Coelidiini, F.Dohrn, 1859

Nielson, M. W., 2011, 2953, Zootaxa 2953, pp. 1-85 : 7-9

Key to the genera of Coelidiini (males)

1. Old world genera*..................................................................................... 2

- New world genera..................................................................................... 7

2. Style with apophysis usually glabrous, sometimes with 1–2 apical spines and or spurs, rarely with digitate processes, if present digitate processes on distal 2/3 of apophysis................................................................. 3

- Style with apophysis with several long digitate processes on distal 1/3 ( Figs. 6, 13, Nielson 1986)........ Stylolidia Nielson

3. Aedeagus with 1–3 prominent apical, subapical or mulitspinulated spines on shaft, sometimes glabrous or with indentations or spicules ( Figs. 261, 275, 460, Nielson 1982)................................................................ 4

- Aedeagus without spines on shaft, frequently with indentations or spicules apically, rarely with apical process ( Figs. 172, 203, Nielson 1982).......................................................................... Taharana Nielson

4. Aedeagus with shaft narrow, tubular, rarely inflated, with long apical or subapical spines (Figs. 306, 361, 412, Nielson 1982) .................................................................................................... 5

- Aedeagus with shaft broad basally or inflated and constricted, glabrous or with very small spines on shaft or with long, paired, spinulated apical spines ( Figs. 249, 254, 268, Nielson 1982).................................................... 6

5. Aedeagus with single glabrous or multispinulated spine ( Figs. 274, 294, 345, Nielson 1982)........... Olidiana McKamey

- Aedeagus with 2 or more glabrous or multispinulated spines (Figs. 463, 543, 595, Nielson, 1982).......... Calodia Nielson

6. Aedeagus inflated and constricted, glabrous or with small spines on shaft ( Figs. 249, 254, Nielson 1982)... Jenolidia Nielson

- Aedeagus broad, tapered distally with 2 long, multispinulated spines on apex ( Figs. 262, 269, Nielson 1982).................................................................................................... Mahellus Nielson

7. Medium size to large, robust species; head rarely angulate, narrower than to wide as pronotum; crown rarely produced distally, nearly aways as wide or wider than eye width, rarely narrower than eye width; dorsal connective never distinctly bifurcate with attachment to aedeagus; style nearly always as long as or longer than aedeagus.................................... 8

- Small to medium size, slender species; head angulate, nearly always distinctly narrower than pronotum; crown nearly always produced distally, nearly always narrower than eye width; dorsal connective nearly always bifurcate basally, sometime enclosing shaft; style shorter than to nearly as long as aedeagus...................................................... 31

8. Clypellus broad, inflated basally, width equal to or greater than base of clypeus, lateral margins constricted medially....... 9

- Clypellus narrow, flat or nearly so basally, width narrower than base of clypeus, lateral margins parallel, tapered or expanded apically............................................................................................. 16

9. Crown narrower than transocular width.................................................................. 10

- Crown as broad to twice as broad as transocular width....................................................... 11

10. Aedeagus with shaft very narrow, tubular throughout, without processes ( Figs. 18, 19, Nielson 1988); dorsal apodeme attached basally to laterobase of aedeagus ( Fig. 19, Nielson 1988)........................................ Fistulidia Nielson

- Aedeagus with shaft broad, enlarged and twisted apically (Figs. 740,741, Nielson 1982); dorsal apodeme typical, attached basally to dorsobase of aedeagus (Figure not shown)......................................... Crassinolanus Nielson

11. Crown 1 to 1 ½ times as broad as transocular width; aedeagus with apical, subapical or row of spines and or teeth........ 12

- Crown twice as broad as transocular width (Fig. 849, Nielson, 1982); aedeagus with long, medial spine (Figs. 852, 856, Nielson, 1982)........................................................................... Clypeolidia Nielson

12. Aedeagus with 1–2 apical or subapical spines, sometimes with row of short subapical spines......................... 13

- Aedeagus with row of medial spines or with spines and teeth in distal half........................................ 15

13. Aedeagus with single, recurved apical spine, directed basally.................................................. 14

- Aedeagus with 1–2 subapical spines directed distally or with row or short spines subapically (Fig. 759, 765, 772, 778, Nielson 1982; Figs. 198, 205 herein)............................................................. Dialodia McKamey

14. Pygofer with long, robust caudodoral process (Fig. 844, Nielson 1982); aedeagus robust with short, recurved apical spine (Fig. 648, Nielson 1982)...................................................................... Hamolidia Nielson

- Pygofer with short, digitate caudoventral process (Fig. 742, Nielson 1982); aedeagus narrow with long, recurved apical spine (Fig. 747, Nielson 1982).................................................................... Codilia Nielson

