Melittomma pervagum ( Olliff, 1889 )

Lawrence, John F., 2020, The Australian Lymexylidae (Coleoptera: Tenerionoidea) with one new genus and two genera new to Australia, Zootaxa 4895 (2), pp. 211-238 : 219-222

Melittomma pervagum ( Olliff, 1889)

(Figs. 13–14, 18, 32, 35)

Hylecoetus pervagus Olliff, 1889: 85 .

Hylecoetus vigilans Lea, 1912: 467 , pl. 17, fig. 8. Syn. nov.

Diagnosis. This species may be distinguished from other species known to this author by several features which also distinguish it from all other Australian melittommatines. The normal Melittomma pronotum is widest at or behind middle and has an evenly rounded anterior edge without anterior angles; in this respect it is similar to those of all known Alcestoma and Australymexylon . The pronotum of M. pervagum , however, is widest near the anterior edge with obtuse but distinct anterior angles between the lateral and anterior margins. The only species which may have a similar pronotum, based on the figure in Villiers (1969, fig. 15) is Melittomma africanum (Thomson) . Other features, such as strongly divergent pronotosternal sutures, highly reduced prosternal process, postcoxal (hypomeral) processes and mesoventrital process (all virtually absent), and mesocoxal cavities partly closed laterally by both mesanepisternum and mesepimeron, all differ from other melittommatines examined. On the other hand, this species possesses the appendiculate paratergites and aedeagal type found in other Melittomma species, the male abdominal apices of which have been examined or illustrated ( Villiers 1969; Wheeler 1986; Fonseca & Vieira 2000; Orozco & Dias 2018).

