Draconarius dialeptus, Okumura, 2013

Okumura, Ken-ichi, 2013, Seven New Species of Spiders of the Subfamily Coelotinae (Araneae: Agelenidae) from Kyushu, Japan, Species Diversity 18, pp. 87-97 : 95-96

Draconarius dialeptus sp. nov.

[Japanese name: Yaku-chibi-yachigumo] ( Fig. 7A–D )

endogynum. For example, it resembles Draconarius venustus Ovtchinnikov, 1999 (q.v.), the type species of this genus, as well as D. pakistanicus Ovtchinnikov and Inayatullah, 2005 (q.v.), D. naranensis Ovthinnikov and Inayatullah, 2005 (q.v.), and D. latellai Marusik and Ballarin, 2011 (q.v.) in having enlarged and convoluted copulatory ducts and small accessorial glands situated in the central portion of the endogynum. Draconarius verrucifer sp. nov. is distinguished from the above four species by the absence of epigynal teeth and the presence of a single, tiny protrusion.

Description. Female (holotype). Total length 4.5; carapace 1.8 long, 1.4 wide; abdomen 2.7 long, 1.6 wide; sternum 1.1 long, 1.0 wide. Ocular area ( Fig. 5G ): anterior eye row almost straight and posterior one slightly recurved in frontal view. Eye sizes: AME 0.04, ALE 0.09, PME 0.09, PLE 0.10. Distances between eyes: AME–AME 0.03, AME–ALE 0.03, PME–PME 0.08, PME–PLE 0.05, AME–PME 0.08, ALE–PLE 0.05. MOA: anterior width 0.11, posterior width 0.26, length 0.21. Leg measurements: I, 6.5 (1.8, 2.3, 1.4, 1.0); II, 6.0 (1.7, 1.9, 1.4, 1.0); III, 5.7 (1.5, 1.8, 1.6, 0.8); IV, 7.9 (2.1, 2.4, 2.3, 1.1).

Chelicera ( Fig. 5H ): promargin with three teeth and retromargin with two widely spaced teeth.

Epigynum and endogynum ( Fig. 5E, F ): epigynal plate almost circular; epigynal teeth absent, but single tiny protrusion present on center of posterior margin of epigynum;

Material examined. Holotype: NSMT-Ar 9867, male, Kusugawa, 230 m alt., Yaku Island , Kagoshima Prefecture, (30°39.7′N, 130°58.8′E) 5 January 2008.

Paratype: NSMT-Ar 9868, male, Nagata, Yaku Island , Kagoshima Prefecture, 3 January 2008 .

Non-type specimen: same data as the holotype, 1 male .

Diagnosis. The new species is similar to D. venustus Ovtchinnikov, 1999 (q.v.), the type species of this genus, in having two retromarginal teeth on the chelicera, an elongat- ed cymbial furrow, a developed conductor, and a long embolus. The new species also resembles D. aspinatus ( Wang, Yin, Peng and Xie, 1990) , and D. bituberculatus Wang, Yin, Peng and Xie, 1990 in the absence of a patellar apophysis, but it can be distinguished from the latter two species by the structure of the conductor. That of D. dialeptus is tripartite, but that of D. aspinatus is spiniform and broad, and that of D. bituberculatus is posteriorly extended ( Wang et al. 1990; Wang 2003).

Description. Male (holotype). Total length 4.0; carapace 2.0 long, 1.4 wide; abdomen 2.0 long, 1.2 wide; sternum 1.0 long, 0.9 wide. Ocular area ( Fig. 7C ): both anterior and posterior eye rows almost straight in frontal view. Eye sizes: AME 0.04, ALE 0.10, PME 0.11, PLE 0.10. Distances between eyes: AME–AME 0.02, AME–ALE 0.01, PME– PME 0.05, PME–PLE 0.02, AME–PME 0.07, ALE–PLE 0.03. MOA: anterior width 0.10, posterior width 0.27, length 0.22. Leg measurements: I, 7.0 (1.9, 2.4, 1.6, 1.1); II, 6.2 (1.7, 2.0, 1.5, 1.0); III, 5.9 (1.6, 1.9, 1.6, 0.8); IV, 8.1 (2.1, 2.5, 2.4, 1.1).

Chelicera ( Fig. 7D ): promargin with three teeth and retromargin with two widely separated teeth.

Palpus ( Fig. 7A, B ): patellar apophysis nearly reduced; LTA large, with dorsal projection; RTA rounded at tip; cymbium narrow; cymbial furrow elongate, occupying threefourths of cymbial length; median apophysis spatulate, slightly curved at tip; conductor large, pointed and tripartite; embolus extremely long, complexly curved.

Coloration: carapace yellowish brown, with indistinct radial flecks; dorsum of abdomen grayish brown, with yellowish-brown chevrons; sternum pale yellow; chelicerae, maxillae, and labium yellowish brown; legs yellowish brown without ring flecks.

Distribution. Yaku Island, Kagoshima Prefecture ( Fig. 6 ).

Etymology. The specific name, an adjective, is derived from Classical Greek, meaning very small and refers to the extremely small size of the new species.

Figures

Fig. 7. Draconarius dialeptus sp. nov., NSMT-Ar 9867, holotype, male. A, Male palpus, ventral view; B, same, retrolateral view; C, ocular area, frontal view; D, left chelicera, ventral view.Scales: 0.5mm.

Fig. 5. A–D, Coelotes oxyacanthus sp. nov.; E–H, Draconarius verrucifer sp. nov. A–B, NSMT-Ar 9863, paratype, male; C–D, NSMT-Ar 9862, holotype, female; E–H, NSMT-Ar 9865, holotype, female. A, Male palpus, ventral view; B, same, retrolateral view; C, E, epigynum; D, F, internal female genitalia; G, ocular area, frontal view; H, left chelicera, ventral view.Scales: 0.5mm.

Fig. 6. Map of Kyushu, Japan, showing the distribution of Coelotes oxyacanthus sp. nov. Draconarius verrucifer sp. nov., D. dialeptus sp. nov., and C. hiradoensis Okumura and Ono, 2006, also showing geographical replacement of C. hiradoensis by C. oxyacanthus.