Zonitoschema melanarthra ( Fairmaire, 1894 ) Bologna & B & B, 2018

Bologna, Marco A., B, Ladislav Černý & B, Ahmed Zubair, 2018, Meloidae (Coleoptera) of Pakistan and Kashmir with the description of three new species, new faunistic and taxonomic records, and a zoogeographic analysis, Turkish Journal of Zoology 42 (6), pp. 637-660 : 641-646

Zonitoschema melanarthra ( Fairmaire, 1894) comb. nov. ( Figure 4 )

New records. Pakistan: 2 exx., NWFP, N of Peshawar , Tangi village, 27.vii.1998, attracted by light, L. Černý and L. Čížek ( CC, CB) .

Distribution. Until now this species was recorded only from India; new species record for Pakistan. It was described from “Barway” in the Jharkhand State and afterwards quoted by Fairmaire (1896) from Pune (as Poona), in the Maharashtra State.

Remarks. This species was described as Zonitomorpha , and we are proposing for it a comb. nov. Its identification was confirmed after the examination of the holotype (MNHN) and a second specimen from Pune (MNHN). Specimens from Tangi differ slightly from the holotype in the shape of pronotum, which is slightly more rounded in the fore sides, but this character could be variable as the specimen from Pune shows a wider and more rounded pronotum than the holotype.

Zonitoschema has a Paleotropical distribution, being distributed in the Afrotropical and Oriental regions, with a partial spread in the southern Palearctic ( Batelka and Bologna, 2014) and Australasian regions ( Bologna et al., 2013). Species relationships are not clarified and the genus requires revision. A key to the Saharo-Sindian species was published by Batelka and Bologna (2014), and keys to the Afrotropical and Asian species are in preparation (Bologna et al., unpublished).

This is one of the few Palearctic and Oriental species of Zonitoschema , some of which are undescribed, with the apical fifth of the elytra black (Bologna and Pan, unpublished). Among them, Z. melanarthra is characterized by the black antennae except antennomere I yellow-orange, and the black legs with the basal 2/3 of the tibiae orange-red. It is very similar to Zonitoschema angustithorax (Pic, 1912) (comb. nov., described as Zonitomorpha ) from E China (Qingdao), which has a narrow pronotum with parallel sides.

Zonitoschema straminea ( Fairmaire, 1894) comb. nov.

New records. Pakistan: 1 ex., Islamabad , sect. E, 7 33°43′N 73°03′E, 600 m, 20.vi.2011, G. Sabatinelli, at light ( CB, in alcohol 95%) .

Distribution. Until now recorded only from eastern India (Jharkhand State); new species record for Pakistan.

Remarks. We refer a single examined specimen to straminea ; the identification was confirmed after the examination of the holotype (MNHN) and a possible paratype from “Barway” (MNHN, labeled respectively: (a) “Barway. P. Cardon; Zonitis straminea Fairm. n. sp. ”; (b) “ Zonitis straminea Fairm. Bengala ; Barway. P. Cardon”).

This species was described as Zonitis , and we are proposing for it a comb. nov. It is characterized by the almost completely orange body with antennal segments III–XI and the last two tarsomeres on each leg including the claw black. The pronotum is slightly widened at middle. No other Old World species has a similar color combination ( Batelka and Bologna, 2014; Bologna and Pan, unpublished).

Hashmi and Tashfeen (1992) recorded, without comment, an undetermined species Zonitoschema from Pakistan.

Subfamily Meloinae

Tribe Lyttini

Lytta (Lytta) antennalis (Marseul, 1873)

Distribution. This species, erroneously cited from Japan by Bologna (2008), due to erroneous identification, actually is distributed from northern Pakistan and northern India to the Himalayan region and Xinjiang.

A single specimen from Rawalpindi (13/ 17.vii.1995, J. Klir; NHMW, late J. Probst collection) possibly refers to a new species of the nominotypical subgenus, close to L. antennalis .

Lytta (Lytta) flavipennis (Motschuslky, 1860)

Distribution. This species is distributed in India and Himalaya, and recorded from Pakistan by Hashmi and Tashfeen (1992). We did not examine specimens from this country, but from the Indian N Punjab, close to Kashmir [2 exx., Kangra valley-Yol, 1300 m, iv–vii.1944, C. Lomi (CB, MRSN)].

