Europiella angulata (Uhler)

SCHUH, RANDALL T., 2004, Revision of Europiella Reuter in North America, with the Description of a New Genus (Heteroptera: Miridae: Phylinae), American Museum Novitates 3463, pp. 1-59 : 12-17

Europiella angulata (Uhler)

Figures 1 , 3 , 6 , 7

Maurodactylus angulatus Uhler, 1895: 53 (n. sp., descr.).

Europiella angulata: Knight, 1968: 41 (n. comb.); Schuh et al., 1995: 386 (n. syn.).

Europiella yampae Knight, 1968: 43 (n. sp.; syn. by Schuh et al., 1995: 386).

DIAGNOSIS: Recognized by the relatively large size, elongate, nearly parallel­sided, broad body form (fig. 6), usually dark general coloration of dorsum, and the vesica with a short, sickle­shaped anterior apical spine and short, weakly curving, posterior spine (fig. 3), a condition shared with E. carvalhoi , E. pilosula , and E. unipuncta . Most similar in size, general appearance, and coloration to E. consors , and apparently not separable from it with certainty except by dissection of the male genitalia. Sexual dimorphism in this species is pronounced, the females being smaller than the males and conspicuously ovoid rather than parallel­sid­ ed (fig. 1).

REDESCRIPTION: Male: Relatively large, broad­bodied species, total length 3.04–3.53, width across pronotum 0.43–0.51. COLOR­ ATION (fig. 1): General coloration of dorsum usually largely dark, but sometimes extensively pale; antennal segments 1–4 black; labium with segments 1 and 2 mostly pale, segments 3 and 4 much darker; much of venter dark in dark specimens, largely pale in pale specimens; coxae, trochanters, and femora mostly dark in dark specimens, largely pale in pale specimens, femora when pale with dark spots or blotches; tibiae with pale background coloration, spines black with black bases. SURFACE AND VESTITURE (fig. 7C, D): Surface and vestiture as in generic diagnosis. STRUCTURE (fig. 7): Elongate, nearly parallel­sided, corial margin weakly convex. GENITALIA (fig. 3): Pygophore moderately large relative to size of abdomen; anterior vesical spine short, sickleshaped, posterior spine short, weakly curving, spines similar in structure to E. carvalhoi , E. pilosula , and E. unipuncta .

Female: Coloration usually light, seldom as dark as in males. Females much shorter than males, ovoid (fig. 1). Total length 2.84– 3.32, width pronotum 0.42–0.48.

HOSTS: Artemisia arbustorum , A. dracunculus , A. tridentata . Also recorded from Chrysothamnus sp. , Solidago sp. (Asteraceae) .

DISTRIBUTION: Widely distributed in the interior of western North America, from British Columbia to Saskatchewan in the north to Arizona and east to west Texas and the foothills of the Rocky Mountains in the south .

DISCUSSION: The distribution of E. angulata is overlapped almost completely by that of E. consors . The great external similarity of these species therefore makes their recognition on the basis of distribution impossible. Consequently, over much of the range of these two taxa dissection of the male genitalia would appear to be necessary to confirm that identifications are accurate.

