HYDROPHILINAE, Latreille, 1802

Peck, Stewart B., 2016, The beetles of the Lesser Antilles (Insecta, Coleoptera): diversity and distributions, Insecta Mundi 2016 (460), pp. 1-360 : 49-51

SUBFAMILY HYDROPHILINAE

TRIBEBEROSINI

Berosus (Berosus) interstitialis Knisch 1924: 270 (replacement name for Berosus tessellatus Fleutiaux and Sallé 1890: 377 ); Blackwelder 1944-1957: 169; Ramos 1946: 32; Miskimenand Bond 1970: 80; Hansen 1999: 88; Turnbowand Thomas 2008: 40. = Berosus (Berosus) stribalus d’Orchymont 1946: 13 ; Spangler 1981: 156; Hansen 1999: 94; Deler-Hernandez et al. 2013: 83 (synonymy); Thomas et al. 2013: 32 of Caymans. = Berosus tessellatus Fleutiauxand Sallé 1890: 377 (secondary homonym) of Guadeloupe. Distribution. Bahamas, Barbuda, Caymans, Cuba, Dominica, Caymans, Guadeloupe, Hispaniola, Jamaica, Mona, PuertoRico, St. Croix, St. John, St. Lucia, St. Thomas; widespread Antilles endemic; not North or Central America .

Berosus (Enoplurus) undatus (Fabricius) 1792: 185 ( Hydrophilus ); Blackwelder 1944-1957: 169; Hansen 1999: 82; Deler-Hernandez et al. 2013: 99. = Berosus guadeloupensis Fleutiaux and Sallé 1890: 376 of Guadeloupe; Wolcott 1951: 242 of Puerto Rico. Distribution. Antigua *, Cuba, Grenada *, Guadeloupe, Puerto Rico. USA (TX), Mexico ; widespread Antillesand North and/or Central America .

Derallus rudis Sharp 1887: 765 ; Fleutiaux and Sallé 1890: 377; Blackwelder 1944-1957: 169; Hansen 1999: 97; Bass 2006a: 13, 2006b: 33. Distribution. Antigua, Cuba, Guadeloupe, Nevis. Mexico , Guatemala (AFN); not South America ; widespread Antillesand North and/or Central America .

Anaceana sp., new genus record, new species record. Distribution. Grenada * (CMNC, USNM), St. Vincent *; Lesser Antillesendemic.

Paracymus confusus Wooldridge 1966: 719, 1971: 402 ; Nilsson 2001: 110; Bass 2006a: 13, 2006b: 33; Ivie et al. 2008b: 240; Turnbowand Thomas 2008: 41; Daltry 2009: 63. Distribution. Antigua, Bahamas, Barbados *, Bequia*, Dominica (AFN), Grenada *, Montserrat, Nevis, St. Lucia, St. Vincent *. USA (widespread), Canada (BC), Mexico ; widespread Antillesand North and/or Central America .

Paracymus delatus Wooldridge 1971: 401 ; Harrison and Rankin 1976: 287, 291; Spangler 1981: 159; Hansen 1999: 110; Daltry 2009: 63. = Paracymus armatus (Sharp) 1882: 64 ( Hydrobius ); Fleutiaux and Sallé 1890: 375 of Guadeloupe; Blackwelder 1944-1957: 172; Hansen 1999: 109 (distribution not in West Indies). Distribution. Antigua *, Barbados *, Cuba, Dominica, Grenada *, Guadeloupe, Martinique*, Mayreau*, PuertoRico, St. Lucia, St. Thomas; widespread Antillesendemic. Notes. In marshes and estuarine pools.

Paracymus nanus (Fall) 1910: 99 ( Creniphilus ); Wooldridge 1966: 715, 1971: 403; Nilsson 2001: 112; Bass 2003: 279; Bass 2006b: 33. = Paracymus sp. ?, Uyttenboogaart 1902: 113 of Barbados. Distribution. Bahamas, Barbados, St. Kitts. USA (FL, MS, LA); widespread Antillesand North and/or Central America ? Notes. Undetermined species are reported from Caymans (Thomas et al. 2013: 34).