Lytta (Lytta) kashmirensis Bologna, 1983

Distribution. Recorded only from the Indian Kashmir [Urdukas, 3900 m, (CB)] ( Bologna, 1983, 2008; Bologna and Osella, 1989).

Remarks. In CK (see http://www.meloidae.com/cs/ obrazky/370/) is preserved one specimen from Pakistan [Northern Areas, 5 km S Babusar, 3400 m, 35.11N 74.03E (30 km south Chilas) 19–21.VII.2000], which is identified as L. kashmirensis . However, this specimen differs from L. kashmirensis in that the elytral apex is yellow and smooth, the elytral punctures less deep and dense, and those on the head and pronotum sparser and tiny. This specimen possibly refers to another undescribed species of the nominotypical subgenus.

Lytta (Asiolytta) limbata (Kollar and Redtenbacher, 1842)

New records. Pakistan: 1 ex., Dir , Lawarac Pass, 2700–3300 m, 17–19.viii.1981, Heinz leg. ( NMPC, from Dvorak’s collection) .

Distribution. This species was recorded from Pakistan by Hashmi and Tashfeen (1992). The nominotypical form, described from Kashmir, is recorded from Afghanistan, Kashmir, northern India (Uttar Pradesh), and Nepal (Bologna, 2008); the subsp. afghanica Kaszab, 1981 , described from Afghanistan, is distributed also in Pakistan (Lulu Sar: Kaszab, 1981) but it was not reported from this country by Bologna (2008).

Remarks. The taxonomy of the subgenus Asiolytta is poorly understood. We briefly examined at HNHM types of the subsp. afghanica , which is easily separated from the nominotypical form based on male genitalia and protarsomeres (see also Kaszab, 1981), and we suspect that it could represent a distinct species. Contrary to our opinion, Axentjev (1987) synonymized both L. limbata afghanica and L. poeciloptera Semenov, 1893 with L. limbata .

Lytta (Asiolytta) discipennis ( Fairmaire, 1891)

Distribution. Apparently endemic to Kashmir ( Fairmaire, 1891). It was recorded from there by Anand (1989), and we know of specimens from the Indian Punjab, Kangra valley (CB).

Remarks. We examined the male holotype of this species (MNHN, labeled “ Cantharis discipennis Fairm. Kashmir ”). A distinguishing feature of this species is the modified male protarsomere I, which is narrowed basally and has a triangularly pointed ventral lamina.

Lytta (Eolytta) flavoangulata ( Fairmaire, 1891)

New records. Pakistan: 1 ex., northern area, Astor , Rama rest-house, 35.3565°N - 14.8080°E, 3170 m, 7.vi.2007, G. Carpaneto ( CB) .

Distribution. New species record for Pakistan. Until now it has been recorded from Kashmir ( Fairmaire, 1891; Borchmann, 1935; Anand, 1989) and Afghanistan ( Kaszab, 1981; see Bologna, 2008). The species is new also for India, Himanchal Pradesh [Mandi, 3500 m, 1.vii.2003 (CB, MSNG)].

Remarks. This species was referred by Kaszab (1962) to the subgenus Poreospasta Selander, 1960 , but recently Shapovalov (2016) placed this and other Central Asiatic species in the new subgenus Eolytta . At present we are unable to evaluate differences between Eolytta and Poreospasta , a subgenus that could be restricted to North America.

We examined the holotype of this species (MNHN, labeled: “Kashmir”; “ Cantharis flavoangulata Fairm. Kashmir ”). The species is recognized by its primarily black body with metallic green or black elytra, and distinctly trapezoidal pronotum, black in color except for its lateral margins of pronotal disk broadly marked with orange.

Lytta (Mesolytta) vrendenburgi Kaszab, 1962

Distribution. Described from Nushki ( Kaszab, 1962), in Pakistan Baluchistan, erroneously considered as an Afghanistan locality by Bologna (2008). It also occurs in Uzbekistan (see below).