SPECIMENS EXAMINED: Holotype of angulata : 3, Type; Colo., 1691; P. R. Uhler Collection (USNM). Holotype of Europiella yampae Knight : 3, Steamboat Springs, July 12, 1964, COLO., 6700̍, H. H. Knight (USNM). Additional specimens: CANADA: Alberta: Elkwater Park, July 25, 1952, L. A. Konotopetz, 33 (CNC). British Columbia: Mt. Kobau, 11 km NW of Osoyoos Summit, 1834 m, August 30, 1993, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 13 (CNC). Saskatchewan: Willow Bunch, July 17, 1955, A. R. Brooks, 13, 7♀ (CNC). USA: Arizona: Greenlee Co.: Apache National Forest, September 20, 1998, J. C. Schaffner, 23 (TAMU). Colorado: Archuleta Co.: 14 mi NE of Pagosa Springs, August 16, 1969, H. M. Ohlendorf, 13 (TAMU). Bent Co.: 15 mi S of Lamar, May 3, 1982, J. T. Polhemus, 163 (JTP). Lamar, 3600 ft, June 4, 1919, 33 (AMNH). Boulder Co.: Nederland, Science Lodge, 9000 ft, July 29, 1961, J. R. Stainer, 63, 3♀ (CNC). Ward, Roosevelt National Forest, July 30, 1968, L. A. Kelton, Solidago sp. (Asteraceae) , 403, 35♀ (CNC). Clear Creek Co.: West Chicago Creek, Arapaho National Forest, 9800 ft, July 28, 1968, L. A. Kelton, 23 (CNC). Eagle Co.: N of Minturn, Grouse Creek Trail, 7840 ft, June 24, 1987, D. A. Polhemus, 13 (JTP). Gunnison Co.: 5 mi NE of Almont, along Taylor River, August 31, 1980, D. A. and J. T. Polhemus, 23 (JTP). Gunnison, August 17, 1925, H. H. Knight, Chrysothamnus sp. (Asteraceae) , 23 (USNM). Hinsdale Co.: 11 mi N of Lake City, August 6, 1997, J. C. Schaffner, 23 (TAMU). Huerfano Co.: Veta Pass, August 9, 1925, H. H. Knight, 13 (USNM). Jackson Co.: 1 mi E of Gould, August 13, 1969, J. C. Schaffner, 13 (TAMU). 4 mi N of Gould, August 5, 1975, J. C. Schaffner, 13 (TAMU). North Park, July 31, 1926, F. C. Hottes, 23 (USNM). Jefferson Co.: Red Rocks Park near Morrison, 5600 ft, July 15, 1983, Schuh and Polhemus, Artemisia sp. (Asteraceae) , 173 (AMNH). Larimer Co.: Estes Park, 8800 ft, August 27, 1920, H. C. Severin, 33, 1♀ (USNM). Fall River Road, Rocky Mountain National Park, 9500 ft, August 16, 1968, L. A. Kelton, 13 (CNC). Fort Collins, June 24, 1920, 13 (USNM). Glen Haven, August 3, 1947, Lawson, 13 (KU). Moffat Co.: Craig, July 14, 1949, R. H. Beamer, 13 (KU). Montezuma Co. : Dolores, August 3, 1900, E. D. Ball, 93 (USNM). Mancos, August 13, 1925, H. H. Knight, 33 (USNM). Routt Co.: Steamboat Springs, July 12, 1964, H. H. Knight, 153, 15♀ (USNM). Saguache Co.: 7 mi W of Cochatopa Pass, August 31, 1980, D. A. and J. T. Polhemus, 473 (JTP). Jct Cochetopa Pass Rd and Cochetopa Creek Rd, August 31, 1980, D. A. and J. T. Polhemus, 153 (JTP). Idaho: Fremont Co.: Henrys Lake, 7000 ft, July 12, 1936, R. E. Miller, 23 (USNM). Montana: Deer Lodge Co.: Silver Lake, 14 mi W of Anaconda on Rt 10A, 6400 ft, August 9, 1986, Schuh, Schwartz, Stonedahl, Artemisia tridentata (Asteraceae) , 1303 (AMNH). Madison Co.: 15 mi SE of Virgina City, July 12, 1982, S. E. Cummings, 13, 1♀ (UNHAMP). Nevada: Elko Co.: Ruby Mts., Lomoille Canyon, 6200 ft, June 16, 1983, Schuh, Schwartz, Artemisia dracunculus (Asteraceae) , 143 (AMNH). New Mexico: Bernalillo Co.: Albuquerque, June 1, 1983, J. T. Polhemus, 13 (JTP). Grant Co. : 4 mi SE of Gila Cliff Dwellings Natl. Monument of Hiway 15, September 20, 1998, J. C. Schaffner, 23 (TAMU). Lincoln Co.: Ruidoso, June 26, 1940, Lipovsky, 13 (KU). Otero Co.: 4 mi E of Cloudcroft, June 22, 1979 – September 26, 1979, J. C. Schaffner et al., 53 (TAMU). 7 mi N of Cloudcroft, August 13, 1982, J. C. Schaffner, 43 (TAMU). Cloudcroft, June 17, 1976, Oman, 193 (OSU). Cloudcroft, June 27, 1940, Beamer, Hardy, Lipovsky, 1003 (KU). Taos Co.: Taos, August 20, 1927, R. H. Beamer, 63 (KU). Oregon: Crook Co.: 23 mi E of Prineville, Mark’s Creek, July 29, 1976, J. D. Lattin, Artemisia arbustorum (Asteraceae) , 63 (OSU). Wallowa Co.: Joseph, July 10, 1969, P. W. Oman, 13 (OSU). Texas: El Paso Co.: No specific locality, July 18, 1927, R. H. Beamer, 13 (KU). Utah: Cache Co.: Blacksmith Fork Canyon, June 28, 1971, G. F. Knowlton, 13 (OSU). Tony Grove Lake, August 17, 1974, G. F. Knowlton, 13 (UCD). Sevier Co.: Richfield, July 15, 1929, E. W. Davis, 13 (USNM). Summit Co.: Emory, August 16, 1929, R. H. Beamer, 13 (KU). Kimball Junction, August 31, 1967, L. A. Kelton, 13, 2♀ (CNC). Wasatch Co.: Soldier, August 13, 1906, 13 (USU). Washington Co.: La Verkin, V. M. Tanner, 13 (CAS). Red Cliff Recreation Area near Leeds, 3200 ft, October 16, 1982, D. A. Polhemus, 43 (JTP). Zion National Park, Watchman C.G., 3900 ft, October 17, 1982, D. A. Polhemus, 13 (JTP). Wyoming: Park Co.: Shoshone National Forest, August 14, 1927 – August 15, 1927, H. H. Knight, 23 (UCB). Shoshone National Forest, August 7, 1927, H. H. Knight, 23 (USNM). Sweetwater Co.: 10 mi E of Rock Springs, July 4, 1968, H. H. Knight, 13 (USNM).

