Cybaeus adenes, Chamberlin & Ivie, 1932

Bennett, Robb, Copley, Claudia & Copley, Darren, 2019, Cybaeus (Araneae: Cybaeidae): the adenes species group of the Californian clade, Zootaxa 4711 (2), pp. 245-274 : 247-248

publication ID	https://doi.org/ 10.11646/zootaxa.4711.2.2
publication LSID	lsid:zoobank.org:pub:06B09245-2722-4F4D-9076-41E68A17E8A7
persistent identifier	https://treatment.plazi.org/id/2F5B4D33-707D-FFA0-B289-446FFA5EFBB6
treatment provided by	Plazi
scientific name	Cybaeus adenes
status

Treatment

Key to the species of the Cybaeus adenes View in CoL group

(males of C. pearcei and C. torosus unknown; females of C. adenes and C. grizzlyi are apparently morphologically indistinguishable)

1. Males............................................................................................. 2

- Females........................................................................................... 8

2(1). Proximal arm of tegular apophysis with single, pointed tip (Figs 12, 18, 25, 56, 61, 63); with ( Figs 25 View FIGURES 25–27 , 61–64 View FIGURES 61–64 ) or without (Figs 12, 56) basal, pointed projection or proximally directed dorsal keel............................................ 3

- Proximal arm of tegular apophysis with tip bifid ( Figs 1 View FIGURES 1–3 , 44 View FIGURES 44–46 , 50 View FIGURES 50–53 , 59–60 View FIGURES 56–60 , 65, 67 View FIGURES 65–68 ) or trifid ( Figs 35 View FIGURES 35–37 , 57 View FIGURES 56–60 ); proximally directed dorsal keel present (but may be inconspicuous) ( Figs 57, 59–60 View FIGURES 56–60 , 65, 67 View FIGURES 65–68 )......................................... 5

3(2). Patellar apophysis with 40–50 peg setae covering dorsal surface (Fig. 13). Proximal arm of tegular apophysis without basal, pointed projection or dorsal keel ( Fig. 56 View FIGURES 56–60 ). West central California: southern San Francisco Bay area to northern Monterey County ( Fig. 69 View FIGURE 69 )................................................................. amicus Chamberlin & Ivie View in CoL

- Patellar apophysis with about 25–30 peg setae dorsally or anterodorsally ( Figs 19 View FIGURES 18–20 , 26 View FIGURES 25–27 ). Proximal arm of tegular apophysis with basal, pointed projection or dorsal keel ( Figs 61–64 View FIGURES 61–64 ). West or east central California ( Fig. 70 View FIGURE 70 )................... 4

4(3). Proximal arm of tegular apophysis with simple untwisted tip and prominent, bluntly pointed dorsal keel ( Figs 61–62 View FIGURES 61–64 ). West central California: Contra Costa and Alameda Counties ( Fig. 70 View FIGURE 70 )................................. grizzlyi Schenkel View in CoL

- Proximal arm of tegular apophysis with simple, slightly corkscrewed tip and dorsal keel a broad, bluntly pointed process dorsal to tip ( Figs 63–64 View FIGURES 61–64 ). East central California: Placer and El Dorado Counties ( Fig. 70 View FIGURE 70 )..... auburn Bennett View in CoL spec. nov.

5(2). Patellar apophysis nearly lacking peg setae ( Fig. 36 View FIGURES 35–37 ); proximal arm of tegular apophysis with broad, trifid tip and prominent dorsal keel ( Figs 57–58 View FIGURES 56–60 ). West central California: Santa Clara County to Monterey County ( Fig. 70 View FIGURE 70 )............................................................................................... reducens Chamberlin & Ivie View in CoL

- Ten to 30 peg setae ( Figs 2 View FIGURES 1–3 , 51 View FIGURES 50–53 ); proximal arm of tegular apophysis with narrower, bifid tip ( Figs 59–60 View FIGURES 56–60 , 65, 67 View FIGURES 65–68 ), dorsal keel reduced ( Figs 59–60 View FIGURES 56–60 ) or prominent ( Figs 65–68 View FIGURES 65–68 ). West central California: north of Santa Clara County ( Figs 69–70 View FIGURE 69 View FIGURE 70 )..... 6

6(5). Proximal arm of tegular apophysis with divergent tips ( Figs 59–60 View FIGURES 56–60 ). West central California: Sonoma to San Francisco Counties ( Fig. 69 View FIGURE 69 ).................................................................... adenes Chamberlin & Ivie View in CoL

- Proximal arm of tegular apophysis with convergent tips ( Figs 65, 67 View FIGURES 65–68 ). West central California: San Mateo and Napa Counties ( Fig. 70 View FIGURE 70 )........................................................................................... 7

7(6). Tips of proximal arm of tegular apophysis larger, relatively strongly convergent, pincer-like ( Fig. 67 View FIGURES 65–68 ); 13 peg setae anterodorsally on patellar apophysis ( Fig. 45 View FIGURES 44–46 ). West central California: San Mateo County ( Fig. 70 View FIGURE 70 )............. sanbruno Bennett View in CoL

- Tips of proximal arm of tegular apophysis relatively weakly convergent, not pincer-like ( Fig. 65 View FIGURES 65–68 ); about ten peg setae dorsally on patellar apophysis ( Fig. 51 View FIGURES 50–53 ). West central California: Napa County ( Fig. 70 View FIGURE 70 )............. schusteri Bennett View in CoL spec. nov.