Redescription. Total length: males, 5.20–14.70 mm (7.90 ± 2.10, n = 65), female 8.00–30.00 (18.90 ± 5.80, n = 78); body 4.13–6.12 (5.04) times as long as combined elytral width. Colour of head black; pronotum reddishbrown to dark brown; elytra yellowish-brown to chestnut brown or dark brown, undersurfaces and legs yellow to yellowish-brown; antennae usually reddish-brown; mandibles red to black, maxillar and labium yellow to yellowish-brown, but male palp organ black. Upper surfaces densely, more or less evenly clothed with short suberect to decumbent, fine, yellow hairs; undersurfaces and legs with similar hairs. Head slightly longer than wide, abruptly constricted to form neck, which is 0.75 times as wide as head behind eyes. Eyes very large, contiguous in male and very narrowly separated in female, protruding and finely facetted, with short, fine interfacetal hairs. Temples very short and abrupt, barely visible from above. Antennal insertions concealed by frontal ridges which extend slightly into eye emargination. Frontoclypeal suture distinct. Clypeus 0.25 times as long as wide with truncate apex. Labrum 0.72 times as wide as clypeus, 0.75 times as long as wide, with apex subangulate and surface densely setose. Antennae in both sexes about as long as head width including eyes. Ratio of antennomere lengths: male: 1.80: 1.20: 2.20: 1.30: 1.20: 1.10: 1.10: 1.00: 1.20: 1.20: 3.40; female: 1.60: 1.00: 1.90: 1.30: 1.20:1.20: 1.20: 1.20: 1.20: 1.20: 2.80. Antennomere length/width ratios: male: 1.50, 1.09, 2.00, 1.08, 1.09, 1.00, 1.00, 0.91, 1.50, 1.50, 4.25; female: 1.45, 0.83, 1.27, 0.68, 0.67, 0.67, 0.67, 0.75, 0.80, 0.92, 2.64. The intermediate antennomeres in the male are about as long as wide and more or less keg shaped, while those in the female are definitely shorter than wide and subserrate; terminal antenomere in both sexes subacute at apex. Mandible about as long as wide; outer edge slightly, evenly curved, apex bidentate; incisor edge simple or with indistinct tooth; mola and prostheca absent. Galea 2.08 times as long as wide, widest near broadly rounded apex, densely setose; lacinia slightly shorter than and 0.25 times as wide as galea, subacute apically; first maxillary palpomere in male 0.9 times as long as wide, 0.75 times as long as second, which is 0.67 times as long as wide; third palpomere about 1.8 times as long as second, 0.73 times as long as wide, more or less globular and deeply excavate and fourth palpomere 1.80 times as long as third, 5.0 times as long as wide and narrowly rounded apically; palp organ relatively complex, with at about 8 primary lobes and numerous secondary lobes. Apical maxillary palpomere in female about 2.50 times as long as wide, barely wider at middle, with obliquely truncate apex. Mentum 0.44 times as long as wide, widest at truncate apex; ligula longer than mentum, widest at apex, which is deeply emarginate forming two lobes; labial palps separated by slightly less than the basal width of one; apical palpomere 2.50 time as long as wide, widest at about middle with narrowly rounded apex. Submentum not separated from gula; gular sutures subparallel. Cervical sclerites each consisting of two complexly curved plates, about equal in length, the anterior one with a small setose patch. Pronotum 0.95–1.15 (1.06) times as long as wide, widest at anterior third; sides straight and diverging from near base to anterior third, where a pair of weak, obtuse anterior angles separate them from the strongly curved anterior edge; posterior angles slightly, posterolaterally produced and subacute with finely notched apex; posterior edge weakly bisinuate, with narrow bead and transverse, submarginal groove; lateral margins very narrow and not extending onto anterior edge; disc moderately convex but with distinctly elevated area posteromesally; finely, densely punctate, with punctures usually separated by less than half a puncture diameter and enclosing one relatively short, fine, yellowish seta plus a small pore. Prosternum about as long as mid length of procoxal cavity, slightly, evenly convex, with straight, notosternal sutures strongly diverging anteriorly and extending to lateral edge just beyond anterior angles. Prosternal process very short, broad and apically truncate, not extending between coxae. Procoxal cavities large and broadly open externally and internally, with small, weak notch at base of notosternal suture; postcoxal (hypomeral) process very short, broad and rounded; protrochantin reduced but externally visible, more or less trapezoidal; endopleuron short, broadly expanded at apex. Scutellar shield slightly longer than wide, widest at base with sides narrowing to truncate apex, neither elevated nor separated from scutum, which lacks median endocarina. Elytra 3.23–5.13 (4.08) times as long as wide and 3.33–5.31 (5.04) times as long as pronotum, with sides more or less parallel, outer angles broadly rounded and apices independently narrowly rounded but narrowly separated; humeri weakly developed; disc with four very weak, longitudinal costae; punctation fine, dense and confused, each puncture bearing a moderately long, fine, inclined seta. Mesoventrite subtriangular, 0.75 times as long as wide, widest at posterior end and gradually narrowed to anterior end which is very narrow and truncate, separating the narrow mesanepisternal coxal rests; anterior portion of mesoventrite simple without longitudinal or transverse ridges. Posterior edge of mesoventrite forming a very short, apically truncate intercoxal process. Mesocoxal cavities large, weakly impressed and contiguous, broadly