Oedignatha ulopatulisus, Barrion & Litsinger, 1995

Barrion, A. T. & Litsinger, J. A., 1995, Family Clubionidae Wagner- Genera Alaeho, Castianeira, Agroeca, Phrurolithus & Scotinella, Riceland Spiders of South and Southeast Asia, Oxon, UK: CAB International, pp. 170-180 : 174-178

Phrurolithus ulopatulisus new species

( Figs 100 a-i, 101 a-e)

Description: Female: Total length 4.35 mm. Cephalothorax 1.85 mm long, 1.35 mm wide, 1.00 mm high. Abdomen 2.50 mm long, 1.20 mm wide, 1.40 mm high. Cephalothorax dark reddish brown with shallow pits, widest above coxae II, cephalic anterior lateral margins almost straight and with a snout anterior to the AME, posterior end of the cephalothorax truncate. Fovea longitudinally short. Eight eyes, all clear, in two rows, AE more procurved than PE. Eye diameter (mm): AME = 0.13, ALE = PME = 0.10, PLE = 0.08. Eye separation: AME-AME one sixth shorter than one AME diameter, AME-ALE as long as one half ALE diameter, PME-PME 1.8 times PME or 1.4 times AME diameter, PM E-PLE as long as one AME diameter, ALE-PLE as long as one ALE diameter. Median ocular quadrangle narrower in front than behind, length shorter than the anterior width. Clypeus height large, nearly twice AME diameter. Sternum same color as cephalothorax, coxae, labium, and endites, anterior margin broadly cleft in the middle for the reception of the labium, lateral margins strongly concave for the coxae and extended to the base o f carapace in between coxae, posterior end widely extended beyond coxae IV, producing a tube encircling the pedicel. Labium longer than wide, above midlength o f maxillae constricted laterally below midhalf, apical end slightly concave at the middle. Maxillae long without concave depression to receive base of pedipalp, brownish yellow sc opula e borne in the yellowish apical end. Chelicerae reddish brown, short and robust with boss-like dorsosubbasal swelling, promargin with four teeth and retromargin with three teeth, all larger than those in the promargin. Legs moderately long and slender, light reddish brown, subbase of femora slightly swollen posteriorly with a spine dorsally. Spination in the femora I = 1-0-1-0, II = III = IV = 1-0-0-0; tibiae I = 0-13-0-0, II = 0-7-4-3, III = 0-2-0-2, IV = 0-0-2-2, metatarsi I = 0-12-0-0, II = 0-5-3-0, III = 0-2-0-0, IV = 0-0-2-2. Metatarsus I/tarsus I = 1.60. Tarsi two-clawed, each claw with four equally spaced teeth. Claw tuft present, split in two rows when smashed, apices of hair tuft moderately clubbed. Leg formula 4123. Pedipalp a little shorter than total length of femora and patella of leg I, and femora with four long ventral setae.

Abdomen longer than wide, dorsum forms a hard sclerotized plate but membranous anterior to anal tubercle, venter sclerotized in the anterior one half along the epigynal area and with a pair of large half-moon-shaped sclerotized plates posterior to the epigastric furrow. Apical one third of abdomen overhangs the cephalothorax. Rest of the abdominal venter and laterals membranous. Spinnerets yellowish brown, anterior pair almost contiguous and relatively longer and stouter than the posterior pair. Epigynum a sclerotized subrectangular plate with a deep cleft in the anterior margin and a pair of globular spots (shadow of spermathecae) dorsad of the epigastric furrow. Spermathecae sacs globular, oblique, and diverging. Fertilization tube about one half the diameter of spermathecae, upcurved and diverging.

Length ofleg and pedipalp segments (mm):

Leg	Femur Patella Tibia Metatarsus Tarsus Total
1	1.50	0.58	1.55	1.10	0.70	5.43
2	1.25	0.55	1.10	0.90	0.68	4.48
3	1.00	0.50	0.85	1.00	0.60	3.95
4	1.55	0.60	1.45	1.50	0.65	5.75
Pedipalp 0.65		0.30	0.43	-	0.65	2.03

Male: Total length 4.25 mm. Cephalothorax 2.00 mm long, 1.45 mm wide, 1.35 mm high. Abdomen 2.25 mm long, 1.38 mm wide, 0.91 mm high. Cephalothorax similar to the female except for a more pronounced black longitudinal fovea, rougher surfaces, and truncate posterior enti. Eight eyes in two rows, all clear except PME. AE shorter than PE, straight to slightly procurved. PE more procurved than AE. Eye diameter (mm): AME = 0.13, ALE = 0.10, PME = 0.09, PLE = 0.08. Eye separation: AME-AME 0.8 eye diameter, AME-ALE 0.60 AME diameter, PME-PME 1.2 times AME or 1.7 times PME diameter, PME-PLEas long as AME separation, ALE-PLE as long as one PME diameter. Median ocular quadrangle wider than long, anterior and posterior width similar in length. Clypeus height large, 2.7 times AME diameter. Sternum, labium, maxillae, and chelicerae as in the female. Legs brownish red except blackish gray patches in the apical one half of femora. Spination in femora I = 1-0-0-0, II = 1-0-1-0, III = 0-0-0-0, IV = 1-0-0-0; tibiae I = 0-7-5-0, II = 0-15-0-0, III = 0-3-0-2, IV = 0-4-0-1; metatarsi I = 0-3-4-0, II = 0-10-0-0, III = 0-1-0-2, IV = 0-2-0-1. Metatarsus I/tarsus I = 1.67. Tarsi two-clawed, each claw with four teeth. Claw tuft in two rows between claws, each hair with a clubbed apex. Leg formula 1423. Pedipalp as long as combined length of femur I and patella I, retrolateral tibial apophysis with a black bidentate apophysis on its side, embolus long and thread-like, conductor forms an L-shaped sclerotized structure with a pointed tip in retrolateral view.

Length ofleg and pedipalp segments (mm):

Leg	Femur Patella Tibia Metatarsus Tarsus Total
1	1.65	1.65	1.70 1.50	0.90	6.40
2	1.40	0.60	1.25 1.18	0.63	5.06
3	1.10	0.50	0.90 1.10	0.60	4.20
4	1.70	0.55	1.65 1.70	0.75	6.35
Pedipalp 0.80		0.30	0.34 -	0.85	2.29

Abdomen blackish with three pairs of ovate faint brown dorsal spots, longer than wide and strongly convex, anterior one third roughly punctated compared to the posterior two thirds. Venter with two sclerotized plates separated by a narrow membranous membrane subposteriorly, genital groove with a transparent orificelike structure in the middle of the two book lung openings. Spinnerets with a membranous margin, anterior pair close to each other and longer than wide.

Etymology: Named after the snout-like cephalic area in Tagalog.

Natural history: Three females and a male were collected from a pitfall trap set adjacent to upland rice and dipterocarp forest in August and October.

Material examined: PHILIPPINES: Luzon Is., Quezon Prov., Real, Llavac VU. , holotype female, 24 October 1984, A.T. Barrion ; two females, and one male paratype, 22 August 1984, same data as holotype. All are in bad state with legs and body separated from each other after examination .

Distribution: Philippines.

Figures

Fig. 100. Female Phrurolithus ulopatulisus n. sp. (a); cheliceral teeth (b); sternum, labium, and maxillae (c); chelicera (d); leg I (e); pedipalp (f); and dorsal (g) and ventral (hi) views of epigynum.

Fig. 101. Male Phrurolithus ulopatulisus n. sp. (a); ventral (b) and lateral (cd) views of pedipalps; and tibial apophysis (e)