Remarks. Dvořak (1983) considered this species a synonym of L. (M.) coccinea (Ménétriés, 1849) , on the basis of specimens from Uzbekistan (Kyzyl-Kum, Kyzyl-Kyr, nr. Bukhara) that we also examined (CB). The synonymy was not accepted by Bologna (2008), because the black leg coloration is a constant character in vrendenburgi specimens from Uzbekistan and Pakistan (see Kaszab, 1962). In addition to this color feature, the shape of the head is different. In vrendenburgi the male temples are slightly widened and more convex than in coccinea . We have examined specimens clearly referable to coccinea from Uzbekistan (Minski, CB) and consequently vrendenburgi and coccinea are widely sympatric.

Cyaneolytta coerulea (Pfaff, 1834)

New records. Pakistan: 2 exx., Sindh, Mithi, Thar desert, 24.x.2015 ( FUUZM) ; 1 ex., Sindh, Tharparkar district, Islamkot , 1.ix.2016 ( FUUZM) ; 1 ex., Sindh, Tharparkar District, Nagarparkar desert, 25.ix.2016 ( FUUZM) .

Distribution. Oriental species, also marginally distributed in the Palearctic Region ( Pakistan, Sikkim, Uttar Pradesh: Bologna, 2008). Hashmi and Tashfeen (1992) recorded it from Pakistan as Epicauta acteon Dejean, and Anand (1980 , 1989) as Cyaneolytta acteon . It was also noted from Pakistan by Selander (1986a, 1986b). Two additional species have been cited in the nearby Indian state of Rajahsthan: C. rajasthanica Kaszab, 1978 and C. indica Anand, 1980 ; both are possibly synonyms of C. coerulea and their validity need confirmation after the comparative examination of types.

Remarks. The complex synonymy of this species was clarified by Selander (1986a).

Lydomorphus (Lydomorphus) tenuicollis (Pallas, 1798)

New records. Pakistan: 2 exx., Baluchistan, Nushki, 8.x.2016 ( CB; FUUZM); 1 ex., N Baluchistan, 40 km W of Kingri village, 23.vii.1998, attracted by light, L. Černý and L. Čížek ( CC); 1 ex., Punjab, Changa Manga forest , 70 km S from Lahore, 9–21.viii.1998 ( NHMW, late J. Probst collection); 15 exx., idem, 19–21.viii.1998; L. Černý and L. Čížek ( CC) .

Distribution. Sahelo-Arabic-Sindian element, widely distributed from Egypt and Sudan, through the Arabian Peninsula to southern Iran, Pakistan, and India. Recorded from Pakistan by Selander (1988), Anand (1989), and Bologna (2008).

Remarks. Usually recorded in the literature as ruficollis. Hashmi and Tashfeen (1992) erroneously cited this species as “ Epicauta tervicolis Pul. ” and “ Lytta tonvicollis Pall. ”.

Lydomorphus (Lydomorphus) angusticollis suturellus (Haag-Rutenberg, 1880)

New records. Pakistan: 2 exx., Baluchistan, Nushki, Kili Jamaldini , 22.vii.2012 ( FUUZM); 2 exx., idem (only Nushki), 25.v.2016 and 3 exx., 8.x.2016 ( CB) .

Distribution. Sahelo-Sindian element, widely distributed from Sudan and Horn of Africa, through the Arabian Peninsula and southern Iran to western India. Recorded from Pakistan (Baluchistan) by Kaszab (1968) and Selander (1988) (see Bologna, 2008). Hashmi and Tashfeen (1992) erroneously recorded this species as Epicauta angusticollis Koll.

Lydomorphus (Lydomorphus) pictus (Laporte de Castelnau, 1840)

New records. Pakistan: 1 ex., N Baluchistan, 40 km west of Kingri village , 23.vii.1998, L. Černý ( CC) ; 1 ex., Sindh, Mithi, Thar Desert , 24.x.2015 ( FUUZM) .

Distribution. Oriental species distributed in Pakistan and India (we know specimens from Rajasthan, Ajmer, CB). Recorded from Pakistan by Hashmi and Tashfeen (1992) as Epicauta picta , but not reported by Bologna (2008).

