Conus fuscocingulatus

Eremija 1959: 187
1959
Loc

Conus (Lithoconus) fuscocingulatus

Moisescu 1955: 163
1955
Loc

Conus (Lithoconus) friedbergi

Moisescu 1955: 165
1955
Loc

Conus

Sacco 1893: 12
1893
Loc

Conus

Sacco 1893: 12
1893
Loc

Conus (Chelyconus) fuscocingulatus Bronn—Hoernes & Auinger 1879 : 47

Auinger 1879: 47
Loc

Conus fuscocingulatus Bronn—Ionesi & Nicorici 1994 : 62

Nicorici 1994: 62