Tylototriton panhai Nishikawa, Khonsue, Pomchote & Matsui 2013a

Phimmachak, Somphouthone, Aowphol, Anchalee & Stuart, Bryan L., 2015, Morphological and molecular variation in Tylototriton (Caudata: Salamandridae) in Laos, with description of a new species, Zootaxa 4006 (2), pp. 285-310 : 299-300

Tylototriton panhai Nishikawa, Khonsue, Pomchote & Matsui 2013a

Tylototriton verrucosus (part) Wongratana 1984: 107.

Tylototriton verrucosus Type II Pomchote, Pariyanonth & Khonsue 2008: 39.

Referred material. NCSM 82961 (one female), NUOL 0 0 437 (one male), Laos, Xaignabouli Province, Botene District, Borsan Village, Phou Samrium Mountain, 17.67194ºN 101.03081ºE, 1,596 m elev., coll. 25–26 July 2011 by Somphouthone Phimmachak and Sengvilay Seateun. NCSM 82954–60 (seven males), same data as NCSM 82961 except 17.67656ºN 101.03214ºE. NUOL 0 0 421 (one female), NUOL 00422–28 (seven males), NUOL 0 0 452 (egg clutch), Laos, Xaignabouli Province, Botene District, Borsan Village, Houay Long Ya Stream, 17.69086ºN 101.04661ºE, 1,347 m elev., coll. 26 July 2011 by Somphouthone Phimmachak and Sengvilay Seateun ( Fig. 5 ).

Measurements are summarized in Table 6 .

Distribution and natural history. All were found in water, with some in a 2 x 4 m pond in montane shrub forest at 1,596 m elev., and some in a small sandy stream in semi-evergreen forest at 1,347 m elev. The eggs were adhered to sticks slightly above the surface of the stream ( Fig. 5 D).

This is the first report of the species in Laos. Elsewhere it occurs in Phu Hin Rong Kla National Park, Phitsanulok Province, Phu Luang Wildlife Sanctuary, Loei Province, and Phu Suan Sai National Park, Loei Province, Thailand ( Nishikawa et al. 2013a; Fig. 2 ).

Figures

FIGURE 2. Principal component analysis of the genus Tylototriton showing two dimensional plots of the first principal component (PC 1) versus the second (PC 2). (A) males of the subgenus Tylototriton, (B) females of the subgenus Tylotriton, and (C) males of the subgenus Yaotriton.

FIGURE 5. Tylototriton panhai: (A) Dorsal view of female (NUOL 00421), (B) dorsal view of male (NCSM 82954), (C) ventral view of female above (NUOL 00421) and male below (NUOL 00422), (D) eggs (NUOL 00452) in situ on sticks just above stream surface, (E) stream habitat at Houay Long Ya Stream, Borsan Village, Botene District, Xaignabouli Province, Laos, and (F) pond habitat at Phou Samrium Mountain, Borsan Village, Botene District, Xaignabouli Province, Laos.

Tables

TABLE 6. Measurements (mm) of adult Tylototriton panhai from Laos. Abbreviations defined in the text.

Measurement SVLA SVLP	Females Range (n =2) 80.4–82.4 85.9–86.7	Males Range; Mean ± SD (n =15) 58.9–75.2; 67.6 ± 4.0 64.9–80.4; 73.5 ± 3.8
TAL TTL AXGR	77.8–80.9 164.5–166.8 43.7–44.4	59.0–81.7; 70.7 ± 6.1 129.9–160.3; 144.2 ± 9.1 29.8–39.1; 35.2 ± 2.6
TAD HL HW	9.6–10.7 23.3–25.3 19.4–21.2	8.4–10.0; 9.1 ± 0.6 19.8–22.3; 20.9 ± 0.7 15.9–18.1; 17.3 ± 0.6
EN IN AL	4.8–5.2 5.7–6.5 29.9–30.7	3.9–5.2; 4.6 ± 0.4 4.7–6.5; 5.5 ± 0.5 27.4–32.1; 29.3 ± 1.4
PL	26.8–30.0	25.0–29.3; 27.0 ± 1.5