Europiella artemisiae (Becker)

Figures 1 , 3

Capsus artemisiae Becker, 1864: 487 (n. sp., descr.); Schuh et al., 1995:385 (n. syn., disc., figs.).

Plagiognathus diversus Van Duzee, 1917a: 283 (n. sp.). NEW SYNONYMY.

Plagiognathus solani Matsumura, 1917: 432 (n. sp.; syn. by Schuh et al., 1995: 386).

Plagiognathus albipennis var. obscura Sahlberg, 1920: 167 (n. var.; syn. by Schuh et al., 1995: 386).

Plagiognathus (Poliopterus) gracilis Wagner, 1956: 74 (n. sp.; syn. by Schuh et al., 1995: 386).

Psallus artemisicola Knight, 1964: 149 (n. sp.); Schuh et al., 1995: 384 (n. syn. with Europiella decolor [Uhler]). REVISED SYNONYMY.

DIAGNOSIS: Always with strongly infuscate hind femora and dark areas on the hemelytra (fig. 1). Similar in appearance to E. decolor and E. lattini , but distinguished by its usually larger size and antennal segment 2 in males always greater than width of pronotum; length of antennal segment 2 not diagnostic for females; male genitalia very similar in structure to those of E. decolor and E. lattini , but pygophore and vesica smaller than those in E. lattini , and anterior apical spine longer (compare figs. 3 and 4).

REDESCRIPTION: Male: Moderately large, elongate species, total length 3.27–3.58, width pronotum 0.97–1.02. COLORATION (fig. 1): Head, pronotum, and scutellum usually mostly dark; hemelytra usually dark with pale areas at base of corium and base of cuneus; some specimens lighter with posterior margin of pronotum and lateral angles of mesoscutum orange and hemelytra generally yellow­orange except for infuscate posterior half of endocorium and posterior portion of cuneus; all antennal segments black; labium generally pale, segment 4 black; venter of body generally infuscate to nearly black; all trochanters pale; forecoxae almost entirely pale; fore and middle femora only weakly darkened, usually with some orange, hind femora heavily infuscate; all tibiae with black spines with distinct black bases. SUR­ FACE AND VESTITURE: Surface and vestiture as in generic diagnosis. STRUCTURE: Elongate, nearly parallel­sided, corial margin only weakly convex; antennal segment about one­third longer than width of head. GENI­ TALIA (fig. 3): Pygophore moderately large relative to size of abdomen; posterior apical spine of vesica short, sharply curving, hookshaped just surpassing gonopore, anterior spine conspicuously longer and weakly decurved, structure very similar to that of E. decolor (fig. 3), also similar to E. lattini (fig. 4), but anterior spine much longer and entire vesica smaller than in that species.

