Burmapyris ohmkuhnlei, Brazidec & Legendre & Perrichot, 2023

Brazidec, Manuel, Legendre, Frederic & Perrichot, Vincent, 2023, Diversity and phylogeny of the extinct wasp subfamily Lancepyrinae (Hymenoptera, Bethylidae) revealed by mid-Cretaceous Burmese amber, Arthropod Systematics & amp; Phylogeny 81, pp. 345-369 : 345

Burmapyris ohmkuhnlei sp. nov.

Fig. 2

Etymology.

The specific epithet is a patronym honouring Dr Christoph Öhm-Kühnle, who generously donated the specimen for study. The specific epithet is to be treated as a noun in the genitive case.

Material studied.

Holotype IGR.BU-060, a complete female; housed in the amber collection of the Geology Department and Museum of the University of Rennes, France (IGR).

Type locality and horizon.

Noije Bum, Hukawng Valley, Kachin State, northern Myanmar; late Albian-early Cenomanian, ca. 99 Ma, mid-Cretaceous.

Diagnosis.

Female. Head elongate, ovoid (Fig. 2A, F ); mandible small; flagellomere 1 shorter than pedicel (Fig. 2D ); notaulus fully impressed (Fig. 2B ); fore wing with 1Rs&1M only slightly angled at junction with Rs+M (Fig. 2C ); profemur swollen; meso- and metalegs slender (Fig. 2E ).

Description.

Body weakly depressed and poorly pubescent (length 3.80 mm). - Head prognathous, elongate, ovoid; LH: 0.83 mm, WH: 0.66 mm; WF: 0.52 mm, HE: 0.23 mm; OOL: 0.31 mm; WOT: 0.10 mm; DAO: 0.05 mm; VOL: 0.43 mm; compound eye elliptical, longer than high, located anteriorly on head; clypeus with median lobe triangular, lateral lobe not developed; space between toruli depressed; antenna filiform, reaching mesoscutellum posteriorly; scape 1.73 times as long as pedicel (length 0.19 mm), dorso-ventrally flattened; 11 flagellomeres cylindrical, distinctly longer than wide except flagellomere 1 (flagellomere 1 length 0.09 mm; flagellomeres 2-10 length 0.10-0.12 mm), slightly longer than wide, shorter than pedicel; flagellomere 11 longest, tapering at apex (length 0.16 mm); mandible decussate, with four to five (?) teeth, first and second apical teeth longest, remaining shorter; ocellar triangle small, anterior ocellus posterior to supra-ocular line; occipital carina present, forming small arch posterior to ocelli. - Mesosoma with dorsum smooth (length 1.46 mm); propleuron only slightly visible dorsally; dorsal pronotal area narrower anteriorly, lateral margin straight, posterior margin widely concave; anteromesoscutum shorter than dorsal pronotal area, notaulus impressed on posterior two thirds of anteromesoscutum, convergent, absent anteriorly, parapsidal signum present; mesoscuto-mesocutellar sulcus reniform, narrow; mesoscutellum posteriorly overlapped by long metanotum; metapectal-propodeal complex rectangular, without postero-lateral spine, median carina present, continuing on propodeal declivity. Fore wing hyaline, at least reaching third metasomal segment (LFW: 2.46 mm); C, Sc+R, M+Cu, 1A tubular; 1Rs&1M only slightly angled at junction with Rs+M; 1Rs shorter than 1M; 1M and cu-a aligned; Rs+M tubular, reaching apex of pterostigma; pterostigma slightly rounded; short stub of post-stigmal abscissa of R1 present; 2r-rs&Rs arising on distal half of pterostigma, long but not closing [2R1] cell; [2R1] cell large. Legs slender, only profemur swollen; tibial spur formula 1-2-2; tarsal claws slightly curved; arolium small. - Metasoma smooth, fusiform and elongate (length 1.51 mm); petiole short; six exposed tergites, covering sternites laterally; sting exserted.

Systematic remarks.

Following the key to the genera of Lancepyrinae of Jouault et al. (2021), our specimen keys to Burmapyris Jouault, Perrichot and Nel, 2021 due to the fore wing with Rs+M vein tubular, the [2R1] cell open, the pterostigma elongate and Rs+M long, extending distally to [1M] cell, and [1M] cell not fully enclosed by tubular veins. Beside these characters, it possesses several other features reminiscent of Burmapyris : the general shape of the body, the very similar fore wing venation, the long antenna and the dorso-ventrally flattened scape. After reexamining the holotype of B. azevedoi , we noticed that it also possesses the median metapostnotal carina, strengthening the attribution of our specimen to Burmapyris . Burmapyris ohmkuhnlei sp. nov. differs from B. azevedoi in the shape of the head (more elongate and less rounded), the smaller mandible, the less pronounced angle of 1Rs&1M, the notaulus not fully impressed and the slender legs with only the profemur swollen.

Figures

Figure 2. Burmapyris ohmkuhnlei sp. nov., holotype female number IGR. BU- 060. A habitus in dorsal view; B mesosoma in dorsal view; C fore wing in dorsal view; D head and antenna in dorsal view; E habitus in lateral view; F head and anterior mesosoma in lateral view. Scale bars: 1 mm (A, E); 0.5 mm (B, C, D, F).