Zospeum pagodulum Inäbnit, Jochum & Neubert, 2019

Inäbnit, Thomas, Jochum, Adrienne, Kampschulte, Marian, Martels, Gunhild, Ruthensteiner, Bernhard, Slapnik, Rajko, Nesselhauf, Claudia & Neubert, Eike, 2019, An integrative taxonomic study reveals carychiid microsnails of the troglobitic genus Zospeum in the Eastern and Dinaric Alps (Gastropoda, Ellobioidea, Carychiinae), Organisms Diversity & Evolution 19 (2), pp. 135-177 : 164-165

publication ID

https://doi.org/ 10.1007/s13127-019-00400-8

persistent identifier

https://treatment.plazi.org/id/300F1A79-B42A-B744-E438-FDE9FECAFD8B

treatment provided by

Felipe

scientific name

Zospeum pagodulum Inäbnit, Jochum & Neubert
status

sp. nov.

Zospeum pagodulum Inäbnit, Jochum & Neubert View in CoL n. sp.

Figure 9a–e View Fig

Type specimens Holotype NMBE 548815 View Materials , Kučka jama, 23.4.2009, leg. Bedek ; Paratypes NMBE 553415 View Materials /4, Kučka jama, 23.4.2009, leg. Bedek ; NMBE 553362 View Materials /1, Kučka jama, 23.4.2009, leg. Bedek.

Specimens examined Croatia: NMBE 548789 View Materials /1, Grnjača špilja, 24.6.2009, leg. Bedek ; NMBE 553361 View Materials /1, Pećina kod sela Puhari, 21.4.2009, leg. Bedek ; NMBE 553359 View Materials /2, Proljetna jama, 28.6.2009, leg. Lukic ; NMBE 553357 View Materials /1, Jama ispod Tominićevog brega, 25.4.2009, leg. Lukic ; NMBE 553358 View Materials /1, Jama ispod Tominićevog brega, 25.4.2009, leg. Bilandzija ; NMBE 553363 View Materials /5, Jama ispod Tominićevog brega, 25.4.2009, leg. Lukic ; NMBE 553360 View Materials / 1, Lučićka špilja, 7.6.2008, leg. Kljakovic ; NMBE 548807 View Materials /1, Grnjača špilja, 24.6.2009, leg. Bedek.

Diagnosis Shell ca. 1.8 mm, conic, transparent, sometimes ribbed, peristome roundish with a straight parietal shield, columella with a parietalis and a weak columellaris.

Measurements (n = 18): sh: 1.61–1.99 mm (mean: 1.764 ± 0.089 mm); sw: 1.08–1.26 mm (mean: 1.167 ± 0.049 mm); ah: 0.8–1.01 mm (mean: 0.924 ± 0.059 mm); aw: 0.66– 0.81 mm (mean: 0.747 ± 0.038 mm); number of whorls: 4.75–5.5 (mean: 5.056 ± 0.22).

Description Shell conical, translucent when fresh; shell usually smooth, but a ribbed population occurs in Grnjača špilja; aperture taller than wide; parietal shield well-differentiated from the lip, its margin usually straight, sometimes convex; parietalis present in the aperture, though comparatively weak; columellaris starting behind the aperture, usually weak.

Not really separable from other smooth-shelled species within the Z. frauenfeldii clade (such as Z. frauenfeldii , Z. bucculentum , Z. robustum ); whorl usually less shouldered than in Z. bucculentum . Both species demonstrate considerable variation.

Etymology Named for the pagoda-like shell shape of the species in general and the holotype in particular.

Distribution (Supplementary Fig. S12) Endemic to the Croatian Primorje-Gorski Kotar region, where it is known primarily from five caves on the Istrian peninsula: Kučka jama ( Fig. 9a View Fig ), Grnjača špilja ( Fig. 9e View Fig ), Proljetna jama, Pećina kod sela Puhari ( Fig. 9c View Fig ; together with Z. pretneri ) and Jama ispod Tominićevog brega ( Fig. 9b View Fig ). An empty shell from Lučićka špilja near Delnice ( Fig. 9d View Fig ) is morphologically assigned to this species.

Kingdom

Animalia

Phylum

Mollusca

Class

Gastropoda

Order

Ellobiida

Family

Ellobiidae

Genus

Zospeum

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