Sybaris praeustus (Redtenbacher, 1842)

New records. Pakistan: 1 ex., Sindh, Tharparkar district, Nagarparkar Desert , 25.ix.2016 ( FUUZM) ; 1 ex., Sindh, Tharparkar district, Islamkot , 2.x.2016 ( CB) .

Distribution. Recorded only from Kashmir and Uttar Pradesh, India (Bologna, 2008).

Remarks. The genus Sybaris is distributed through the sub-Saharan African, eastern Sind, and Oriental regions. It has never been studied comprehensively and needs revision. The Asiatic species, in particular, are difficult to distinguish.

Sybaris testaceus (Fabricius, 1792)

Distribution. Species recorded from Pakistan ( Hashmi and Tashfeen, 1992; Bologna, 2008), India, and Nepal (Bologna, 2008).

Sybaris tunicatus (Redtenbacher, 1842)

Distribution. Endemic to Kashmir ( Anand, 1989; Bologna, 2008).

Remarks. The validity of the species needs confirmation because it was based on the presence of an anterior black spot on the elytra, which could represent only a variation of the coloration of S. praeustus (Redtenbacher, 1842) .

Sybaris semivittatus (Redtenbacher, 1842)

Distribution. Recorded only from Kashmir and Himanchal Pradesh (Bologna, 2008).

Eolydus conspicuus (Waterhouse, 1889)

Distribution. Occurring from Iran to Afghanistan and Pakistan. Also recorded from “Turkestan” (Bologna, 2008) and Baluchistan ( Kaszab, 1956).

Tribe Epicautini

Epicauta (Epicauta) atkinsoni Kraatz, 1880

New records. Pakistan: 6 exx., Baluchistan, Nushki , 16.vii.2015 ( CB) ; 1 ex., idem, 8.x.2016 ( CB) ; 3 exx., Sindh, Tando Jam Univ. Agr., 14.v.2012 ( CB) .

Distribution. New species record for Pakistan; previously recorded from Iran (Baluchistan), Afghanistan, and India (Kaszab, 1952, 1958; Bologna, 2008).

Epicauta (Epicauta) hirtipes (Waterhouse, 1871

Distribution. Species widely distributed in Kazakhstan, Iran, Pakistan, India, Nepal (Bologna, 2008).

Epicauta (Epicauta) mannerheimi (Mäklin, 1875)

Distribution. This species is distributed in both Oriental and Palearctic regions (Buthan, Nepal, Uttar Pradesh), and some southern regions of China and Vietnam). Recorded from Pakistan by Anand (1989) and Hashmi and Tashfeen (1992), but not reported by Bologna (2008). It could be confused with other species such as E. rubriceps Kollar and Redtenbacher, 1842 .

Epicauta (Epicauta) nepalensis (Hope, 1831)

Distribution. Distributed in both Oriental and Palearctic regions ( Nepal, and some states of northern India; Bologna, 2008). Recorded from Pakistan by Hashmi and Tashfeen (1992), but not reported by Bologna (2008). It could be confused with E. rubriceps Kollar and Redtenbacher, 1842 .

Epicauta (Epicauta) rubriceps Kollar and Redtenbacher, 1842

New records. Pakistan: 2 exx., Kashmir , Azad, Kotli, 14.vii.2014 ( CB, FUUZM); 1 ex. , idem, 26.viii.2016 ( CB) .

Distribution. This species, described from Kashmir, is widely distributed in Pakistan, India, and western China (Bologna, 2008).

Remarks. The identification of specimens from Kotli, both females and damaged, needs confirmation and is questionable, after the study of male specimens.

Epicauta (Epicauta) quadraticollis Fairmaire, 1891

Distribution. Species poorly known in the literature, recorded from Pakistan by Anand (1989) and from Kashmir and northern India (Himanchal Pradesh) by Bologna (2008).

Psalydolytta villipes (Haag-Rutenberg, 1880)

New records. Pakistan: 3 exx., Kashmir , Azad, Kotli, 14.vii.2014 ( CB; FUUZM) .

Distribution. New species record for Pakistan; previously recorded only from India ( Kaszab, 1954; Anand, 1977). The genus Psalydolytta was never previously recorded from Pakistan or Kashmir.

