Iberattus semiglabratus (Simon, 1868) Prószyński & Noordam & Oger & Schäfer, 2018

Prószyński, Jerzy, Noordam, Aart, Oger, Pierre & Schäfer, Michael, 2018, Delimitation of Mediterranean genus Iberattus gen. n., with comments on genus Saitis (Araneae: Salticidae) *, Ecologica Montenegrina 18, pp. 82-98 : 85-86

Iberattus semiglabratus (Simon, 1868) comb. n.

Figure 1

Attus semi-glabratus Simon, 1868b: 561 (Dm) .

Phlegra semiglabrata Simon, 1876a: 127 .

Phlegra semiglabrata Simon, 1937: 1221 , 1267, f. 1957-1959 (m, Df).

Euophrys lundbladii Schenkel, 1938 b: 15 , f. 6 (f, provisional name only).

Euophrys semiglabrata Heciak & Prószyński, 1984a: 378 , f. 1-14 (Tmf from Phlegra ).

Habrocestum semiglabratum Prószyński, 2009a: 320 , f. 62 (f, no justification provided for any generic transfer). Euophrys semiglabrata Barrientos et al., 2014: 36 , f. 15-17 (mf).

Euophrys semiglabrata Lecigne, 2017: 34 , f. 10G-I (m). [Synonyms listed after Word Spider Catalog].

Type species. Attus semi-glabratus Simon, 1868 [= Iberattus semiglabratus (Simon, 1868) ].

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Material. Lectotype male, paralectotypes: 4 males and 4 females, original collection label is: "902. Phl. [egra] semiglabrata E. S. Astur., Portug., la Rhune " - collection Simon - MNHN-Paris (lectotype designated by Hęciak & Prószyński (1984)).

Other documentation used: color photographs in photogalerie of Pierre Oger (https://arachno.piwigo.com/index?/category/1232-euophrys _semiglabrata ) ( Figs 1 A-G).

Remarks. In spite of renewed interest in latest time ( Barrientos et al., 2014, Lecigne, 2017) the species remains little known, apparently due to its rarity and cryptic coloration ( Fig. 1A ) but also to radical lost of natural coloration of live specimen ( Fig. 1B ) during preservation ( Figs 1 C-D), which hampers recognition in the field.

Diagnosis. Live male has black carapace with eyefield light reflecting, apparently due to iridescent scales, and whitish scales making median thoracal streak and marginal lines, abdomen is dark gray with two parallel, thin black streaks. Palps of the EUOPHRYINES type ( Figs 1E , G-I). Epigyne with a pair of posterior "windows" and a pair of round translucent darker spots in front of them ( Figs 1F, P, R ), spermathecae three chambered ( Figs 1Q , S-T).

Description. Facsimile of the detailed description ( Fig. 6 ) is demonstrated below, in the section "Appendixfacsimiles". Body coloration of live male specimen dark ( Fig. 1A ), carapace with light reflecting eyefield and whitish median streak indistinct and ending before posterior end of the carapace, abdomen cryptic colored - gray, with brown shade and indistinct, still darker pattern. Palps is shown on Figs 1E , G-I. Epigyne with a pair of posterior "windows" and a pair of round translucent darker spots in front of them ( Figs 1F, P, R ), spermathecae three chambered ( Figs 1Q , S-T).

Distribution. Males from Rasmalhos ( Portugal) and Escaut - Pyrénées atlantiques ( France), female from 64 Bielle, Turon de la Técouère ( France). Matching of sexes and conspecificity not verified.

Nomenclatorical correction

Name " Euophrys lundbladii Schenkel, 1938: 15 , f. 6 ( Fig. 7 ) was not described but proposed by Schenkel conditionally, as a replacement for specimen identified provisionally by him as " Euophrys difficilis [?]”, to be substituted if the species would appear new. It appeared that the species was not new, but misidentified " Phlegra” semiglabrata , so provision by Schenkel never become valid. Roewer (1954: 1175) apparently decided to introduce order and implemented Schenkel's provision, without reason and authority to do that. Name Euophrys " lundbladi " Schenkel, 1938 never become available (in the nomenclatorical sense) because there was never such species described and there is no type specimen to be connected with. Therefore:

Euophrys lundbladi Roewer (1945[1955]: 1175) should be considered nomen nudum (see Fig. 7 ).

Informal subgroup of genera SAITEAE (sensu Prószynski)

Remarks. Informal group of genera EUOPHRYINES is so large (800 to 1000 species according to various classifications) that any subdivision is welcomed, providing that is based on clear cut, well noticeable diagnostic characters. The difficulty is that EUOPHRYINES are relatively uniform group, with little diversity of genitalic characters, so even if some genera are standing out, one does not know where to place remaining. Morphological and behavioral features seem to separate "third leg gesturing" genera into informal subgroup SAITEAE from the "first leg gesturing " Euophrys , which could be considered as a nucleus of another, delimited subgroup EUOORYDEAE ( Prószynski et al., 2018 c: 34), while a hundred of other genera are pending classification into their subgroups within EUOPHRYINES. The present, incomplete subdivision job may constitute, perhaps, some small progress in setting modern system of Salticidae , but that still leaves hundreds of other species pending their grouping inside EUOPHRYINES.

Figures

Figure 1. Diagnostic characters of Iberattus semiglabratus documented by color macrophotographs (A-F) and diagnostic drawings (G-M). A - live male, B-C - color pattern of a male, in different stage of fading during preservation, D - color pattern of preserved female, E - palp, F - epigyne, G-I - palp, J - dorsal pattern of preserved male, K - cheliceral dentition, L - frontal view of male, note reduction of clypeus, M-N - dorsal pattern of preserved two females, O - male by Barrientos et al., P-T - epigyne and internal structure of epigyne in two females.

Figure 6. Facsimile of description of Euophrys semiglabrata by Hęciak & Prószyński (1984).

Figure 7. Diagnostic drawings of Euophrys semiglabrata misidentified as "Euophrys difficilis" (its conditional replacement name E. lundbladii never become valid).