15. General color dark brown to black with distinctive transverse yellow band on pronotum, mesonotum and base of clavus, large yellow spot on middle of forewing; aedeagus toothed subapically with medial spines and distinctive serrated spine medially (Fig. 787, Nielson 1982)................................................................... Daridna Walker

- General color dark brown without transverse yellow band, yellow and ivory markings on pronotum and mesonotum; aedeagus with row of spines medially (1982, Fig. 793, Nielson 1982)...................................... Nedangia Nielson

16. Crown carinate laterally, usually elevated, produced about 1/3 of its distal length beyond anterior margin of eyes.......... 7

- Crown not carinate laterally, usually not elevated, produced much less than 1/3 of its distal length beyond anterior margin of eyes............................................................................................... 19

17. Crown with anterior margin acutely angled; dorsal habitus at rest more or less linear; pygofer in lateral view without paired caudal processes; aedeagus with shaft broad, spines medial or subapical........................................ 18

- Crown with anterior margin obtusely angled; dorsal habitus at rest deltoid; pygofer in lateral view often with paired caudal processes (Figs. 858, 864, 869, 874, 878, Nielson 1982); aedeagus with shaft often tubular, often with 1–2 apical spines (Figs. 862, 867, 873, 874, 882, Nielson 1982)....................................................... Coelidia Germar

18. Subgenital plate deltoid in apical 1/3 (Fig. 918, Nielson 1982); style with long apophysis (Fig. 919, Nielson 1982)............................................................................................... Deltolidia Nielson

- Subgenital plate convex apically (Fig. 910, Nielson 1982); style with very short apophysis (Fig. 909, Nielson 1982)............................................................................................ Triquetolidia Nielson

19. Pygofer in lateral view with caudal processes, caudodorsal process extended or lobelike, caudoventral process present or absent.............................................................................................. 20

- Pygofer in lateral view without caudal processes............................................................ 30

20. Pygofer with caudodorsal process widely separate from caudoventral process, if present............................ 21

- Pygofer with caudodorsal and caudoventral processes closely appressed, processes very long and digitate (Fig. 938, Nielson 1982); style setaceous (Figs. 940, 941, Nielson 1982).......................................... Crinolidia Nielson

21. Segment X without ventral process; pygofer with caudodorsal process narrow, elongate or small lobelike............... 22

- Segment X with prominent ventral spine (Fig. 945, Nielson 1982); pygofer with very large, robust caudodorsal process (Fig. 945, Nielson 1982)..................................................................... Megalidia Nielson

22. Small to moderately large, narrow to somewhat robust species; pygofer with caudodorsal process elongate or small to large, single lobe; aedeagus with spines, setae, flanges on shaft or single spine apically................................... 23

- Large, broad, very robust species; pygofer with caudodorsal process short, broad, bilobed apically ( Fig. 3, Nielson 1988); aedeagus with tiny spines apically ( Figs. 4, 5, Nielson 1988)................................... Pygmaelidia Nielson

23. Aedeagus with shaft moderately tubular to broad, with config urations of flanges or partially inflated in lateral and dorsal views .................................................................................................. 24

- Aedeagus with shaft very narrow, evenly tubular in lateral and dorsal views, without flanges or inflations............... 25

24. Aedeagus in lateral view sinuate or nearly so with single apical spine (Figs. 804, 810, 816, 825, 831, 837, Nielson 1982); style very long, narrow (Figs. 802, 807, 813, 821, 828, Nielson 1982)............................... Kramerolidia Nielson

- Aedeagus in lateral and dorsal views not sinuate, with row of short subapical spines (Figs. 1005, 1006, Nielson 1982); style moderately long, robust (Fig. 1003, Nielson 1982)............................................. Jassolidia Nielson

25. Style with long, very slender apophysis................................................................... 26

- Style with short to long, robust apophysis................................................................. 27

26. Aedeagus with serrate flange on shaft (Figs. 967, 968, Nielson 1982); style bulbous apically, with apical spine (Figs. 964, 965, Nielson 1982); subgenital plate glabrous (Fig. 966, Nielson 1982)................................. Nudulidia Nielson

- Aedeagus with toothed spines on shaft (Figs.961, 962, Nielson 1982); style attenuated apically (Fig. 959, Nielson 1982); subgenital plate pilose (Fig. 960, Nielson 1982).................................................. Dicolecia Nielson

27. Style in lateral or dorsal view with long, robust to very broad, curved apophysis; aedeagus with numerous setae, teeth and or serrate flanges....................................................................................... 28

- Style in lateral or dorsal view with very short, robust apophysis (Fig. 954, Nielson 1982); aedeagus somewhat inflated, with 3 moderately long setae on shaft (Figs. 956, 957, Nielson 1982).................................... Gicrantus Nielson