closed laterally by both mesanepisternum and mesepimeron; mesotrochantin visible, small, ovate. Metaventrite 1.40 times as long as wide, strongly convex, longest anterolaterally, with discrimen very weakly indicated but extending anteriorly almost as far as posterior edges of coxal cavities; posteror edge of metaventrite undulate on either side of small projection cleft at apex. Metanepisternum 4.00 times as long as wide, widest at anterior end and gradually narrowed to truncate apex; metepimeron concealed. Metendosternite with very long stalk, paired, elongate-oval laminae about twice as long as wide, slender lateral arms extending dorsally and a very short anterior process more or less continuous with laminae and bearing a pair of relatively widely separated anterior tendons. Hind wing about 2.75 times long as wide; apical field 0.22 times as long as wing length, with anterior oblique and posterior oblique linear sclerites meeting at base of field and extending to wing membrane. Radial cell about 0.04 times wing length, 2,0 times as long as wide, with straight base; cross-vein r3 absent, r4 long, oblique and slightly curved; radio-medial loop forming a 45° angle with apex of MP 1+2 and basal portion of RP extending to basal fourth. MP 1+2 strongly developed, with medial spur straight, extending to wing margin. Medial field with four veins extending to wing margin: MP 3+4 undivided, without basal crossvein or stub; wedge cell 6.3 times as long as wide, apically obliquely truncate, giving rise to CuA 1+2, which is moderately long, oblique and forked to form short CuA 1 meeting MP 3+4 and a long CuA 2; AA 4 very close to CuP + AA 3. Anal embayment absent; anal lobe large with two veins (AP 1+2, AP 3+4). Procoxa 3.18 times as long as wide, widest near base and slightly narrowed to apex, strongly projecting. Mesocoxa 2.08 times as long as wide, widest near base and narrowed apically, projecting posteriorly. Metacoxa 0.60 times as long as wide, strongly oblique and projecting, with distinct femoral grooves. Trochanters all small and subtiangular; trochanterofemoral joints strongly oblique so that femur just meets coxa. Profemur 3.07 times as long as wide; protibia 5.6 times as long as wide; protarsus about as long as tibia, with segment length ratio 4.8: 2.4: 1.2: 1.0: 2.8. Mesofemur 3.62 times as long as wide, widest at middle, with slight tibial groove; mesotibia 8.2 times as long as wide, more or less parallel sided; mesotarsus about 1.6 times as long as tibia, with segment length ratio 3.25: 1.63: 1.25: 1.00: 1.25. Metafemur 3.27 times as long as wide, widest at middle, with slight tibial groove; Metatibia 10 times as long as wide, very slightly wider apically and inwardly curved; mesotarsus about 1.25 times as long as tibia, with segment length ratio 3.82: 2.17: 1.67: 1.00: 1.67. Tibial spurs all relatively short and of equal lengths. Pretarsal claws moderately long and slender with relatively small, bisetose empodium. Abdomen with first ventrite (sternite III) slightly shorter than second, with intercoxal process very weakly developed or absent; concealed sternite II well-developed with weak median carina; ventrites 2–4 more or less equal in length and 5 slightly shorter than 4 and broadly rounded or subtruncate apically; abdominal tergites III–VI weakly sclerotised, VII (pygidium) moderately sclerotised with apex broadly rounded in female, subtruncate and slightly raised in male. Abdominal sternite VIII in male about 0.87 times as long as wide, widest near base and produced to form narrowly rounded apical lobe, lightly sclerotised basolaterally and membranous apicomesally; tergite VIII subquadrate, moderately sclerotised, with truncate base, slightly produced, rounded, setose apex, basomesal endocarina extending to about middle of sclerite, a pair of laterotergal appendages, each of which is about twice as long as wide, slightly curved mesally, narrowly rounded at apex with a number of short spines along internal edge, plus a flattened, sclerotised lobe at the base of each and extending to about middle of appendage. Segment IX with pregenital ring narrowly rounded anteriorly and tergite membranous. Aedeagus normally oriented, with tegmen dorsal to penis; tegmen with basale 2.33 times as long as wide, widest near base, base slightly acute; median linear sclerite extending from apex almost to base; apicale 0.4 times as long as basale, 1.8 times as long as wide, parallelsided with deeply emarginate apex forming a pair of obliquely rounded parameroids; base of apicale attached to a pair of long tegminal struts extending anteriorly and mesally to join at midline near base of penis. Penis 4.15 times as long as wide, widest at basal third with narrowly rounded apex; penis distinctly curved ventrally, with slender median projection at base which is attached to tegminal struts. Segment VIII in female weakly sclerotised at base and membranous apically, with long spiculum ventrale forked at base. Ovipositor about 6.5 times as long as wide, with distinct proctigeral and paraproctal but reduced coxital bacula; paraprocts 1.8 times as long as gonocoxites, which are 2.0 times as long as combined width; basal coxital lobes moderately heavily, evenly sclerotised, without basal bacula, broad at base, abruptly narrowed apically to form a pair of slender, contiguous, apically narrowly acute processes; apical lobes obliquely attached to basal lobes, membranous, densely setose, subcontiguous and apically truncate; gonostylus 0.23 times as long as coxite, apicolaterally attached, expanded apically and densely setose.

