Agalenocosa gamas, Piacentini, Luis N., 2014

Piacentini, Luis N., 2014, A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae), Zootaxa 3790 (1), pp. 1-35 : 30-31

Agalenocosa gamas sp. nov.

( Figs 23 , 24 h, 27)

Type material. Female holotype from Argentina, Santa Fe, Las Gamas , 20 km W Vera, Ramírez, M. J. & Faivovich, J., 27.x.1994 (MACN-Ar 22732) and two female paratypes with three eggsacs with the same location and data as the holotype (MACN-Ar 30548).

Etymology. The specific epithet is a noun in apposition taken from the type locality.

Diagnosis. Females of A. gamas sp. nov. resemblesthose of A. gentilis by the coloration pattern ( Figs 17 , 23 ), but can be recognised by the small lightly sclerotised area that surrounds the copulatory openings and the relatively long, nearly parallel, anteriorly directed stalks of the spermathecae and the small, rounded heads ( Fig. 23 d).

Description. Male. Unknown.

Female. Holotype (MACN-Ar 22732). Carapace brown with diffuse light brown median band and with a Yshaped dark mark that extends from the fovea to the PLE, pale submarginal bands ( Figs 23 a–c). The border of the carapace clothed with white pubescence ( Figs 23 a–c). Sternum yellow, sparsely covered with brown bristles, more abundant on the margins. Labium dark grey. Chelicerae lighter than labium, light brown, covered with brown bristles. Dorsum of abdomen brown with irregular pale dots, longitudinal lines of white pubescence on its lateral sides, and two lines of bunches of glistening setae on its posterior part ( Figs 23 a–c). Venter pale brown. Legs pale brown without annulations. Leg formula IV> I> II> III.

Spination: femur I p 0-0-d1 d 1-1-1, II p d1-0-d1 d 1-1-1 r d1-0-d1, III p d1-0-d1 d 1-1-1 r d1-0-d1, IV p d1- 0-d1 d 1-1-1 r 0-0-d1; patella I d 0- 1 II p 1 d 1-1, III p 1 d 1-1 r 1, IV p 1 d 1-1 r 1; tibia I v 2-2-0, II p d1-0-d1 v 2-2 -0, III p d1-0-1 d 1-1-0 r 0-1- 1 v p1-p1-2ap, IV p 0-d1-d1 d 1-0-1 r 0-1- 1 v p1-p1-2ap; metatarsus I v 2-2-2ap, II p 1-0-2ap r 0-0-1ap v 2-2 -1ap, III p d1-d1-2ap r d1-d1-2ap v 2-2 -1ap, IV p d1-1-1 d 0-0-2 r d1-1-1 v 2-2 -1ap.

Epigynal plate without pockets or atria, copulatory openings with a lightly sclerotised area posteriorly ( Fig. 23 b). Vulva: head of spermatheca rounded, stalk of spermathecae nearly parallel and longer than the head ( Fig. 23 e). Vulval chamber small and rounded, next to the base of spermathecae in dorsal view ( Fig. 23 e).

Measurements. Female, MACN-Ar 22732: TL 5.32, CL 2.60, CW 1.87, CH 0.93, AL 2.53. Eyes: AME 0.12, ALE 0.08, PME 0.18, PLE 0.15. Row of eyes: AER 0.48, PME 0.50, PLE 0.72. Sternum (length/width) 1.42/1.08. Labium (length/width) 0.40/0.38. Legs: length of segments (femur + patella/tibia + metatarsus + tarsus = total length): I 2.07 + 2.47 + 1.67 + 0.93 = 7.14, II 1.87 + 2.13 + 1.33 + 0.80 = 6.13, III 1.47 + 2.07 + 1.47 + 0.80 = 5.81, IV 2.53 + 3.20 + 2.67 + 1.07 = 9.47.

Variation. Females (range, mean ± s.d.): TL 4.39 – 5.32, 4.97 ± 0.50; CL 2.00 – 2.60, 2.40 ± 0.35; CW 1.40 – 1.87, 1.65 ± 0.24 n = 3.

Distribution. Only known from the type locality, in northern Santa Fe Province ( Fig. 27 )

Figures

FIGURE 17. Agalenocosa gentilis Mello-Leitão. a – c habitus, a dorsal, b lateral, c ventral; d epigyne ventral; e vulva, cleared, dorsal. Scale bars, a – c 2.00 mm, d 0.20 mm, e 0.10 mm. Abbreviations: BS Base of spermatheca, EP epigynal plate, FD fertilization ducts, HS head of the spermatheca, SS stalk of spermatheca.

FIGURE 23. Agalenocosa gamas sp. nov.. a – c habitus, a dorsal, b lateral, c ventral; d epigyne ventral; e vulva, cleared, dorsal. Scale bars, a – c 2.00 mm, d 0.20 mm, e 0.10 mm. Abbreviations: BS Base of spermatheca, DP Dictynoid pore, EP epigynal plate, FD fertilization ducts, HS head of the spermatheca, SS stalk of spermatheca.

FIGURE 24. Female, habitus frontal, a A. velox comb. nov.; b A. tricuspidata comb. nov.; c A. luteonigra (Mello-Leitão); d A. punctata Mello-Leitão; e A. gentilis Mello-Leitão; f A. grismadoi sp. nov.; g A. pirity sp. nov.; h A. gamas sp. nov.. Scale bars, a, c, e 0.50 mm, b, d, g 1.00 mm.

FIGURE 27. Geographic distribution of A. grismadoi sp. nov. (red stars), A. pirity sp. nov. (blue squares) and A. gamas sp. nov. (black circle). The asterisk inside of the mark point that is only the province as locality in the label.