Scotaena reversa Carnimeo, 2018

Carnimeo, Fernando Henrique & Noll, Fernando Barbosa, 2018, On the dumping ground genus Scotaena Klug, 1810 (Hymenoptera: Tiphiidae: Thynninae): Phylogeny, taxonomy and geographic distribution, Zootaxa 4399 (4), pp. 451-490 : 460-461

Scotaena reversa Carnimeo sp. nov.

( Figs 27–32 , 91, 98, 100, 105 )

Holotype. BRAZIL: Serra da Bocaina, São José do Barreiro , São Paulo, 11/xi/1964, 1♂ (Kovacs) ( AMNH) . Paratype, BRAZIL: Parque Nacional da Serra dos Órgãos, Rio de Janeiro, 04/i/2014 – 01/ii/2014, 3♂ (R.F. Monteiro and eq. col.); Parque Nacional do Itatiaia , Rio de Janeiro, 27/ii/2014, 1♂ (R.F. Monteiro and eq. col.) ( UFRJ) .

Male description. Structure: Supra-antennal plate with slightly carinate margins. A slight elevation between the plates; clypeal apical emargination as deep as its sublateral margins. Base of clypeal emargination subtriangular, what origins a slight longitudinal bevel raised on the medial region of the clypeus; wings hyaline, ferruginous on the anterior margins, stigma and marginal cell. The anterior half of the forewing slightly castaneus; scutellum raised and pointed at the top; Dorsal surface of the propodeum with dense and coalescent punctures, laterals striated at the base; mesepisternal suture strongly raised apically, the lamellae narrow and acute; first metasomal tergite longer than wide. Smooth transverse fasciae on apical margins of tergites almost absent, very short and covered by a dense row of setae; epipygium sides rugose by elongate punctures, smooth centrally; hypopygium subquadrate, rounded laterally.

Genital capsule: Dorsal surface of the basal ring more than 2x longer than its ventral surface. With a longitudinal line medially in dorsal view; gonocoxite tapering towards the apex, with more than half the gonostylus extension. The apex rounded and almost contiguous, slightly separated medially, with a longitudinal line dorsally; aedeagus longer than the gonocoxite and shortest than the gonostylus. Expanded laterally in two longitudinal lamellae, curved ventrally and with a small apical lobe; gonostylus slightly wider at the base, narrowing medially and broadening again before the apex. With golden pubescence on the half apical region. The apex slightly rounded, triangularly curved ventrally.

Coloration. body predominantly black, except for the following castaneus markings: flagellomeres V–VII. A large spot on the scutellum dorsal surface. The metanotum centrally. The legs except for the outer surface of the femora. Diffuse spots on the sternites II–IV.

Female: unknown.

Etymology. In Latin reversus means “returned”, referring to the lack of the strong carina on the margins of the supra-antennal plate, which is a diagnostic character of the Scotaena but seems to be reverted or at least less pronounced in this species.

Distribution. The known records of Scotaena reversa are restricted to the coastal region of the Brazilian Atlantic Forest.

Figures

FIGURES 27–32. Scotaena reversa, holotype male. 27, frontal head; 28, dorsal habitus; 29, lateral habitus. Genital capsule: 30, dorsal view; 31, ventral view; 32, lateral view.Scale bar = 1mm.

FIGURES 90–105. Scotaena characters. 90, 96, 99: S. trifasciata, 91, 98, 100, 105 S. reversa; 92, 94, 97, 101, 103, 104: S. vetusta; 93, 95, 102: S. horni. Scale bar = 1mm.