Tribe Mylabrini

Mylabris (Mylabris) cfr. cernyi Pan and Bologna, 2014

New records. Pakistan: 6 exx., Gilgit-Baltistan, Dassu, S. Prepsl (CB, CC).

Distribution. Mediterranean Turkey, Israel and Palestine, Jordan. According to Pan and Bologna (2014) this Pakistan record is doubtful and needs to be corroborated.

Remarks. The Pakistan specimens examined could represent a new species. Molecular techniques should help to resolve this question.

Mylabris (Mylabris) quadripunctata (Linnaeus, 1761)

Distribution. This species was recorded from Pakistan by Iablokoff-Khnzorian (1983), but this identification is doubtful (see Pan and Bologna, 2014) and possibly refers to Mylabris cernyi .

Mylabris (Calydabris) cfr. mirzayani ( Kaszab, 1968)

New records. Pakistan: 1 ex., Baluchistan, Nushki , 12.vi.2012 ( FUUZM); 3 exx., Baluchistan, City Loralai ,

30.37N 68.6E, 1426 m a.s.l., 18.vi.2012 ( CB) ; 1 ex. idem, 1.vi.2013 ( FUUZM) ; 2 exx., Lower Dir, Tehsil, Samar Bagh , 17.vii.2012 ( CB) ; 3 exx., idem, 22.vii.2012 ( FUUZM) .

Remarks. The available specimens are all damaged and cannot be adequately identified. They are close to Mylabris (Calydabris) mirzayani because of the elytral pattern but some differences need a deep comparison with typical specimens of this species and others from Iran, Kerman province (CB), which probably represent a new species. The mesosternum has distinct setae at the posterior apex as in the nominotypical subgenus and in the subgenus Calydabris Kaszab, 1960 , but it is more similar to that of Calydabris because of the yellow coloration of the anterior sides and the posterior setae less dense. Although the ventral blade of the claws is not reduced as in most Calydabris , we know undescribed Iranian specimens possibly referable to M. mirzayani or to a distinct species, which have claws as in the Pakistan ones. The male genitalia are similar to those of Calydabris and differ from those of the nominotypical subgenus in that the parameres are slender and the aedeagus, also very slender at its apical third, has its two hooks distant from the apex. The identification of these specimens is appended, waiting for material in better condition.

Mylabris (Micrabris) afghanica Kaszab, 1953

New records. Pakistan: ca 90 exx., Northern Areas , Shandur Pass, 4200 m, 60 km ENE Chitral, 16/ 17.vii.1998, J. Kaláb ( CB; CC; CK) ; 6 exx., NWFP, Chitral distr. , Bumboret valley, Gumbak pass, 3000 m, 17.viii.1999, M. Šlachta ( CC) ; 1 ex., WFP, southwest Utrot , 2400 m, 35.28N - 72.26E, 26–28.vii.2000, J. Kaláb ( CC) ; 3 exx., N Pakistan, Diamir distr. , Nanga Parbat Mt. , Rialkhot valley, 3300 m, Fairy Meadows, 18.vii.1999, M. Šlachta ( CC) ; 3 exx., N Pakistan, Baltistan pr., Hushe valley, Nangbrok road, 3500 m, 29.vii.1999, M. Šlachta ( CC) .

Distribution. Central Asiatic element, endemic to the Hindu-Kush Mts., previously recorded from Afghanistan and Pakistan (Bologna, 2008).

Remarks. Specimens from Shandur Pass differ slightly from typical afghanica in the puncturation and need further study.

Mylabris (Micrabris) macilenta Marseul, 1873

New records. Pakistan: numerous exx., NWFP, Ayuba , Ghora distr., 75 km N from Islamabad, 1/ 16.vi.1998 ( CB) ; 2 exx., Northern Areas , Astor, Rama rest-house, 35.3569°N 74.8080°E, 3170 m, 7.6.2007, G. Carpaneto leg. ( CB) ; 1 ex., Baltistan , Mimimark, 3900 m, 29.vii.1996, U. Pessolano ( CB) ; 52 exx., Kashmir , Drass, 3330 m, 2.viii.1976, B.G. Osella ( CB; MSNV) ; more than 100 exx., NWFP, Ayubia- Ghora Dhaka , 75 km NE of Islamabad, 12–16.viii.1998, L. Černý and L. Čížek ( CC) .