Female: Coloration similar to male. Body form much more strongly ovoid. Total length 2.94–3.22, width pronotum 0.94–1.10; antennal segment 2 relatively shorter than in males, less than or equal to width of pronotum.

HOSTS: Breeds on various species of Artemisia , including both herbaceous and woody taxa. Also recorded from Chrysothamnus , Haplopappus , and Solidago spp. (Asteraceae) .

DISTRIBUTION: In the Nearctic ranging from Alaska in the north to Zacatecas, Mexico, in the south. In the Rocky Mountain system ranging south into New Mexico, and at higher latitudes as far east as Minnesota and Wisconsin. Although many localities are what would be called boreal, others—such as those from southern California and Mexico —are not.

DISCUSSION: The status of this holarctic species was discussed by Schuh et al. (1995), who were the first authors to recognize its occurrence in North America.

Van Duzee (1917a) described a species Plagiognathus diversus from Santa Cruz County , California. Examination of the holotype and dissection of a male paratype of P. diversus from the type locality, as well as examination of a number of other paratypes, indicate that this taxon is a junior synonym of Europiella artemisiae , new synonymy.

Schuh et al. (1995), without comment, synonymized Psallus artemisicola Knight , described from St. Paul, Minnesota, with Europiella decolor . I can only conclude that this decision was made in haste, because upon further examination of specimens used by Knight in the preparation of his original description, this nominal taxon is clearly the same as E. artemisiae , not E. decolor (Uhler) revised synonymy.