TRIBEHYDROPHILINI

SUBTRIBEACIDOCERINA

Enochrus (Methydrus) bartletti Short 2004: 352 ; Valentineand Ivie 2005: 275; Ivie et al. 2008b: 240; Turnbowand Thomas 2008: 40. = Philhydrus aequalis Sharp 1882: 68 , Fleutiauxand Sallé 1890: 375 of Guadeloupe; Harrison and Rankin 1976: 279, 291 of St. Vincent; Daltry 2009: 63 of St. Lucia. = Enochrus orchymonti Mouchamps 1956: 10 (of Peru); misidentification in Bennett and Alam 1985: 20; Turnbowand Thomas 2008: 40. = Enochrus nebulosus Say 1822: 277 , Ramos 1946: 32 of Mona; Wolcott 1951: 244 of Puerto Rico. =” Enochrus sharpi ” (nomen nudum), Bass 2003: 279 of Barbados. = Philhydrus spec. ? Uyttenboogaart 1902: 113 of Barbados. Distribution. Antigua *, Bahamas, Barbados, Cuba, Dominica, Grenada *, Guana, Hispaniola, Martinique*, Mona, Montserrat, Puerto Rico, St. Croix, St. Lucia, St. John, St. Thomas, St. Vincent; widespread Antillesendemic. Notes. Found in marshes and stony bottom rivulets. the Lesser Antilles records of Enochrus (Methydrus) debilis (Sharp) 1882: 69 ( Philydrus ) in Hansen 1999: 180 of Hispaniola, Guatemala and Mexico may refer to E. bartletti .

Enochrus pseudochraceus Gundersen 1977: 256 ; Hansen 1999: 186; Short 2004: 355; Bass 2003: 279, 2006a: 13, 2006b: 33; Thomas et al. 2013: 34. = Enochrus ochraceus Melsheimer 1845a: 101 ; Wolcott 1951: 244 of Puerto Rico; Thomaset al. 2013: 33 of Caymans. Distribution. Antigua, Barbados, Caymans, Cuba, Dominica, Caymans, Grenada *, Hispaniola, Jamaica, Mayreau*, Mustique*, Puerto Rico, St. John, St. Kitts, St. Lucia *, St. Vincent *, Union*. Mexico to Panama; widespread Antilles and North and/or Central America .

Helochares (Helochares) abbreviatus (Fabricius) 1801: 251 ; Blackwelder 1944-1957: 172; Spangler 1981: 158; Hansen 1999: 159; Short 2005: 215; Daltry 2009: 63. = Helochares (Hydrobaticus) rufobrunneus Balfour-Browne 1939: 293 of Grenada; Wolcott 1951: 2 , 44 of Puerto Rico. Distribution. Antigua *, Barbados *, Cuba, Grenada, Guadeloupe, Martinique*, Puerto Rico, St. Lucia. Costa Rica, Panama, Colombia, Venezuela and Surinam to Argentina and Bolivia; widespread Antilles and South America .

Helochares (Helochares) guadeloupensis Orchymont 1926: 233 ; Blackwelder 1944-1957: 172; Hansen 1999: 160. = Helochares pallidus (Laporte) 1840: 53 ; Fleutiauxand Sallé l889: 376 of Guadeloupe. Distribution. Guadeloupe; single island endemic. Notes. Possibly a synonym of the above widespread species.

Helochares (Sindolus) femoratus (Brullé) 1841: 59 ; Orchymont 1926: 236; Blackwelder 1944-1957: 172; Hansen 1999: 158; Bass 2006: 33. Distribution. Antigua?, Nevis. Argentina, Brazil, Colombia?, French Guiana, Venezuela (AFN); the Lesser Antillesand Latin America . Notes. The Lesser Antilles records of Bass and Orchymont need confirmation.

SUBTRIBEHYDROPHILINA

Hydrobiomorpha (Hydrobiomorpha) phallica (Orchymont) 1928: 165 ( Neohydrophilus ); Blackwelder 1944-1957: 171; Mouchamps 1959: 331; Hansen 1999: 216; Bass 2003: 279, 2006b: 33; Daltry 2009: 63. = Hydrocharistenebrioides Jacquelin du Val 1856: 50 ; Fleutiaux and Sallé 1890: 375 of Guadeloupe. = Hydrobiomorpha casta (Say) 1835: 170 ( Hydrophilus ); Miskimenand Bond 1970: 81 of St. Croix. Distribution. Antigua *, Barbados, Grenada *, Guadeloupe, Hispaniola, Martinique, PuertoRico, St. Croix, St. Kitts, St. Lucia. Panama, probably all of Central America ; Venezuela; widespread Antillesand Latin America .