Distribution. Himalayan element distributed from northern Pakistan to southeastern China, through northern India, Nepal, and Bhutan (Bologna, 2008). Already recorded from Kashmir ( Borchmann, 1935; Axentjev, 1987) and Pakistan (Saha, 1972; Hashmi and Tashfeen, 1992). We also know of specimens from N Punjab, close to Kashmir [1 ex., Kangra valley-Yol, 1300 m, iv–vii.1944, C. Lomi (MRSN)]. Records from the Afghanistan Hindukush ( Borchmann, 1935) refer, in part, to M. frolovi (see Kaszab, 1973).

Remarks. Axentjev (1987) erroneously synonymized M. afghanica with M. macilenta .

Mylabris (Micrabris) nuristanica nuristanica Kaszab, 1958

New records. Pakistan: 12 exx., NWFP, Chitral distr. , Bumboret valley, Broon village, 29.vii–6.viii.1998, L. Černý and L. Čížek ( CC); 1 ex. , Hindukush Mts. , Dir, 1700 m, 13–14.vii.1995, Jiří Klír ( CC) .

Distribution. Mylabris nuristanica is a polytypic species; the nominotypical subspecies was recorded from both Afghanistan and Pakistan.

Mylabris (Micrabris) nuristanica subalpestris Kaszab, 1958

New records. Pakistan: 15 exx., Baluchistan, Ziarat , 29–30.vi.1991 S. Prespl ( CB, CC); 1 ex. , NWFP, road Chitral-Mastuj , 85 km NE Chitral, 15.vii.1998, J. Kaláb ( CC) .

Distribution. The subsp. subalpestris Kaszab, 1958 was recorded only from Afghanistan but, according to the new records, it is present also in Pakistan. It was quoted from Pakistan by Hashmi and Tashfeen (1992) and from the locality of Chitral by Saha (1972a).

Remarks. Described by Kaszab (1958) in the subgenus Eumylabris , but correctly transferred to the subgenus Micrabris by Pardo Alcaide (1969).

Mylabris (Micrabris) waziristanica ( Kaszab, 1958)

Distribution. Distributed in Afghanistan and Pakistan ( Kaszab, 1958, 1981; Bologna, 2008; Bologna et al., 2008a). Erroneously recorded from northwestern India ( Kaszab, 1958; Anand 1989), but actually the type locality is in Pakistan.

Remarks. The species was erroneously described as Coryna Billberg, 1813 by Kaszab (1958) and compared to another species now included in the genus Actenodia Laporte de Castelnau, 1840 , and later referred to Mylabris (Micrabris) by Bologna et al. (2008a) after the examination of the holotype and one paratype from Waziristan (NHM).

Mylabris (Zitunabris) cfr. panjoensis Kaszab, 1953

New records. Pakistan: 1 ex., Lower Dir , Tehsil, Samar Bagh, 12.vii.2016 ( CB) .

Distribution. Previously recorded only from Afghanistan ( Kaszab, 1953); new species record for Pakistan.

Remarks. The taxonomy and phylogenetics of the subgenus Zitunabris Pardo Alcaide, 1968 is incompletely resolved, as confirmed by molecular data ( Salvi et al., 2018). M. panjoensis was described as an Afghanistan subspecies of M. syriaca Klug, 1845 , but, in our opinion, according to the description ( Kaszab, 1953), it could be a distinct species.

The Pakistan specimen slightly differs from the description that was based on Afghanistan material. The elytra are more opaque and with dense punctures, the mesosternum shape slightly differs, and the black pattern on the elytra is less undulate. This last character is more similar to the elytral pattern in M. atrofasciata Pic, 1921 , a species endemic to Iran. The Pakistan specimen differs from the latter by its subopaque elytra, less pronounced temples, and narrower pronotum; its pronotal disk also is less depressed anteriorly. Pending a revision of the subgenus, we prefer to tentatively assign this specimen to panjoensis .