SPECIMENS EXAMINED: Holotype of Psallus artemisicola Knight : 3, St. Paul, Minn., Sept. 3, 1923, H. H. Knight, Artemisia ludoviciana ; deposited in the USNM. Holotype of Plagiognathus diversus Van Duzee : 3, Niles Canyon, Cal., W. M. Gifford, Alameda Co., 12­VI­17, Cal., Type diversus , Type No. 357; deposited in the CAS. Additional specimens: CANADA: Alberta: Head Smashed­in Buffalo Jump Interpretive Park, September 1, 1993, M. D. Schwartz, Artemisia cana (Asteraceae) , 283 (CNC). British Columbia: 15.8 km N of S end of Muncho Lake Provincial Park, August 1, 1982, G.G.E. Scudder (UBC). 5 mi W of Toad River Lodge, August 24, 1980, R. J. Cannings (UBC). 9 mi E of Grand Forks, June 8, 1959, L. A. Kelton (CNC). 9 mi E of Grand Forks, June 8, 1959, L. A. Kelton, 23 (CNC). Blue River Bridge, km 705.6 Cassiar Highway, July 21, 1983, G.G.E. Scudder (UBC). Cassiar Hiway, km 705.6, Blue River Bridge, July 21, 1983, G.G.E. Scudder, 13 (UBC). Grand Forks, June 8, 1959, E. E. MacDougall (CNC). Hyland River, July 20, 1983, G.G.E. Scudder (UBC). Kamloops, July 1, 1959, L. A. Kelton (CNC). Kamloops, July 1, 9159, L. A. Kelton, 13 (CNC). Kootenay Pass, August 8, 1970, L. A. Kelton, Artemisia longifolia (Asteraceae) (CNC). Kootenay Pass, August 8, 1970, L. A. Kelton, Artemisia longifolia (Asteraceae) , 83, 9♀ (CNC). Lillooet, June 1, 1921, J. McDunnough, 13 (USNM). Lillooet, June 1, 1926, J. McDunnough (CNC). Lillooet, May 29, 1926, J. McDunnough (CNC). Lillooet, May 30, 1926, J. McDunnough (CNC). Lillooet, May 31, 1926, J. McDunnough (CNC). Lillooet, Seton Lake, June 16, 1926, J. McDunnough (CNC). Lilooet, June 1, 1926, J. McDunnough, 123, 20♀ (CNC). Meyer Flat, June 29, 1959, L. A. Kelton (CNC). mi 392 Alaska Highway, Summit Lake, 4200 ft, July 19, 1959, R. E. Leech (CNC). mi 392 Alaska Highway, Summit Lake, July 21, 1959, R. E. Leech (CNC). Muncho LPP, 15 km N of S end of park, August 1, 1982, G.G.E. Scud­ der, 2♀ (CNC, UBC). Nicola, September 1, 1993, G.G.E. Scudder (UBC). Nicola, September 1, 1993, G.G.E. Scudder, 13, 9♀ (CNC). Oliver, July 19, 1970, L. A. Kelton (CNC). Oliver, July 19, 1970, L. A. Kelton, 13 (CNC). Oliver, Meyer Flats, June 29, 1959, L. A. Kelton, 203 (CNC). Oliver, Vaseaux Lake, May 29, 1959, E.E. MacDougall, L. A. Kelton, 13, 9♀ (CNC). Osoyoos, Richter Pass, July 28, 1959, L. A. Kelton, 23 (CNC). Osoyoos, Richter Pass, June 28, 1959, L. A. Kelton (CNC). Spillmacheen Rest Area, September 12, 1992, G.G.E. Scudder, 1♀ (UBC). Summit Lake, mile 392 on Alaska Hiway, July 19, 1959, R. E. Leech, 4♀ (CNC). Surprise Lake, August 18, 1980, R. J. Cannings (UBC). Surprise Lake, August 18, 1980, R. J. Cannings, 33, 3♀ (CNC, UBC). Terrace, July 24, 1960, W.R. Richards, 33, 3♀ (CNC). Tetsa Br. no. 1, August 16, 1982, L. A. Kelton, Artemisia tilesii (Asteraceae) , 103, 20♀ (CNC). Tetsa Bridge no. 1, August 16, 1982, L. A. Kelton, Artemisia tilesii (Asteraceae) (CNC). Trutch, 1260 m, August 25, 1980, R. J. Cannings (UBC). Trutch, 1260 m, August 25, 1980, R. J. Cannings, 3♀ (CNC, UBC). Yahk, July 22, 1959, L. A. Kelton, 13 (CNC). Manitoba: Glenlea near Red River, Agr. Res. Sta., August 19, 1993, M. D. Schwartz, Artemisia frigida (Asteraceae) , 413 (CNC). Herchmer, August 10, 1937, D. G. Denning, 1♀ (USNM). Northwest Territories: Aklavik, July 25, 1931, Bryant, 