Entada borneensis Ridl., J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 67: 307. 1898.

O'Donnell, Shawn A., Ringelberg, Jens J. & Lewis, Gwilym P., 2022, Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade), PhytoKeys 205, pp. 99-145 : 99

Entada borneensis Ridl., J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 67: 307. 1898.

Type.

MALAYSIA. Borneo, Sarawak, Sarawak River, Penkulu Ampat, G.D. Haviland s.n. (holotype: K [K000635744]) .

Description.

Liana> 40 m long. Leaves: rachis 5-9.5 cm, sub-glabrous to tomentose, terminating in a bifurcating tendril; pinnae 2 opposite pairs per leaf, each with 3-7 pairs of alternate to sub-opposite leaflets, except for the distal opposite pair; leaflets narrowly oblong to obovate, 1.4-4 × 0.7-1.8 cm, base asymmetrically rounded, apex rounded and emarginate, both surfaces glabrous, main vein puberulous. Inflorescence: a 19-40 cm long, solitary, axillary spike, rachis tomentose. Flowers: yellowish or greenish-white to white, sessile, staminate or bisexual; calyx cupular, 0.5-0.6 mm long, glabrous; petals 2 × 0.6-0.8 mm; stamen filaments 4-6 mm long. Fruit: a gigantic, torulose craspedium, 50-120 × 10-13 cm, with transverse septa between seeds dividing the fruit into one-seeded segments which, upon ripening, fall from the persistent replum; segments 9-10 cm long; epicarp coriaceous, endocarp chartaceous. Seeds: circular, laterally compressed, 4 cm in diameter, hard, brown, lacking a pleurogram.

Distribution.

Borneo.

Habitat and ecology.

Primary and secondary rainforest, especially along rivers; in sandy clay substrates, loams and soils derived from limestone; 0-800 m alt.

Note.

Nielsen (1992) noted that E. borneensis is locally common but rarely collected.