Campylopus trachyblepharon (Müll. Hal.)Mitt.

J. Florschutz-de Waard, H. R. Zielman & M. A. Bruggeman- Nannenga, 2011, Flora of the Guianas, Series C, fascicle 2., Kew: Kew Publishing : 13

13. Campylopus trachyblepharon (Müll. Hal.)Mitt. , J. Linn. Soc., Bot. 12: 80. 1869.

- Dicmnum trachyblepharon Müll. Hal. , Syn. Musc. Frond. 1: 389. 1848. Type: Brazil, Beyrich s.n. (NY).

- Campylopus arenicola (Müll. Hal.)Mitt. , J. Linn. Soc., Bot. 12: 77. 1869.

- Dicranum arenicola Müll. Hal. , Bot. Zeitung (Berlin) 13: 761. 1855. Type: Brazil, Serra do Itatiaia, Pabst s.n. (NY).

Rather large plants growing in loose tufts. Stems to 7 cm high, usually interruptedly foliate, densely tomentose, between the tufts scarcely foliate. Leaves in the comal and lower tufts spreading, slenderly lanceolate, to 7 mm long, between the tufts shorter, more or less appressed; apex acute, Serrate in upper half, dentate at the tip, costa ca. 1/3 of leaf base, percurrent to short excurrent, in cross-section with stereids at ventral and dorsal side, at back with conspicuous, serrate, 2-3 cell high lamellae; alar cells coloured and iniiated, in the smaller stem leaves less conspicuous; inner basal laminal cells rectangular, slightly incrassate, towards margin rounded quadrate; upper laminal cells subquadrate or rhomboid, incrassate. Perichaetial leaves ovate, abruptly narrowed to a long and narrow subula. Seta to 1 cm long, capsule curved, operculum obliquely rostrate. Calyptra cucullate, fringed at base.

Distribution: E Africa, Bermuda, Guyana, Coastal plains of SE Brazil.

Ecology: Terrestrial in low savanna vegetation on white sand; not known from Suriname and French Guiana.

Selected specimens: Guyana: Timehri, Dakara Cr., Gradstein 3734 (L); Santa Mission, trail W of village, Florschütz-de Waard 6059 (L).

Note: In the Guyanan collections the verticillate foliation is not always distinct, in the Gradstein collections even absent; the prominent lamellae at the back of the costa and the strongly serrate upper lamina with subquadrate lamina] cells distinguish this species from all other Guyanan species. The basal laminal cells are variably incrassate and pitted; the cells just above the auricles may be thin-walled but quickly become more incrassate in distal direction.