13 (CAS). Fort McPherson, MacKenzie River, July 18, 1957, R. Hurley, 93, 2♀ (CNC). Fort Mc­ Pherson, MacKenzie River, July 20, 1957, R. Hurley, 13 (CNC). Fort McPherson, MacKenzie Riv­ er, July 24, 1957, R. Hurley, 63, 1♀ (CNC). Fort McPherson, MacKenzie River, August 6, 1922, C.H. Crickmay, 1♀ (CNC). Inuvick, July 25, 1983, L.A. Kelton, Artemisia sp. (Asteraceae) 543, 74♀ (CNC). Norman Wells, August 2, 1949, W.R.M. Mason, 33 (CNC). Norman Wells, August 7, 1949, W.R.M. Mason, 13 (CNC). Reindeer Depot, MacKenzie Delta, August 5, 1948, W.J. Brown, 43, 5♀ (CNC). Reindeer Depot, MacKenzie Delta, August 15, 1948, J.R. Vockeroth, 33, 5♀ (CNC). Reindeer Depot, Mac­ Kenzie Delta, August 2, 1948, W.J. Brown, 13 (CNC). Reindeer Depot, MacKenzie Delta, August 18, 1948, J.R. Vockeroth, 13, 2♀ (CNC). Reindeer Depot, MacKenzie Delta, August 11, 1948, J.R. Vockeroth, 13, 2♀ (CNC). Saskatchewan: 10.8 km E of Maidstone on Hwy 16, August 22, 1993, M. D. Schwartz, Artemisia biennis (Asteraceae) , 353 (CNC). Fort Walsh Natl. Hist. Park nr visitor center, September 3, 1993, M. D. Schwartz, Artemisia frigida (Asteraceae) , 263 (CNC). N. Battleford, August 22, 1993, M. D. Schwartz, Artemisia frigida (Asteraceae) , 243 (CNC). Yukon Territory: Arctic Circle, Dempster Highway, July 29, 1983, L.A. Kelton, 1♀ (CNC). Blackstone River, Dempster Hiway km 92, July 12, 1983, G.G.E. Scudder, 33, 3♀ (UBC). 14 mi E of Dawson, 1300 ft, July 29, 1962, R. E. Leech, 123, 16♀ (CNC). Dawson, July 17, 1949, W. W. Judd, 13 (CNC). Dawson, July 23, 1983, L.A. Kelton, Artemisia frigida (Asteraceae) , 93, 12♀ (CNC). Mi 87 Dempster Highway, August 4–8, 1973, G. and D.M. Wood, 13 (CNC). Destruction Bay, July 26, 1982, L.A. Kelton, Artemisia tilesii (Asteraceae) 433, 27♀ (CNC). Engineer Creek, Dempster Highway, July 23, 1983, L.A. Kelton, Rosa sp. (Rosaceae) , 13 (CNC). Gravel Lake, 58 mi E of Dawson, 2050 ft, August 11, 1962, P.J. Skisko, 183, 11♀ (CNC). Junction of Dempster and Klondike Highway, July 20, 1982, L.A. Kelton, 13 (CNC). Moose Creek, July 18, 1982, L.A. Kelton, 13 (CNC). Rancheria, August 11, 1982, L.A. Kelton, Lathyrus sp. (Fabaceae) , 453, 65♀ (CNC). Takahane River, July 20, 1981, C. S. Guppy, 13 (UBC). Tatchun, August 11, 1983, L.A. Kelton, 1♀ (CNC). MEXICO: Durango: 20 mi E of El Salto, Navajos, 8000 ft, July 27, 1964 – August 11, 1964, L. A. Kelton, 613 (CNC). Puebla: San Miguel Texmelucan, August 27, 1969, L. A. Kelton, 13 (CNC). Zacatecas: Guadalupe, July 6, 1969, L. A. Kelton, 13 (CNC). USA: Alaska: Big Delta, July 13, 1951, J.R. McGillis, 13 (CNC). Big Delta, July 26, 1951, W.R.M. Mason, J.R. Mc­ Gillis, 13 (CNC). Chicken, July 22, 1982, L.A. Kelton, Aster sp. (Asteraceae) , 473, 35♀ (CNC). Chitina, August 10, 1934, C. P. Alexander, Artemisia sp. (Asteraceae) , 33 (USNM). Gardiner Creek Camp, Alaska Hiway, August 5, 1978, P. H. Arnaud, Jr., 13 (CAS). Gulkana River, Richarson Hwy, August 2, 1952, 33 (USNM). Isabel Pass, mi 206 Richardson Highway, 2900 ft, July 13, 1962, P. J. Skisko, 13, 1♀ (CNC). Kotzebue, June 26, 1951, R. I. Sailer, 13 (USNM). Lower Tonsina, July 15, 1953, W.C.F., 43 (USNM). Palmer, August 1, 1948, R. I. Sailer, 13 (USNM). 