Javaulius Marshall, 1956

Riedel, Alexander, 2024, Taxonomic review of weevils of the tribe Celeuthetini, mainly from Sundaland, Sulawesi and the Lesser Sunda Islands (Coleoptera, Curculionidae, Entiminae), Zootaxa 5464 (1), pp. 1-75 : 31

Javaulius Marshall

Javaulius Marshall, 1956: 61–62 .

Type species: Javaulius setosus ( Marshall, 1925) , by original designation.

Description. Integument covered with appressed, microgranulate, polygoniform scales of mosaic-like arrangement, not overlapping; elytra, legs and antennal scape with sparse, erect, narrow or spatulate scales. Head short; with angulate, transverse sulcus separating rostral base deeply incised or rather indistinct; vertex with radial furrows (partly obscured by scales in J. subvirens Marshall and J. rudis Marshall ,). Eyes convex, protruding; separated from apex of prothorax by ca. length of eye. Rostrum. Dorsum medially with longitudinal furrow (indistinct in J. mixtus Marshall ); apical declivity distinct, subglabrous (sparsely squamose in J. subvirens Marshall ); dorsal profile subangulate. Antenna. Scape curved, with erect setae. Funicle with length of article 1 and 2 subequal. Pronotum longer than wide; coarsely punctate to scabrous. Metanepisternum narrow, anteriorly with impression. Elytra subovate; with 12 striae, some abbreviated (only 10 striae in J. subvirens Marshall ). Legs. Protibia ventrally simple, not denticulate. Venter. Procoxae subcontiguous, separated by 0.05 X coxal diameter, positioned slightly closer to apex than base of prothorax. Mesoventral process trapezoid, 1.25 X as long as posterior width. Abdomen. Female ventrites 3–5 exposed, laterally slightly shorter than ventrites 1–2. Genitalia. Penis ( Figs. 16b , 17b , 18b , 19b , 20b , 21b ) with apodeme firmly attached to body. Endophallus Z-folded, near base of body forming ventral diverticulum. Transfer apparatus spiniform, shorter than body of penis.

Notes. Traditionally this genus comprises smaller species, usually inhabiting the leaf litter. All currently described species are from Java and Bali. Javaulius Marshall is closely related to Nusasyntrophus gen. n., Syntrophus Marshall , and Parasyntrophus gen. n.

Key to the species of Javaulius Marshall

1 Body larger, pronotum plus elytron 4.12–4.52mm. Species from Bali. ( Fig. 16a )............... J. carinulifer ( Heller, 1914)

1´Body smaller, pronotum plus elytron 2.95–3.38 mm. Species from Java ........................................... 2

2 Elytra with rows of small tubercles on alternate intervals ( Fig. 21a )........................ J. tuberculatus ( Faust, 1896)

2´Elytra with intervals simple except prominence on intervals 5–7................................................. 3

3 (2´) Scaling grey or brown, but not green. Apical declivity of rostrum subglabrous, without scales......................... 4

3´Scaling grey and green. Apical declivity of rostrum with sunk-in scales ( Fig. 20a )............. J. subvirens Marshall, 1956

4 (3) Rostrum dorsally with median glabrous costa, posteriorly containing median longitudinal impression. Elytral interval 6 of male at middle with distinct elongate swelling. ( Fig. 17a )...................................... J. mixtus Marshall, 1956

4´Rostrum dorsally with distinct median furrow. Elytral interval 6 of male less distinct if near middle, or forming tubercle above declivity............................................................................................ 5

5 (4´) Rostrum weakly converging to apex; dorsally with simple median furrow, without ridges. Elytral intervals 5–6 of male above declivity with tubercle. ( Fig. 18a )...................................................... J. rudis Marshall, 1956

5´Rostrum parallel-sided; dorsally with median furrow bordered by fine glabrous ridges. Elytral interval 6 of male at middle with blunt swelling. ( Fig. 19a )........................................................... J. setosus ( Marshall, 1925)

Figures

FIGURE 16. Javaulius carinulifer (Heller). Heteroglymma carinuliferum Heller, lectotype: (a) Habitus (b) Penis (c) Labels.

FIGURE 17. Javaulius mixtus Marshall, holotype: (a) Habitus (b) Penis (c) Labels.

FIGURE 18. Javaulius rudis Marshall, lectotype: (a) Habitus (b) Penis (c) Labels.

FIGURE 19. Javaulius setosus (Marshall). Heteroglymma setosa Marshall, lectotype: (a) Habitus (b) Penis (c) Labels.

FIGURE 20. Javaulius subvirens Marshall, lectotype: (a) Habitus (b) Penis (c) Labels.

FIGURE 21. Javaulius tuberculatus (Faust). Trigonops tuberculata Faust, lectotype: (a) Habitus (b) Penis (c) Labels.