8(1). Atrium with lateral margins strongly curved antero-medially ( Figs 47–48 View FIGURES 47–49 ); copulatory ducts contiguous (or nearly so) ( Figs 48–49 View FIGURES 47–49 ). West central California: San Mateo County ( Fig. 70 View FIGURE 70 ).................................... sanbruno Bennett View in CoL

- Atrium with lateral margins simple, not curved anteriorly ( Figs 4 View FIGURES 4–8 , 9 View FIGURES 9–11 , 15 View FIGURES 15–17 , 21 View FIGURES 21–24 , 30 View FIGURES 28–30 , 41 View FIGURES 38–43 ); copulatory ducts widely separated ( Figs 6 View FIGURES 4–8 , 11 View FIGURES 9–11 , 17 View FIGURES 15–17 , 24 View FIGURES 21–24 , 29 View FIGURES 28–30 , 43 View FIGURES 38–43 ). East or west central California ( Figs 69–71 View FIGURE 69 View FIGURE 70 View FIGURE 71 )............................................. 9

9(8). Atrium weakly to moderately concave and broad, length from epigastric groove to anterior margin <width between lateral margins ( Figs 38, 41 View FIGURES 38–43 , 21–22 View FIGURES 21–24 ).......................................................................... 10

- Atrium strongly concave and narrower, length 1.2–3.0 times> width ( Figs 6 View FIGURES 4–8 , 15 View FIGURES 15–17 , 33 View FIGURES 33–34 , 54 View FIGURES 54–55 ).......................... 11

10(9). Atrium at anterior margin of vulva ( Figs 38, 41 View FIGURES 38–43 ); vulva compact, apparently strongly reduced ( Figs 39–40, 42–43 View FIGURES 38–43 ). West central California: Santa Clara County to Monterey County ( Fig. 70 View FIGURE 70 )..................... reducens Chamberlin & Ivie View in CoL

- Atrium antero-medially located ( Figs 21–22 View FIGURES 21–24 ); vulva “normal” ( Figs 22–24 View FIGURES 21–24 ). East central California: Placer and El Dorado Counties ( Fig. 70 View FIGURE 70 )............................................................... auburn Bennett View in CoL spec. nov.

11(9). Atrium inverted U-shaped or lyriform, widest anteriorly ( Figs 4, 7 View FIGURES 4–8 , 9 View FIGURES 9–11 , 30 View FIGURES 28–30 ); copulatory ducts smoothly curved posteriorly, not turning anteriorly at spermathecal heads ( Figs 6, 8 View FIGURES 4–8 , 28–29 View FIGURES 28–30 ).................................................. 12

- Atrium inverted U- or vase-shaped (not lyriform), widest posteriorly ( Figs 15 View FIGURES 15–17 , 53–54 View FIGURES 50–53 View FIGURES 54–55 ); copulatory ducts smoothly curved posteriorly ( Figs 16–17 View FIGURES 15–17 ) or turning anteriorly at spermathecal heads ( Figs 34 View FIGURES 33–34 , 55 View FIGURES 54–55 )................................ 13

12(11). California: north and west San Francisco Bay area ( Fig. 69 View FIGURE 69 ).............................. adenes Chamberlin & Ivie View in CoL

- California: east San Francisco Bay area ( Fig. 70 View FIGURE 70 ).............................................. grizzlyi Schenkel View in CoL

13(11). Atrium inverted vase-shaped, lateral margins convex ( Fig. 53 View FIGURES 50–53 ). California: Napa County ( Fig. 70 View FIGURE 70 )................................................................................................. schusteri Bennett View in CoL spec. nov.

- Atrium inverted U-shaped, lateral margins concave ( Figs 15 View FIGURES 15–17 , 33–34 View FIGURES 33–34 , 54 View FIGURES 54–55 ). California: San Francisco Bay area counties south of San Francisco and Berkeley to Monterey County ( Figs 69–71 View FIGURE 69 View FIGURE 70 View FIGURE 71 )............................................. 14

14(13). Copulatory ducts turning anteriorly at spermathecal heads ( Figs 34 View FIGURES 33–34 , 55 View FIGURES 54–55 ); length of atrium (from epigastric groove to anterior margin) 2–3 times> width ( Fig. 33 View FIGURES 33–34 , 54 View FIGURES 54–55 ). West central California: Alameda County ( Fig. 71 View FIGURE 71 )........................ 15

- Copulatory ducts smoothly curved posteriorly, not turning anteriorly ( Figs 16–17 View FIGURES 15–17 ); length of atrium 1.2–1.6 times> width ( Fig. 15 View FIGURES 15–17 ). West central California: southern San Francisco Bay area to northern Monterey County ( Fig. 69 View FIGURE 69 )............................................................................................ amicus Chamberlin & Ivie View in CoL

15(14). Copulatory ducts broad anteriorly (extending laterally beyond spermathecal bases), heavily sclerotized posteriorly ( Fig. 55 View FIGURES 54–55 ); lateral margins clearly visible ventrally through integument of uncleared epigynum ( Fig. 54 View FIGURES 54–55 ).... torosus Bennett View in CoL spec. nov.

- Copulatory ducts narrow anteriorly (not extending laterally beyond spermathecal bases), not heavily sclerotized posteriorly ( Fig. 34 View FIGURES 33–34 ); not clearly visible through epigynal integument ( Fig. 33 View FIGURES 33–34 )........................ pearcei Bennett View in CoL spec. nov.