Types. Hylecoetus pervagus Olliff, 1889 : Holotype, sex?, “Lord Howe Island” (AMS). Hylecoetus vigilans Lea, 1912 : Holotype ♀, “Little Mulgrave River (H. Hacker) (SAM). Syn. nov.

Other specimens examined. QLD: Atherton , 4 mi. S, 11.2.1975 , H. & A. Howden (1♁, ANIC); Babinda , 28.xii.1972 , M. S. Moulds (1♀, ASCU); Babinda , 4 mi. W, 10.iii.1964 , I. F. B. Common, M. S. Upton (1♀, ANIC); Baldy Mtn. Rd. , 6 mls SW of Atherton, 1100m, 27.xii.1972 , B. Cantrell (1♀, QMB); Barron Falls, Kuranda , 24.x.1944 , at light, J. G. Brooks (4 ♀, QDAF; 1♁, 8♀, ANIC); Barron Gorge Lookout, Kuranda , 27.i.1989 , A. Sund-holm, J. Bugeja (1♀, ANIC); Binna Burra , 27.xii.1969 , MV light, R. Hardie (1♀, ANIC); Binna Burra, Lamington Nat. Park, J. Balderson (1♀, ANIC); Boar Creek , xii.1969 , J. G. Brooks (1♁, ANIC); Boar Pocket , x.1970 , J. G. Brooks (1♀, ANIC); Bramston Beach Rd. on Tower Rd. (17°23.136’S, 145°59.596’E), 93m, 4.ix.2018 , D. C. Rentz (1♀, ANIC); Bunya Mts. , 12.ii.1940 , N. Geary (1♀, AMS); Cairns, Macleay Museum (1♀, ANIC); Cairns, E. W. Ferguson colln. (2♀♀, ANIC); Cairns , xii.1970 , MV light, R. Hardie (1♀, ANIC); Cape Tribulation , 29.xii.1980 , MV lamp, G. & A. Daniels (2♁♁, AMS); Cape Tribulation Area (16°03’S to 16°05’S, 145°28’E), 1–11.v.1992 , J. F. Lawrence (1♁, ANIC); Cardstone , 15.xi.1966 , J. G. Brooks (1♀, ANIC); Cardstone , v.1972 , D. Perkins (1♀, ANIC); Claudie River, near Mount Lamond , 1.1972, at light, D, K, McAlpine, G. Holloway (6♀♀, AMS); Coo-rumba, 11.1983, 1985 , R. H. Mudder (4♀♀, AMS); Crystal Cascades, Cairns , 8.ix.1977 , I. F. B. Common (1♀, ANIC); Cuningham’s Gap , 26.xi.1964 , H. A. Rose (1♀, QMB); Daintree , 12 km S (16°19’S, 145°24’E), 22.xi.1981 , J. Balderson (1♁, ANIC); Forty Mile Scrub Nat. Park (18°06’S, 144°52’E), 2–3.iii.1988 , A. A. Calder (1♁, ANIC); Freedom Cunty, nr. Kuranda , 14.xii.1954 , R. De Keyzer (1♀, ASCU); Gap Creek (15°51’S, 145°20’E), 20.xi.1992 , at light, A. Calder, P. Zborowski (1♁, ANIC); Gap Creek , 5 km ESE Mt. Finnigan (15°50’S, 145°20’E), 13– 16.v.1981 , at light, I. D. Naumann (1♀, ANIC); Gap Creek , 5 mi. N of Bloomfield River, 100’, 8–9.v.1970 , G. B. Monteith (1♀, QMB); Gap Creek, Mt. Finlayson Ra., nr. Bloomfield , 2.i.1981 , MV lamp, G. & A. Daniels (2♀♀, AMS); Gordonvale , 13 km SwbyW (17°09’S, 145°41’E), 16.xi.1981 , J. Balderson (2♀♀, ANIC); Hutchinson Creek, near Daintree , R., 7.i.1967 , MV lamp, D, K, McAlpine, G. Holloway (1♀, AMS); Innisfail, F. H. Taylor, Hylecoetus vigilans, H. J. Carter det. (2♀♀, ANIC); Innisfail , 21.x.1925 , C. E. Simms (1♀, AMS); Innisfail , 4.5 mi. E, 50 ft., rainforest, 4.xi.1966 , at light, E. B. Britton (1♀, ANIC); Intake, nr. Redlynch , 30.xi.1967 , R. Dobson (2♀♀, ANIC); Iron Range , v.1961 , J. M. (1♀, ANIC); Iron Range , 9–12.iv.1964 , I. F. B. Common, M. S. Upton (1♁, 1♀, ANIC); Iron Range , 2.ii.1972 . P. Zborowski (1♀, ANIC); Iron Range , 9.i.1972 , M. Moulds (1♀, ANIC); Iron Range, vi–vii,1981, D. W. Frith (1♀, ANIC); Iron Range Nat. Park, Cooks Hut , 1 km N Lammon Hill (12°43’S, 143°18’E), 17.vii.1988 , at light, T. A. Weir, A. A. Calder (1♁, 2♀♀, ANIC); Iron Range, Cape York Pen. , 28.iv– 4.v.1968 , G. Monteith (1♀, QMB); Iron Range, West Claudie River , 6.xii.1985 , D. Yeates (1♀, QMB); Jubilee Rd. , 4.5 mi. NE Innisfail, 4.xi.1966 , at light, E. B. Britton (1♁, ANIC); Julatten , xii.1996 – ii.1997 , at light, B. P. Moore (6♁♁, 3♀♀. ANIC); Julatten , 3.xii.2004 , at light, B. P. Moore (1♀, ANIC); Kuranda , 12.i.1972 , P. Zborowski (1♀, ANIC); Kuranda , 2.ii.1982 , W. N. B. Quick (1♀, ANIC); Kuranda, 19 Butler Drive (top of range), 335m, X.2003 – II.2003 - 2007, 2010, 2015–2018, D. C. F. Rentz (13♁♁, 86♀♀, ANIC); Kuranda, 19 Butler Drive (top of range), 335m, 1–15.xi.2015 , D. C. F. Rentz (1♁, 2♀♀, AMS); Kuranda , 1 mi. E, 11.iii.1964 , I. F. B. Common, M. S. Upton (1♀, ANIC); Kuranda , 1 mi. N, 23.iv.1969 , I. F. B. Common, M. S. Upton (2♀♀, ANIC); Kuranda , 13 km NW, 6– 7.xii.1982 , J. T. Doyen (1♁, 3♀♀, ANIC); Kuranda Range St. For. , 10.i.1967 , MV lamp, D, K, McAlpine, G. Hol-loway (2♁♁, AMS); Lake Barrine, Atherton Tableland (17.15’S, 145.38’E), 13.xii.1985 , J. Balderson (1♀, ANIC); Lake Eacham (17°17’S, 145°38’E), 23.xi.1998 , at light, A. Calder (3♁♁, ANIC); Lake Eacham Nat. Park , 2– 5.ii.1998 , T. A. Weir (1♁, ANIC); McLeod River, 14 km WbyN Mt. Carbine (16°30’S, 145°00’E), 23.xi.1981 , J. Balderson (2♀♀, ANIC); Mission Beach , 1–5.x.1981 , G. O’Reilly (1♁, 4♀♀, ANIC); Mission Beach, ix–x.1991, at light, G. O’Reilly (1♁, 5♀♀, AMS); Mission Beach , ix.1993 , at light, G. O’Reilly (1♁, AMS); Mission Beach , 20.ix.1994 , at light, G. O’Reilly (1♁, AMS); Moses Creek, 4 km NbyE Mt. Finnigan (15°47’S, 145°17’E), 14– 16.x.1980 , T. Weir (1♀, ANIC); Mossman , 3 mi. W, 14.iii.1964 , I. F. B. Common, M. S. Upton (1♀, ANIC); Moss- man Gorge , 300 ft., 31.x.1966 , at light, E. B. Britton (2♁♁, 3♀♀, ANIC); Mt. Glorious , 16.i.1972 . G. Monteith (1♀, QMB); Mt. Glorious , 14.xi.1986 , flight intercept/trough trap, T. Hiller (1♀, ANIC); Mt. Hypipamee N.P. (17°26’S, 145°29’E), closed forest, 9–14.xi.1998 , at light, T. A. Weir (1♁, ANIC); Mt. Hypipamee N.P. (17°26’S, 145°29’E), 18–26.xi.1998 , A. Calder (4♁♁, 5♀♀, ANIC); Mt. Lamond, 4 mi. NE, Iron Range , 3.i.1972 , MV light, D, K, McAl-pine, G. Holloway (1♀, AMS); Mt. Misery , SW of Cooktown (15.52’S, 145.13’E), 867m, 20.i.1994 , mv lamp, G. & A. Daniels, R. Eastwood (1♀, QMB); Mt. Tambourine , 17.xii.1925 , A. Musgrave (1♀, AMS); Mt. Tennison Woods, via Mt. Glorious , 17.iii.1975 , rainforest, at light. K. J. Lambkin (1♀, QMB); Mt. Tozer , 2 km ENE (12°44’S, 143°13’E), 1.vii.1986 , at MV light, J. C. Cardale (1♁, ANIC); Mt. Tozer , 9 km ENE (12°43’S, 143°18’E), 5– 10.vii.1986 , at light, T. Weir, A. Calder, J. Cardale (2♁♁, 3♀♀, ANIC); Mt. Tozer , 11 km ENE (12°43’S, 143°18’E), 11–16.vii.1986 , at light, T. Weir, A. Calder (1♁, 1♀, ANIC); Mulgrave River , 4 mi W Gordonvale, 29.xii.1966 , MV lamp, D. K. McAlpine, G. Holloway (1♁, AMS); Nat. Park, H. Illidge (1♀, QMB); Palmerston Nat. Park, Tully-Cairns powerline, 