Anthobium gaoligongshanense Shavrin & Smetana, 2019

Shavrin, Alexey V. & Smetana, Aleš, 2019, A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). III. Consanguineum, crassum and reflexum groups, and an additional species of the fusculum group, Zootaxa 4688 (4), pp. 451-482 : 476-477

Anthobium gaoligongshanense Shavrin & Smetana , sp.n.

( Figs. 68 , 75 , 85–87 )

Type material examined: Holotype ♂ [dissected]: ‘ CHINA: Yunnan [ CH 07-22], Nujiang | Lisu Aut. Pref., Gaoli- gong Shan, valley 21 | km W Gongshan, 3320 m, 27°47’03’’N, | 98°27’39’’E, moss, alder, bamboo, | Rhodod [endron], sifted, 6. VI .2007, M. Schülke’ <rectangular label, printed>, ‘ HOLOTYPE | Anthobium | gaoligongshanense sp.n. | Shavrin A.V. & Smetana A. 2018’ <red rectangulat label, printed> ( CSC) .

Paratype ♀: same data as the holotype, with additional red rectangular printed label ‘ PARATYPE | Anthobium | gaoligongshanense sp.n. | Shavrin A.V. & Smetana A. 2018’ ( CSC) .

Description. Measurements (n=2): HW: 0.75–0.79, HL: 0.42–0.45; AL(holotype): 1.32; OL: 0.15–0.16; PL: 0.62; PW: 1.25–1.35; ESL: 1.17–1.22; EW: 1.57; AW: 1.38; MTbL(holotype): 0.67; MTrL(holotype): 0.30 (MTrL 1–4: 0.15; MTrL 5: 0.15); AedL: 0.65; TL: 3.40(holotype)–3.54.

Body yellow-brown, with somewhat darker apical portion of head and antennomeres 8–11; mouthparts, antennomeres 1–7 and legs yellow. Forebody without meshes between punctures; abdomen with very indistinct isodiametric microsculpture. Head with irregular, fine and deep punctation, slightly denser on infraorbital ridges, with paired impunctated transverse portions between middle impression; neck with moderately large and deep, sparse punctation; pronotum with irregular punctation, finer in middle, larger, markedly sparser and deeper on lateral portions, with moderately wide transverse impunctated area on mediobasal third; scutellum with several fine punctures; punctation of elytra moderately sparse, large and deep, denser and finer on prescutellar area, each elytron forming six vague longitudinal rows of punctures; abdominal tergites with indistinct, dense and fine punctation. Habitus as in Fig. 75 .

Head 1.7 times as wide as long; middle portion of head slightly elevated, with wide, triangular impression at level of eyes between distinct and moderately deep grooves, reaching posterior margin of eyes; surface between punctures on infraorbital ridges irregularly and slightly elevated; postocular ridge distinct, acute. Distance between ocelli about one anf half times as long as distance between ocellus and posterior margin of eye. Length × width of antennomeres (holotype): 1: 0.20 × 0.08; 2–3: 0.11 × 0.05; 4: 0.10 × 0.06; 5–8: 0.11 × 0.06; 9–10: 0.10 × 0.07; 11: 0.16 × 0.07.

Pronotum twice as wide as long, 1.6–1.7 times as wide as head, widest about middle, evenly narrowed both postreriad and anteriad; posterior angles obtuse, without posterolateral emargination; edges of lateral margins of pronotum with irregular, small crenulation; middle elevated portion with wide and moderately deep longitudinal impression in apical half and indistinct, semicircular wide impression in mediobasal third; lateral portions very wide and markedly explanate.

Elytra moderately short, markedly wider than long, 1.8–1.9 times as long as pronotum, with straight apical margin, distinctly widened posteriad, reaching middle part of abdominal tergite IV; surface of each elytron with indistinct, five longitudinal elevation between punctures in middle.

Abdomen without visible tomentose wing-folding spots in middle of tergite V.

Male. Protarsomeres 1–4 slightly widened. Apical margins of abdominal tergite VIII and sternite VIII slightly emarginated.Aedeagus ( Fig. 85 ) with very wide basal portion, gradually narrowed toward truncated apex of median lobe; parameres rather short and wide, slightly asymmetrical, distinctly exceeding apex of median lobe, with small apical and preapical setae; internal sac very long, wide and spirally rolled in basal portion. Aedeagus laterally as in Fig. 86 .

Female. Protarsomeres 1–4 moderately narrow. Apical margin of abdominal tergite VIII somewhat straight. Apical margin of abdominal sternite VIII rounded.

Comparative notes. Anthobium gaoligongshanense sp.n. differs from remaining species of the crassum -group by the the elytra less than twice as long as pronotum, and by the external and internal structures of the aedeagus.

Distribution. The species is known only from the type locality in Gaoligong Shan range in Yunnan, China ( Fig. 86 ).

Bionomics. Specimens were taken by sifting mosses in mixed forest with alder, bamboo and Rhododendron (locality CH 07-22: Fig. 68 ).

Etymology. The specific epithet is the Latinized adjective derived from the name of the Gaoligong Shan and the added suffix – ense, denoting the place of the origin.

Figures

FIGURES 65–68. Type locality of Anthobium crassum (65–66), A. crenulatum (67) and A. gaoligongshanense (68) in Yunnan, China (photograph: M. Schülke).

FIGURES 73–77. Habitus of Anthobium: 73—A. reflexum (Kazakhstan) 74—A. deplanatum (holotype), 75—A. gaoligongshanense (holotype), 76—A. kashmiricum, 77—A. transversale (holotype). Scale bar: 1.0 mm.

FIGURES 83–86. Aedeagus of Anthobium deplanatum (83–84: paratype) and A. gaoligongshanense (85–86: holotype): 83, 85—parameral view, 84, 86—lateral view. Scale bar: 0.1 mm.

FIGURE 87. Distribution of Anthobium deplanatum (black circles), A. gaoligongshanense (black square) and A. transversale (black triangle) in the south-western China.