Eryma veltheimii ( Münster, 1839 )

Devillez, Julien & Charbonnier, Sylvain, 2021, Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda), Geodiversitas 43 (2), pp. 25-73 : 40

Eryma veltheimii ( Münster, 1839)

( Fig. 7 D-F)

Glyphea veltheimii Münster, 1839: 22 , pl. 10, fig. 1. — Fraas 1855: 94. — Glaessner 1929: 159. — Woods 1930: 80, pl. 22, figs 5-6. — Van Straelen 1936: 9. — Förster 1966: 124, fig. 23, pl. 17, figs 2, 4. — Taylor 1979: 34. — Feldmann & Titus 2006: 64. — Karasawa et al. 2013: table 1.

Eryma veltheimii – Bronn 1849: 579. — Oppel 1861: 358; 1862: 36, pl. 7, fig. 5. — Beurlen 1928: 156. — Schweigert et al. 2000: 7, fig. 1c, pl. 4, figs 1-3. — Garassino & Schweigert 2006: 8, fig. 8, pl. 1, fig. 2, pl. 11, figs 3-4. — Feldmann & Titus 2006: 64. — Fürsich et al. 2007a: table 2. — Odin et al. 2019: 652.

Clytia veltheimii – Beurlen 1928: 170.

Galicia veltheimii – Schweitzer et al. 2010: 25. — Charbonnier & Garassino 2012: 864. — Audo et al. 2014: 462. — Schweigert 2015: fig. 544.

TYPE MATERIAL. — Holotype BSPG AS VII 186. TYPE LOCALITY. — Kapfelberg quarry near Kelheim, Bavaria, Germany. TYPE AGE. — Kimmeridgian.

DESCRIPTION

Carapace

Short, spiny rostrum; fusiform intercalated plate; elongated cephalic region, representing almost half of the length of the carapace; wide, deep cervical groove, strongly inclined, joined to dorsal margin; deep gastro-orbital groove, elongated and oblique; postcervical groove not identified; wide, deep branchiocardiac groove, strongly inclined.

Pleon and uropods

Somites with subtriangular pleurites; telson with a median line and two longitudinal crests along its lateral margins, crests ended by a small spine; uropods as long as telson, with a small spine on the external margin; uropodal endopods with a longitudinal carina; uropodal exopods with a diaeresis, and a longitudinal carina.

Thoracic appendages

Chelate P1; P1 propodus subrectangular, strongly elongated, dorso-ventrally compressed; presence of a strong spine at the inner distal extremity of the propodus, above the dactylus; slender fingers, progressively narrowing to their distal extremity, slightly curved inward; occlusal margin without teeth; P1 carpus short, subtriangular; thin P2-P5.

Ornamentation

Carapace covered by small tubercles; intercalated plate covered by small tubercles; oblique orbital row of tubercles in cephalic region; presence of a strong antennal spine; smooth P1 propodus and fingers.

DISCUSSION

This species is known by some specimens, mainly from the Kimmeridgian plattenkalks of Wattendorf ( Audo et al. 2014). Because of the compression of the holotype, it is difficult to clearly identify the grooves of the carapace. Indeed, only the cervical, gastro-orbital and branchiocardiac grooves are wellmarked. However, the assignation to Eryma is supported by the gastro-orbital groove wich is not divided in two branches and by the shape of the P1 chelae (elongated subrectangular propodus; elongated fingers, curved inward and narrowing to their distal extremity).

The difficulty to identify the carapace grooves does not allow the comparison with that of other species. The presence of a spine at the distal extremity of the inner margin of the propodus ( Fig. 7D ) is characteristic of this species. Moreover, the lack of ornamentation on the P1 chelae is a characteristic only found in E. veltheimii and E. westphali . A spiny rostrum like in E. veltheimii is not found in E. mandelslohi , E. modestiforme , and E. ventrosum . Garassino & Schweigert (2006) pointed out the absence of spines on the margin of the telson, contrary to E. modestiforme . However, the careful examination of the type specimen shows that two marginal spines are present ( Fig. 7F ). Finally, the carapace of E. veltheimii is only covered by tubercles, contrary to E. georgeii , E. mandelslohi , E. quadriverrucatum , E. ventrosum , and E. westphali .

Figures

FIG. 7. — Eryma major Oppel, 1861, Eryma veltheimii (Münster, 1839) and Eryma westphali Schweigert, Dietl & Röper, 2000 from the Kimmeridgian of Germany: A, holotype SMNS 3682 of E. major from Egesheimer;B, specimen SMNS 67655 of E. major from Gerhausen; C, specimen SMNS 64371 of E. major from Nusplingen; D-F, holotype BSPG AS VII 186 of E. veltheimii from Kehlheim: part (D), counter-part (E), details of telson and uropods (F); G-J, holotype SMNS 24227 of E. westphali from Nusplingen: general view (G), schema of the carapace (H), schema of left chela (I), schema of right chela (J); C, specimen SMNS 63733 from Nusplingen. Abbreviations: a, branchiocardiac; as, antennal spine; b, antennal groove; b1, hepatic groove; c, postcervical; d, gastro-orbital groove; e1e, cervical groove; ip, intercalated plate. Black arrows show the spines. Photographs and line drawings: J. Devillez. Scale bars: 1 cm.