28. Forewing with small ivory or yellow spots or large yellow spot in basal half of clavus; aedeagus in dorsal and lateral views equal to nearly equal in width, with several to numerous, moderately long setae or row of short setae, spicules or spines; style rarely with apical spines................................................................................ 29

- Forewing with small ivory to yellow spots on veins and with large ivory to yellow spots or with transverse or longitudinal yellow bands; aedeagus with shaft in dorsal view often much broader medially or subapically than laterally, often with serrate lateral flanges or row of spicules, short setae or short spines on shaft (Figs. 1038, 1029, 1045, 1050, 1060, 1072, 1085, 1091, Nielson 1982); style robust, often with apical or subapical spines (Figs. 1052, 1058, 1065, 1082, 1090, 1096, 1100, Nielson, 1982)................................................................................ Omanolidia Nielson

29. Aedeagus with several to numerous, moderately long setae on shaft (Figs. 975, 989, Nielson 1982 and Figs. 214, 222, 230, 238, 246, 262 herein.................................................................... Evansolidia Nielson

- Aedeagus with row of short setae, spicules or spines on shaft (Figs. 1012, 1020, 1026, 1033, Nielson 1982)..................................................................................................... Spinolidia Nielson

30. Style profusely setose (Fig. 931, Nielson 1982); aedeagus with shaft serrate, long sharp apical spine (Fig. 934, Nielson 1982)....................................................................................... Calodicia Nielson

- Style glabrous (Fig. 996, Nielson 1982); aedeagus not as above, shaft with 2 subapical spines (Fig. 998, Nielson 1982)............................................................................................. Crinorus Nielson

31. Style nearly as long as or longer than aedeagus; dorsal connective with or without basal bifurcation................... 32

- Style about 1/4 as long as aedeagus, apophysis absent or very short (Fig. 665, Nielson 1982); dorsal connective without basal bifurcation............................................................................... Dicodia Nielson

32. Aedeagus with pair of basoventral processes (Figs. 682, 683, Nielson 1982) or single basoventral keel ( Fig. 25, Nielson 1988; Fig. 7, Nielson 1992).................................................................................. 33

- Aedeagus without basoventral processes or basoventral keel................................................... 34

33. Aedeagus with pair of very long basoventral processes; dorsal connective with arms enclosing aedeagal shaft, attached to base of processes (Fig. 710, Nielson 1982)..................................................... Ventrolidia , gen. nov.

- Aedeagus with subbasal ventral keel; dorsal connective with arms not enclosing aedeagal shaft, attached laterally to aedeagal shaft ( Fig. 25, Nielson 1988; Fig. 7, Nielson 1992)....................................... Carinoscapula , gen. nov.

34. Aedeagus with shaft inflated or partially inflated in lateral or in lateral and dorsal views, with flanges or spines from subbase to subapex (Figs. 697, 698, 703, 704, Nielson 1982); dorsal connective with or without basal bifurcation, if so, not completely enclosing aedeagal shaft (Figs. 689, 704, Nielson 1982).......................................... Boliviela DeLong

- Aedeagus with narrow, tubular shaft in lateral and dorsal views, with or without such armature; dorsal connective deeply to shallowly bifurcate or not, if deeply bifurcate completely enclosing aedeagal shaft.................................. 35

35. Dorsal connective deeply bifurcate basally, arms enclosing aedeagal shaft (Fig. 683, Nielson 1982). Collasuyusana , gen. nov.

- Dorsal connective shallowly bifurcate or not bifurcate....................................................... 36

36. Aedeagal shaft glabrous between subbase and subapex....................................................... 37

- Aedeagal shaft not glabrous, armed with spines or spicules ( Figs. 6, 27, Nielson 1986)............ Armaturolidia , gen. nov.

37. Pygofer with long, very narrow, highly membranous caudoventral process or process may be very small or absent; aedeagus with 1–2 apical spines................................................................................. 38

- Pygofer with very long, robust caudoventral process (Fig. 717, Nielson 1982; Fig. 37, Nielson 1988); aedeagus with apical spine or digitate process (Fig. 721, Nielson 1982; Fig. 39, Nielson 1988)............................ Tinocripus Nielson

38. Subgenital plate profusely setose, setae very long ( Figs. 15, 23, 39, Nielson 1986); pygofer usually without membranous lobe on caudal margin between caudal processes; aedeagus with 1–3 apical spines ( Figs. 13, 20, Nielson 1986). Paralidia Nielson

- Subgenital plate usually sparsely setose, seta very short ( Fig. 10, Nielson 1988); pygofer with membranous lobe on caudal margin between caudal processes ( Fig. 15, Nielson 1988); aedeagus with single apical or subapical spine ( Fig. 11, Nielson 1988)................................................................................. G racilidia Nielson

*A complete key to 15 genera in the Ethiopian region is presented in Nielson (1992) and is not repeated here.