Ichnea digna Wolcott, 1927

Opitz, Weston, 2010, Classification, Natural History, and Evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VIII. The Genera Acanthocollum Opitz, Stegnoclava Opitz, and Ichnea Laporte, The Coleopterists Bulletin (mo 9) 64, pp. 1-65 : 38-39

Ichnea digna Wolcott, 1927

( Figs. 57, 58, 143 , 160 , 178 , 200 ; Map 5 )

Ichnea digna Wolcott, 1927: 93 . Holotype. Gender not known. San Carlos , Costa Rica, Schild and Burgdorf (USNM). (Specimen pin mounted, no other information available.) Blackwelder 1945: 389; Corporaal 1950: 270.

Diagnosis. Specimens of this species have a narrow, essentially planar frons (although in some specimens, including the holotype, there is a faint, longitudinal impression). In these characteristics, beetles of I. digna differ from superficially similar specimens of I. gremia in that the elytral disc does not show a pale transverse fascia.

Description. Size: Length 7.7 mm; width 2.6 mm. Form ( Fig. 143 ): Oblong subovoid. Integ- umental color ( Figs. 57, 58): Cranium mostly yellow, dark markings behind eye; pronotum roseate; elytra mostly dark brown, with yellow lines along sutural and epipleural margins; profemora with anterior facies yellow, remainder brown, mesofemora and metafemora mostly yellow, with dorsal brown line. Head: HW/PW 1.0; frons minutely indented; EW/VW 3.9; antenna as in Fig. 160 , funicle not compressed. Thorax: PL/PW 1.0; elytra broad subovoid, EL/PL 5.6; EL/EW 2.2; anterior margin of protibia with 12 spines. Male genitalia: Aedeagus as in Fig. 178 . Male mesodermal internal reproductive organs ( Fig. 200 ): Medial accessory gland about as long as lateral gland; testes comprised of 22 follicles.

Variation. Size: length 5.5–6.1 mm; width 1.7– 2.5 mm. The pronotum may be entirely roseate, but with anteromedial and posteromedial dark spots, or yellow at the sides and a variously expressed dark vertical vitta on the disc.

Natural History. Specimens have been collected from December through June; some in a Malaise trap in primary and secondary rainforest and some on a fallen log of P. macroloba . The known altitudes from which these beetles have been captured range from 150 to 610 m.

Distribution. The known range of this species extends from Nicaragua to western Colombia ( Map 5 ).

Specimens Examined. 4 4. NICARAGUA: Chontales. COSTA RICA: Guanacaste, Hacienda El Oro, LaCruz, Santa Cecilia, 1992, 500 m, C. Moraga; Maritza, 1988, 600 m Malaise trap; 3 km SE Rio Naranjo, 25-31-X-1992, F. D. Parker; idem, 8-12-VI-1993, F. D. Parker: Heredia, La Selva, 10°26′N 84°01′W, 2-III-1993, 50– 150 m; idem, 15-III-1993; idem, 16-III-1993; idem, 4-IV- 1994; idem, 1-II-1996; idem, 2-IV-1998; Puerto Viejo, La Selva, 28-XII-1985, E. Giesbert; La Selva, 3 km S Puerto Viejo, 10°26′N 84°01′W, 2-IV-1984, on fallen branch of Pentaclethra macroloba, H. A. Hespenheide ; idem, 4-IV-1984, H. A. Hespenheide; idem, 27-IV-1990, H. A. Hespenheide; idem, 7-V-1990, H. A. Hespenheide; idem, 21-I-3-II-1991, H. A. Hespenheide; idem, 4-VIII-1996, H. A. Hespenheide; 11 km ESE La Virgen, 10°21N 54°03W, 6-IV- 2004, 350 m, Malaise trap: Limón: 35 km N Guapiles, 18-20- VI-1978, E. Giesbert. PANAMA: Bocas Del Toro, 40 km W Chiriquí Grande, 10-V-1999, Morris & Wappes; 18–20 km NE Fortuna Dam, 23-26-V- 1984, 610 m, E. Giesbert: Chiriquí, Hacienda Las Lagunas, 8-I-1981, dead trunk, G. Ekis: Panamá, Barro Colorado, Canal Zone, V-1929, Darlington; Altos de Pacora, 4-10-I, E. Giesbert; Cerro Azul, 4 km N Goofy Lake, 9°12′N 79°23′W, 24-VII- 1970, 700 m, H. A. Hespenheide; 3 km S Cerro Jefe, 24-VII- 1970, 700 m, R. P. Hespenheide; Cerro Campana, 29-VI- 1974, 350 m, Erwin & Whitehead; 10 km N El Llano, 16-20-V- 1984, 427 m, E. Giesbert; Madden Dam, 21-V-1978, G. B. Marshall. COLOMBIA: Cauca, Isla Gorgona Mancora, 2°58′N 78°11′W, 29-IV-9-V-2000, 60 m Malaise trap, H. Torres; Isla Gorgona Alto, El Mirador, 2°58′N 78°11′W, 26-V-10-VI- 2000, 180 m, Malaise trap, H. Torres; Gorgona, El Helechal, 2°58′ N 78°11′ W, 7-20-II-2001, 30 m, Malaise trap, H.

Torres. Specimens are deposited in CHAH, FSCA, IAVH, INBC, JEWC, USNM, and WOPC.

Figures

Map 5. Geographic distribution of Ichnea spp.

Figs. 142–144. Line habiti of 142) Ichnea incerta;

Figs. 157–168. Antennae of Ichnea spp. 157) I. aequinoctialis; 158) I. acanthomelina; 159) I. dimidiatipennis; 160) I. digna; 161) I. divisa; 162) I. gremia; 163) I. incerta; 164) I. plumbea; 165) I. marginella; 166) I. atra; 167) I. aterrima; 168) I. gregata.

Figs. 176–190. Aedeagi of Ichnea spp. 176) I. plumbea; 177) I. gremia; 178) I. digna; 179) I. callanga; 180) I. frenata; 181) I. procera; 182) I. opaca; 183) I. divisa; 184) I. incerta; 185) I. aequinoctialis; 186) I. gregata; 187) I. acanthomelina; 188) I. marginella; 189) I. aterrima; 190) I. lycoides.

Figs. 197–202. Mesodermal reproductive organs of Ichnea spp. 197) I. aequinoctialis, male; 198) I. plumbea, male; 199) I. incerta, male; 200) I. digna, female; 201) I. aequinoctialis, female; 202) I. incerta, female.