1000 ft., rainforest, 5–6.xi.1966 , E. B. Britton (3♀♀, ANIC); Palm Grove Nat. Park (Palm Grove Section of Tamborine National Park), Tamborine Mtn. , 24–29.x.1993 , under bark, in rotten wood, S. A. Ślipiński, J. F. Lawrence 1♁, 1♀, ANIC); Paluma, 6 km NW on Paluma Dam Rd. (18°59’S, 146°09’E), rainforest, 14.i.1970 , at light, E. Britton (1♀, ANIC); Paluma ( Mt. Spec ), 5.i.1964 , MV light, J. G. Brooks (1♀, AMS, K517765 ); Polly Creek, Garradunga (17°458’S, 146°021’E), 15.i.2010 , MV light, J. Hasenpusch (1♀, ANIC); Shiptons Flat (15°47’S, 145°14’E), 17–19.x.1980 , T. Weir (2♁♁, ANIC); South Johnstone , 15 km WNW, 15.xi.1985 & x.1986 , light trap, Fay & Halfpapp (1♁, 2♀♀, ANIC); Tamborine Mtn. , i.1940 , H. Hacker (1♀, QDAF); Tinaroo Lake , west short, 2500 ft., 7.xi.1966 , at light, E. B. Britton 3♀♀, ANIC); Townsville, F. H. Taylor (1♀, ANIC); Tully Falls , 13.i.1972 , P. Zborowski (1♀, AMS); Tully Gorge Nat. Park (17°45’S, 145°32’E), 1.iii.1988 , at light, A. A. Calder (1♁, ANIC); Upper Currumbin , 10.xi.1931 (1♀, QMB) ; Upper Mulgrave River , 30.iv.1970 , G. B. Monteith (1♀, QMB); West Claudie River , 25.vi.1982 , mv lamp, G. Daniels, M. A. Schneider (1♀, QMB); West Claudie River, Iron Raqnge , 30.ix.1974 M. S. Mounds (1♀, AMS); Whitfield Range Timber Road, c 13–24 km frm Cairns (16°53’S, 145°44’E), 1600–2200 ft., various dates, 1970–1972, at light, J. G. Brooks, J. A. G. Brooks (4♁♁, 54♀♀, ANIC); Windsor Tableland , NE of Mossman (16.12’51”S, 145.04’09”E), site 1, 810m, 4.i.1984 , mv lamp, G. & A. Daniels, R. Eastwood (3♀♀, QMB) . NSW: Cherry Tree St. For., Pennefathers Rdo. Via Casino , 10.ii.1997 , MV light, S. Watkins (2♀♀, ANIC); Illawarra, Kiama, Hylecoetus pervagus, Macleay Mus. (2♀♀, ANIC); Lord Howe Island, on banyan, A. M. Lea, Hylecoetus pervagus Oll. (1♀, ANIC, 1♀, AMS); Lord Howe I., 1.i.1991 , in light trap, R. Schwartz (1♀, ASCU); Lord Howe Is. 15.xii.1992 , at UV light, G. R. Brown (1♁, ASCU); Lord Howe Island, Old AM Collection (1♀, AMS); Lord Howe Island, Research Station (31°31’37”S, 159°3’58”E), 12.ii.2001 , S. J. Fellenberg (1♀, AMS); Nighcap Nat. Park , Terania Ck., N of Lismore, 15–16.xi.1988 , UV light, G. Williams (1♀, AMS); Tooloom Scrub, West of Urbenville , 14.xii.1979 , at light in rainforest, G. & T. Williams (1♀, ANIC); Tooloom Plateau, via Urbenville , 16–17.xii.1972 , at light, G. B. Monteith (4♀♀, QMB); Upper Allyn River , 800 ft., 6.ii.1961 , I. F. B. Common, M. S. Upton (1♁, 1♀, ANIC). Wilson River F . R., NE of Wauchope , 5–6.xii.1988 , G. Williams (1♀, AMS, K517750 ) .

Distribution. Widely distributed in NSW and QLD.

Biology. Adults are usually collected at lights, but some have been taken in flight intercept traps. Specimens from Lord Howe Island were associated with banyan ( Ficus macrophylla Desf. Ex Pers. f. columnaris). Adults, pupae and larvae were collected in white rotted wood of a piccabeen palm ( Archontophoenix cunninghamiana (H. Wendl.) H. Wendl. & Drude ) in what is now the Palm Grove Section of Tamborine Mountain National Park, QLD.