Mylabris (Zitunabris) syriaca Klug, 1845

New records. Pakistan: 1 ex., Baluchistan, Loralai , 1.vi.2013 ( FUUZM) .

Distribution. The species is widely distributed from Turkey (and possibly Greece) to Levant, Iran, and Afghanistan. This is a new species record for Pakistan.

Remarks. Only one photo of the Loralai specimen was examined. It clearly shows the elytral pattern, pronotum shape, and body punctures, which correspond to this species.

Mylabris (Eumylabris) cincta Olivier, 1795

New records. Pakistan: 9 exx., W Baluchistan, Tump, 90 km west of Turbat , 13–15.iv.1993, S. Bečvář ( CC) ; 2 exx., western Baluchistan, Turbat , 8–19.iv.1993, S. Bečvář ( CC) ; 3 exx., western Baluchistan, Gwadar , 17–18.iv.1993, S. Bečvář ( CC) .

Distribution. Widely distributed from Morocco to Arabia, and from Greece and Macedonia, through southern Russia, Turkey, Levant, and the Caucasus, to Central Asia, Afghanistan, Pakistan, and northwestern India (Himanchal Pradesh, Uttar Pradesh) (Bologna, 2008).

Remarks. The subgenus Eumylabris Kuzin, 1954 needs taxonomic revision; species remain undescribed or confused, and some of the taxa could ultimately be synonymized.

Mylabris (Eumylabris) klapperichi klapperichi Kaszab, 1958

New records. Pakistan: 25 exx., NWFP, Chitral distr. , Kalash valleys, Bumburet valley, Broon village, 29.vii–6. viii.1998, L. Černý and L. Čížek ( CB, CC) .

Mylabris (Eumylabris) klapperichi hirtipedes Kaszab, 1958

New records. Pakistan: 1 ex., Baluchistan, Nushki, Kilki Jamaldini , 22.vii.2012 ( FUUZM) ; 2 exx., idem, 16.vi.2016 ( CB) ; 2 exx., idem, 8.x.2016 ( CB) ; 3 exx., Baluchistan, Khuzdar , vi.1991, S. Prespl ( CB) .

Distribution. Polytypic species;both the nominotypical and the subsp. hirtipedes Kaszab, 1958 have been recorded from Afghanistan and Pakistan (Bologna, 2008).

Remarks. The nominotypical form, described from Afghanistan, differs from the subsp. hitipedes in having less dense punctures on the head and pronotum. We suspect that hirtipedes could be a distinct species or a subspecies of another Eumylabris species.

Mylabris (Eumylabris) magnoguttata (Heyden, 1881)

New records. Pakistan: 13 exx., Gilgit , 21.vi.1991, S. Prespl ( CB, CC, CK) .

Distribution. Central Asiatic element, distributed from southeastern Russia through the whole of Central Asia to northwestern China (Bologna, 2008). The intraspecific taxonomy needs classification and the subspecies pardoi Kaszab, 1958 is endemic to Afghanistan and northern Pakistan. The presence in Pakistan of the nominotypical form, tentatively cited by Bologna (2008), must be confirmed.

Remarks. This is a polytypic species but, pending a revision of the subgenus, the recognition of subspecies is impossible.

Mylabris (Eumylabris) posticalis (Dokhtouroff, 1889)

Distribution. Species widely distributed in Central Asia: Iran, Afghanistan, Pakistan, Kyrgyzstan, Uzbekistan, Chinese Xinjiang.

Remarks. The taxonomy of this species and that of the M. calida complex (the Maghrebian populations probably are distinct from others) needs revision and requires the examination of types of all related taxa.

Mylabris (Eumylabris) pulchra Kaszab, 1958

New records. Pakistan: 1 ex., Baluchistan, Central Zarghun , 21.iv.1957 ( CK, from the website http://www. meloidae.com/en/pictures/36260) .

Distribution. Recorded from Afghanistan ( Kaszab, 1958); new species record for Pakistan.

Mylabris (Eumylabris) quinqueplagiata Kaszab, 1958

Distribution. Previously recorded only from Afghanistan and Pakistan (Bologna, 2008). Cited by Saha (1972a) from the Salt Range.