Hydrophilus (Hydrophilus) ensifer Brullé 1837: 52 ; Hansen 1999: 229; Turnbowand Thomas 2008: 41; Thomas et al. 2013: 34. = Hydrophilus ater Olivier 1792: 125 (primary homonym of Hydrophilus ater Gmelin 1790 ); Blackwelder 1944-1957: 171; Wolcott 1951: 243 ; Ramos 1946: 32; Thomas et al. 2013: 34 of Caymans. Distribution. Subspecies Hydrophilus ensifer duvali Hansen 1999: 229 (new name for Hydrophilus intermedius Jaquelin du Val 1857: 48 ; Daltry 2009: 63 of St. Lucia, in Hansen 1999: 229); Barbados, Bahamas, Caymans, Cuba, Culebra, Hispaniola, Mona, Puerto Rico, the Lesser Antilles (unspecified islands). Subspecies Hydrophilus ensifer ensifer ; Colombia and French Guiana to Argentina and Bolivia; widespread Antilles and South America .

Hydrophilus (Hydrophilus) insularis Laporte 1840: 50 ; Fleutiauxand Sallé 1890: 374; Uyttenboogaart 1902: 113; Lengand Mutchler 1914: 399 ( Stethorus ); Blackwelder 1944-1957: 171; Wolcott 1951: 243 ; Miskimenand Bond 1970: 81; Hansen 1999: 230; Bass 2003: 279, 2006a: 13, 2006b: 33; Valentineand Ivie 2005: 275; Ivieet al. 2008b: 239; Turnbowand Thomas 2008: 41; Thomaset al. 2013: 34; Short and McIntosh 2014: 189. Distribution. Antigua, Bahamas, Barbados, Caymans, Cuba, Grenada *, Guadeloupe, Guana, Hispaniola, Martinique, Mayreau*, Mona, Montserrat, Nevis, St. Croix, St. Kitts, St. Vincent *, Puerto Rico. USA (CA-AZ-TX-FL), Mexico to Costa Rica, northern South America ; widespread Antilles and North and/or Central America . Not South America , contrary to Blackwelder 1944-1957: 171.

[ Neohydrophilus medius (Brullé) 1837: 54 ( Hydrophilus ); Blackwelder 1944-1957: 171 of Guadeloupe, of Martinique; Wolcott 1951: 243 of PuertoRico; Miskimenand Bond 1970: 81 of St. Croix; Hansen 1999: 215. Distribution. Bolivia; other records dubious; not the Lesser Antilles records in Blackwelder 1944-1957: 171.]

Tropisternus (Pristoternus) apicipalpis (Chevrolat) 1834 : no. 44 ( Hydrophilus ); Blackwelder 1944-1957: 218; Hansen 1999: 218. = Tropisternus agilis Laporte 1840: 53 ; Uyttenboogaart 1902: 113; Lengand Mutchler 1914: 400 of St. Vincent. Distribution. Barbados, Cuba, St. Vincent. USA (AZ), Mexico to Argentina, Bolivia; widespread New World.

Tropisternus (Pristoternus) chalybeus (Laporte) 1840: 53 ( Hydrophilus ); Blackwelder 1944-1957: 170; Hansen 1999, 219; Ivieet al. 2008b: 240. = Tropisternus nitidus Laporte 1840: 53 , Fleutiauxand Sallé 1890: 375 of Guadeloupe. Distribution. Cuba, Dominica, Guadeloupe, Montserrat, Mustique*, Puerto Rico. Mexico to Panama, to Brazil (not Argentina, Hansen 1999: 219); widespread Antilles and Latin America .

Tropisternus (Pristoternus) laevis Sturm 1826: 64 (Hydrophilus) ; Blackwelder 1944-1957: 170, as subspecies Tropisternus laevis mergus (Say) 1835: 171 ( Hydrophilus ); Hansen 1999: 220; Thomas et al. 2013: 34. Distribution. Caymans, Cuba, the Lesser Antilles (individual islands not specified). Mexico to Venezuela and Colombia to Argentina and Bolivia; widespread Antilles and Latin America .

Tropisternus (Tropisternus) lateralis (Fabricius) 1775: 228 ( Hydrophilus ); Fleutiauxand Sallé 1890: 375; Uyttenboogaart 1902: 113; Blackwelder 1944-1957: 170; Ramos 1946: 32; Bennettand Alam 1985: 20; Hansen 1999: 223; Bass 2003: 279, 2006a: 13, 2006b: 33; Ivieet al. 2008b: 240; Turnbow and Thomas 2008: 41. Distribution. Antigua, Bahamas, Barbados, Barbuda, Cuba, Dominica, Caymans, Guadeloupe, Hispaniola, Jamaica, Martinique*, Mona, Montserrat, Nevis, PuertoRico, St. Croix, St. Kitts, St. Lucia, St. Thomas (probably all Tropisternus lateralis lateralis ). USA, Mexico to Ecuador (including Galapagos), Brazil, Uruguay, Argentina (all other subspecies); widespread New World, including Hawaii (introduced). Plate 3.