21 mi N of Paxson, July 16, 1987, S. G. Cannings, 1♀ (UBC). Steese Hwy MP 145.2, September 4, 1948, G. Jefferson, 13 (USNM). Teller, July 11, 1951, R. I. Sailer, 103, 10♀ (USNM). Umiat, July 16, 1959, R. Madge, 53, 8♀ (CNC). Umiat, July 17, 1959, J.E.H. Martin, 1♀ (CNC). Umiat, July 18, 1959, R. Madge, 93, 10♀ (CNC). Umiat, July 19, 1959, J.E.H. Martin, 1♀ (CNC). Umiat, July 23, 1959, J.E.H. Martin, 13, 5♀ (CNC). Unalakleet, July 6, 1961, R. Madge, 33, 2♀ (CNC). Unalakleet, July 8, 1961, R. Madge, 53, 2♀ (CNC). Unalakleet, July 10, 1961, R. Madge, 83, 4♀ (CNC). Unalakleet, July 13, 1961, R. Madge, 23, 2♀ (CNC). Unalakleet, July 15, 1961, R. Madge, 143, 7♀ (CNC). Unalakleet, August 5, 1961, B.S. Heming, 13, 5♀ (CNC). Arizona: Apache Co. : Apache National Forest on Hwy 78, 5.4 mi SW Arizona­New Mexico state line, September 20, 1998, J. C. Schaffner, 23, 2♀ (TAMU). Coconino Co.: 7 mi NW Flagstaff, August 13, 1975, J. C. Schaffner, 33 (TAMU). Navajo Co.: Show Low, Sitgreaves National Forest, August 10, 1967, L. A. Kelton, 33, 1♀ (CNC). California: Alameda Co.: Niles Canyon, June 13, 1917, W. M. Giffard, 53 (CAS, USNM). Butte Co.: Oroville, April 30, 1927, H. H. Knight, Artemisia heterophylla (Asteraceae) , 23 (CAS). Fresno Co.: 2 km W of Kaiser Pass on USFS 80, 2600 m, July 25, 1999, M. D. Schwartz, Artemisia sp. (Asteraceae) , 293, 12♀ (CNC). Humboldt Co.: Beatrice, July 10, 1949, B. P. Bliven, 13 (CAS). Blocksburg, October 10, 1937, B. P. Bliven, 43 (CAS). Dinsmores, June 25, 1939, B. P. Bliven, 13 (CAS). Laribee Valley, July 3, 1938, B. P. Bliven, 13 (CAS). Mad River Beach, June 20, 1969, W. G. Goodman, 33 (UCD). Miranda on Hwy 101, June 8, 1952, B. P. Bliven, 13 (CAS). Myers, August 21, 1938, B. P. Bliven, 13 (CAS). Redway, May 29, 1938, B. P. Bliven, 23 (CAS). Inyo Co. : Bishop, July 28, 1940, E. E. Kenaga, 13 (KU). Kern Co.: 5 mi N of Kernville, May 15, 1968, S. Earnshaw, 13 (UBC). Lake Co.: Laurel Dell, August 3, 1916, E. P. Van Duzee, 13 (CAS). Los Angeles Co.: Whitter, February 19, 1935, Artemisia californica (Asteraceae) , 33 (USNM). Mariposa Co. : Yosemite, Nevada Falls, June 7, 1931 – July 29, 1946, E. Essig and R.Usinger, 23 (UBC). Mendocino Co.: 4 mi W of Eel River Ranger Station, Mendocino National Forest, 1450 ft, June 9, 1972, J. Powell, Artemisia douglasiana (Asteraceae) , 83 (UBC). Willits, July 3, 1948, W. F. Chamberlain, 113 (TAMU). Modoc Co.: Hackamore, July 12, 1947, R. Usinger, Chrysothamnus sp. (Asteraceae) , 23 (UBC). Orange Co.: Cleveland National Forest, 1.5 mi E of San Juan Campground, 500 m, May 12, 1978, R. T. Schuh and J. D. Pinto, 113 (AMNH). Cleveland National Forest, El Cariso Campground on Hwy 74, 750 m, May 12, 1978, J.D. Pinto and R. T. Schuh, Artemisia sp. (Asteraceae) , 133 (AMNH). Green River Camp, Lower Santa Ana Canyon , May 11, 1933, E. P. Van Duzee, 33 (CAS). Santa Ana , April 1, 1936, C. E. Norland, 13 (LACM). Plumas Co.: 4 mi W of Quincy, June 19, 1949, P.D. Hurd, 13 (UBC). Riverside Co.: Saboba Springs, June 2, 1917, E. P. Van Duzee, 153 (CAS). Sacramento Co.: Sacramento, April 3, 1920, E. P. Van Duzee, 13 (CAS). Sacramento, August 10, 1935 – May 11, 1960, H. H. Keifer, R. F. Wilkey, Artemisia douglasiana (Asteraceae) , 103 (CAFA). Sacramento, August 10, 1935, H. H, Keifer, Artemisia sp. (Asteraceae) , 13 (CAS). San Diego Co.: San Diego, March 11, 1914 – May 8, 1913, E. P. Van Duzee, 83 (CAS). San Joaquin Co.: No specific locality, May 30, 1917, W. M. Giffard, 23 (CAS). Santa Barbara Co. : 13 mi E of Lompoc, April 21, 1932, E. P. Van Duzee, 13 (CAS). Lompoc, April 21, 1922 – August 6, 1938, R. H. Beamer, E. P. Van Duzee, 23 (KU, CAS). Santa Clara Co. : Felton, June 12, 1952, W. H. Lange, 13 (UCD). Santa Cruz Co. : No specific locality, June 7, 1917, W. M. Giffard, 63 (CAS). Siskiyou Co.: Hornbrook, June 14, 1959, L.A. Kelton, R. Madge, 13 (CNC). 