Mylabris (Eumylabris) tiflensis Billberg, 1813

New records. Pakistan: 3 exx., Baluchistan, Zhob , v.1991, S. Prepsl ( CC); 5 exx., Punjab prov., Bhakar (Mianwali), 15.v.2008, M. A. Akhter ( CC) .

Distribution. Recorded from northwestern India and Pakistan (Lahore: Lyallpur) by Kaszab (1958 as M. klapperichi adam ) and by Saha (1972a), who proposed the synonymy of these taxa. Erroneously cited by Hashmi and Tashfeen (1992) as M. tiplensis .

Remarks. We examined specimens of this species at MNHN in Marseul’s collection. The synonymy with M. klapperichi adam Kaszab, 1958 needs confirmation.

Mylabris (Argabris) impedita bogatschevi Kaszab, 1958

New records. Pakistan: 1 ex. northwestern Pakistan, northern areas, Shandur Pass , ca. 60 km ENE Chitral, 16.– 17.vii.1998, J. Rejsek ( CC)

Distribution. Polytypic species distributed in Tajikistan (the nominate and the subsp. kryzhanovskii Pripisnova, 1987 , and lopatini Pripisnova, 1987), Kazakhstan, Afghanistan (both the nominate and the bogatshevi subspecies), Uzbekistan. New for Pakistan.

Mylabris (Argabris) klugi Redtenbacher, 1850

Distribution. Polytypic species distributed from Turkmenistan and Iran to the Chinese Xinjiang, through Kazakhstan, Tajikistan,andAfghanistan ( subsp. kabuliensis Kaszab, 1958 ). Recorded by Hashmi and Tashfeen (1992) from Pakistan.

Croscherichia goryi (Marseul, 1870)

New records. Pakistan: 1 ex., Baluchistan, Nushki , 16.vi.2016 ( CB) ; more than ca. 50 exx., western Baluchistan, Turbat , 8–19.iv.1993, S. Bečvář ( CC) ; 8 exx., SW Baluchistan, Gwadar , 17–18.iv.1993, S. Bečvář ( CC) ; 2 exx., Lower Dir, Khyber Pakhtunkhwa, Samar Bagh , 17.vii.2012 ( CB) ; 1 ex. idem, 22.vii.2012 ( FUUZM) ; 1 ex., Khyber Pakhtunkhwa, Dera Ismail Khan , 12.v.2016 ( FUUZM) ; 4 exx., Pazni Rek , 23.iii.1935 ( NHM) ; Jangul, Makran , 1.vi.1963, G. Popov ( NHM) ; Ormara , 26.iv.1931, M. Sharif ( NHM) .

Distribution. Arabic-Sindian element, distributed in northeastern Africa ( Egypt, N Sudan), Levant and Arabic Peninsula, southern Iraq and Iran, southern Pakistan and western India ( Bologna and Coco, 1991). The species also is distributed in northeastern Afghanistan and this is a new species record for this country: Jalalabad env., Nangahar, 550 m, J. Pljushtch, 13.v.2010 (3 exx. CC) and 13.v.2012 (1 ex, CB; 2 exx. CC).

Croscherichia salavatiani Kaszab, 1968

New records. Pakistan: 2 exx., Baluchistan, Nushki , 16.vi.2012 ( CB; FUUZM) ; 3 exx., idem, 8.x.2016 ( CB) ; 4 exx., Baluchistan, Nushki, Kili Jamaldini , 22.vii.2016 ( FUUZM) ; 1 ex., western Baluchistan, Tump , 90 km W of Turbat, 13–15.iv.1993, S. Bečvář ( CC) ; 8 exx., western Baluchistan, Turbat , 8–19.iv.1993, S. Bečvář ( CC) .

Distribution. Arabic-Sindian species, distributed in the Arabic Peninsula, southern Iran, southern Afghanistan, southern Iran, and southern Pakistan. Previously recorded from two Pakistan localities by Bologna and Coco (1991).

Figures

Figures 3–4. Zonitis afghanica Kaszab, 1958: habitus (3); Zonitoschema melanarthra (Fairmaire, 1894): habitus (4).