Kalloconus moravicus ( Hoernes & Auinger, 1879 )

Harzhauser, Mathias & Landau, Bernard, 2016, A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea, Zootaxa 4210 (1), pp. 1-178 : 63-64

Kalloconus moravicus ( Hoernes & Auinger, 1879)

Figs 3 N, 11A1–A5, 11B1–B3

Conus fusco-cingulatus Bronn—Hörnes 1851: 21 (partim), pl. 1, figs 4a–c [non Phasmoconus fuscocingulatus (Hörnes, 1851) ]. [ Lithoconus ] [ Conus ] moravicus n. f. — Hoernes 1878a: 195 (nomen nudum).

Conus (Lithoconus) Moravicus nov. form.— Hoernes & Auinger 1879: 29 (partim) [nov. nom. pro Conus fusco-cingulatus in Hörnes 1851, pl. 1, figs 4a–c].

non Conus (Lithoconus) moravicus R. Hoernes et M. Auinger—Švagrovský 1982: 404 , pl. 5, fig. 4 [= Monteiroconus supracompressus (Sacco, 1893) ].

Type material. 7 syntypes NHMW 1851 /0010/0001, including specimen illustrated in Hörnes (1851, pl. 1, fig. 4) ; 3 syntypes NHMW 1851 /0020/0001, all Mikulov-Kienberk ( Czech Republic); middle Miocene , Badenian (late Langhian).

Studied material. Syntypes and 12 specimens, NHMW 2016 /0033/0001, Mikulov-Kienberk ( Czech Republic) ; 15 spec. NHMW 1846 /0037/0057, Gainfarn ( Austria) ; 4 spec. NHMW 1848 /0003/0004, Ritzing ( Austria) ; 1 spec. NHMW 1865 /0035/0025, Szob ( Hungary) .

Illustrated material. Figs 11A 1 –A5: Mikulov-Kienberk ( Czech Republic): SL: 40.2 mm, MD: 26.2 mm, NHMW 2016/0033/0001; Figs 11 B1–B3: syntype, Mikulov-Kienberk ( Czech Republic): SL: 47.5 mm, MD: 32.0 mm, NHMW 1851/0010/0001, illustrated in Hörnes (1851, pl. 1, fig. 4); Fig. 3 N: Mikulov-Kienberk ( Czech Republic): SL: 35.2 mm, MD: 23.1 mm, NHMW 2016/0033/0001.

Revised description. Medium sized solid shells with very low conical spire and short, broad conical last whorl. Spire whorls nearly flat to weakly concave in adapical half and slightly convex abapically; early spire whorls distinctly striate, with two striae demarcating impressed suture. Very broad, poorly defined swellings along shoulder of 3rd -6th teleoconch whorl, causing undulating suture. Last two whorls smooth with very irregular suture. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Last whorl broad with rounded, prominent shoulder. A weak concavity appears in many specimens slightly below shoulder, resulting in stout, ficoid outline. Last whorl straight-sided or weakly ventricose below concavity; indistinctly constricted at base. Siphonal canal short and wide; fasciole short, moderately swollen, well demarcated from short inner lip; aperture moderately wide with subparallel margins. Densely spaced wavy spiral grooves and cords on base. Colour pattern frequently preserved, consisting of 12–16 thin, continuous spirals separated by broader interspaces.

Shell measurements and ratios. n = 13 specimens: largest specimen: SL: 47.5 mm, MD: 32.0 mm, mean SL: 39.4 mm (σ = 4.4), mean MD: 26.2 (σ = 3.3), spire angle: µ = 132.1° (σ = 8.0°); last whorl angle: µ = 37.9° (σ = 1.6°), LW: µ = 1.5 (σ = 0.04), RD: µ = 0.72 (σ = 0.02), PMD: µ = 0.91 (σ = 0.03), RSH: µ = 0.08 (σ = 0.03); the large specimen of 70 mm height, mentioned by Hoernes & Auinger (1879), is based on a misidentification.

Discussion. The colour pattern is highly reminiscent of Phasmoconus fuscocingulatus (Hörnes, 1851) , which was the reason why Hörnes (1851) mixed both species. The broad conical and often fig-shaped last whorl of K. moravicus , however, allows a separation of both species (Fig. 12). Nevertheless, the original species concept of Hoernes & Auinger (1879) was blurred by lumping different species. Especially juvenile shells of Monteiroconus supracompressus (Sacco, 1893) were mixed with K. moravicus in the type lots. Similarly, Švagrovský (1982) misidentified a Monteiroconus supracompressus as K. moravicus (both species are found at the locality described by Švagrovský 1982). The more elongate last whorl of M. supracompressus and its nearly flat spire, separate both species distinctly.

Paleoenvironment. Typically found in shallow marine, sandy nearshore environments with seagrass.

Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Niederleis, Gainfarn, Enzesfeld, Steinebrunn, Pötzleinsdorf, Baden, Bad Vöslau ( Austria), Mikulov-Kienberk ( Czech Republic), Borský Mikuáš ( Slovakia) ( Hoernes & Auinger 1879; Švagrovský 1982); Eisenstadt-Sopron Basin: Forchtenau ( Austria) ( Hoernes & Auinger 1879); Oberpullendorf Basin: Ritzing ( Austria) ( Hoernes & Auinger 1879); Styrian Basin: Gamlitz ( Austria) ( Hoernes & Auinger 1879); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus ( Romania) ( Boettger 1906).

Figures

FIGURE 1. Map of central and south – eastern Europe showing the geographic position of selected localities with rich Conidae diversities (modified from Kováč et al. 2007). 1: Korytnica (Poland), 2: Podhorce, Zukowce, Biłka, Tarnopol (Ukraine), 3: Grund, Guntersdorf, Immendorf (Austria), 4: Steinebrunn (Austria), Mikulov (Czech Republic), 5: Gainfarn, Enzesfeld, Baden, Bad Vöslau, Baden-Sooß (Austria), 6: Ritzing (Austria), 7: Weitendorf, Pöls (Austria), 8: Letkés, Szob (Hungary), 9: Borsodbóta (Hungary), 10: Várpalota (Hungary), 11: Hidas (Hungary), 12: Coşteiu de Sus, Lăpugiu de Sus (Romania).

FIGURE 11 A 1 – A 5. Kalloconus moravicus (Hoernes & Auinger, 1879), Mikulov-Kienberk (Czech Republic), NHMW 2016 / 0033 / 0001. 11 B 1 – B 3. Kalloconus moravicus (Hoernes & Auinger, 1879), Mikulov-Kienberk (Czech Republic), NHMW 1851 / 0010 / 0001, syntype. 11 C 1 – C 3. Kalloconus neumayri (Hoernes & Auinger, 1879), Lăpugiu de Sus (Romania), NHMW A 1628. 11 D 1 – D 3, 33 D. Kalloconus neumayri (Hoernes & Auinger, 1879), Lăpugiu de Sus (Romania): NHMW 1999 z 0077 / 0027, syntype, 11 E 1 – E 2. Kalloconus neumayri (Hoernes & Auinger, 1879), Gainfarn (Austria), NHMW 1843 / 0099 / 0058. 11 F 1 – F 2. Kalloconus neumayri (Hoernes & Auinger, 1879), Lăpugiu de Sus (Romania), NHMW A 1628. 11 G 1 – G 3. Kalloconus ponderoaustriacus (Sacco, 1893), Lăpugiu de Sus (Romania), NHMW 1854 / 0035 / 0041, holotype. 11 H 1 – H 3. Kalloconus ponderoaustriacus (Sacco, 1893), Gainfarn (Austria), NHMW 1855 / 0045 / 0358.