Stephanollona pauper ( REUSS , 1874)

ZÁGORŠEK, KAMIL, 2010, BRYOZOA FROM THE LANGHIAN (MIOCENE) OF THE CZECH REPUBLIC PART II: SYSTEMATIC DESCRIPTION OF THE SUBORDER ASCOPHORA LEVINSEN, 1909 AND PALEOECOLOGICAL RECONSTRUCTION OF THE STUDIED PALEOENVIRONMENT, Acta Musei Nationalis Pragae Series B 66 (3 - 4), pp. 139-255 : 139-255

Stephanollona pauper ( REUSS, 1874)

Pl. 137, Fig. 1-6

v. * 1874 Lepralia pauper m. – Reuss p. 164, Pl. 5, Fig. 4

v. * 1974 Hippoporella pauper ( REUSS, 1874) – David and Pouyet p. 173, Pl. 7, Fig. 8

1977 Hippoporella pauper ( REUSS, 1874) – Vávra p. 130 (cum syn.)

non 1989 Hippoporella pauper ( REUSS, 1874) – Schmid p. 48, Pl. 14, Fig. 1-3

2006 “ Hippoporella” pauper ( REUSS, 1874) – Berning p. 124, Fig. 164-166 (cum syn.)

T y p e: Holotype deposited in the Natural History Museum Vienna under the number 1878.11.49 and illustrat- ed by David and Pouyet (1974).

M a t e r i a l: More than 30 specimens from different sections studied in detail. An additional 3 specimens from the Reuss collection stored in the Natural History Museum Vienna under the number 1878.11.36 from the section Baden .

D i a g n o s i s: Autozooeciawithwelldevelopedmarginal pores. Primary orifice with a wide sinus, aperture small, slightly cleithridiate. Secondary aperture with sinus. Oral spines very large, autozooecia without ovicells have 4 spines, ovicelled ones 2 spines only. Large, spatulate adventitious avicularia with a pivotal bar and a large rostrum situated laterally from the aperture. A second, small, suboral avicularium may be present. Ovicell globular with a wide frontal fissure not immersed into the distal part of autozooecium.

R e m a r k s: The type has 4 oral spines, small avicularia, clearly visible marginal areolar pores, but no ovicell and large adventitious avicularia. David and Pouyet (1974) and also Berning (2006) already described ovicells and large adventitious avicularia and added these features to the description of the species.

Stephanolona has been discussed in detail by Gordon (1994). He argued that the characteristic features of the genus are the presence of condyles, marginal areolar pores, oral spines, large adventitious avicularia and small globular ovicells. Because of the presence of large adventitious avicularia, a sinus, oral spines and small globular ovicells in Lepralia pauper REUSS, 1874 , this species is listed in the genus Stephanolona .

The species Stephanolona pauper differs from Schizotheca fissa in having adventitious avicularia ( Schizotheca fissa has vicarious avicularia) and a wide sinus. Schedocleidochasma incisa differs from Stephanolona pauper in having a cleithridiate aperture, small avicularia and no oral spines.

Schmid (1989) described specimens with very prominent cleithridiate apertures and with no adventitious avicularia or oral spines. These specimens resemble Schedocleidochasma incisa ( REUSS, 1874) .

Berning (2006) illustrated a specimen of “ Hippoporella” pauper with large spatulate adventitious avicularia, oral spines and also ovicells with two slits at their margin. The confusion about types was discussed by Berning (2006). I agree with him and understand the species in the same manner as he does. However, he suggested a new genus for this species. I believe that Stephanolona completely covers all the features visible in S. pauper .