2.5 mi N of Medicine Lake on Medicine Lake Rd, 750 m, July 10, 1985, G. M. Stonedahl and J. D. McIver, 33 (AMNH). Yreka, June 15, 1959, L. A. Kelton, Solidago sp. (Asteraceae) , 13 (CNC). Yreka, June 15, 1959, L.A. Kelton, R. Madge, 113, 24♀ (CNC). Sutter Co.: Nicolaus, July 27, 1935, R. H. Beamer, 23 (KU). Trinity Co.: Coffee Creek, July 30, 1931, R. Usinger, 13 (UBC). Tulare Co.: Woodlake, April 24, 1932, E. P. Van Duzee, Artemisia sp. (Asteraceae) , 23 (CAS). Ventura Co.: Santa Paula , June 28, 1955, W. E. Simonds, 13 (CAFA). Colorado: Chaffee Co.: 20 mi NE of Buena Vista, August 31, 1914, A. K. Fisher, 13 (USNM). Elbert Co.: 10 mi E of Kiowa, August 29, 1982, D. A. Polhemus, 73 (JTP). Jefferson Co.: Red Rocks Park near Morrison, 5600 ft, July 15, 1983, R. T. Schuh, D. A. Polhemus, 13 (AMNH). La Plata Co.: Durango, August 12, 1925, H. H. Knight, 53, 5♀ (USNM). Lake Co.: Leadville, 10,300 ft, August 3, 1919 – August 5, 1919, 13, 3♀ (AMNH). Larimer Co.: Fort Collins, June 7, 1898, 43 (USNM). Montezuma Co. : Mesa Verde Natl. Park , 7000 ft, July 17, 1968, L. A. Kelton, 33, 1♀ (CNC). Pitkin Co.: Aspen, 8000 ft, July 24, 1919, 13 (AMNH). Rio Blanco Co. : 30 mi N of Rifle on Rt 13, 6800 ft, August 14, 1986, R. T. Schuh, Artemisia ludoviciana (Asteraceae) , 1033 (AMNH). Routt Co.: Steamboat Springs, July 12, 1964, H. H. Knight, Artemisia sp. (Asteraceae) , 203, 20♀ (USNM). Saguache Co.: Alder Creek, July 14, 1993, W. F. Chamberlain, 43 (TAMU). Washington Co.: Lindon, June 21, 1937, C. L. Johnston, 63 (KU). Yuma Co. : Joes, June 27, 1937, C. L. Johnston, 13 (KU). Idaho: Franklin Co.: Cub River Canyon, August 12, 1973, G. F. Knowlton, 53 (USU). Idaho Co.: E of Lowell, Wilderness Access Campground (m.p. 123), 2200 ft, August 7, 1986, Schuh, Schwartz, Stonedahl, Artemisia ludoviciana (Asteraceae) , 133 (AMNH). Lochsa River Valley, E of Wilderness Access Campground (mp 127), 2400, August 8, 1986 – August 1, 1987, Schuh, Schwartz, Stonedahl, Artemisia ludoviciana (Asteraceae) , 1473 (AMNH). W of Lowell on Rt 12, 3 Devils Picnic Site, 1700 ft, August 7, 1986, Schuh, Schwartz, Stonedahl, Artemisia ludoviciana (Asteraceae) , 133 (AMNH). Oneida Co. : Ireland Canyon, October 1, 1969, G. F. Knowlton, 23 (USU). Unknown Co.: Silver Cow Camp, July 23, 1991, W. J. Hanson, 23 (USU). Iowa: Dickinson Co.: Cayler Prairie, July 11, 1963, J. C. Schaffner, Artemisia sp. (Asteraceae) , 33 (USNM). Story Co.: Ames, August 9, 1932, F. Andre, 13, 1♀ (USNM). Ames, May 27, 1929, H. H. Knight, 23, 1♀ (USNM). Unknown Co.: Cedar Falls, July 26, 1927, Harris and Johnston, 23 (USNM). Minnesota: Chisago Co.: North Branch, August 5, 1922, H. H. Knight, 403, 40♀ (USNM). Hennepin Co.: Fort Snelling, August 2, 1923, H. H. Knight, 103, 10♀ (USNM). Ramsey Co.: St. Paul, September 3, 1923, H. H. Knight, 103, 10♀ (INHS). Montana: Bighorn Co.: Custer Battlefield, August 4, 1927, H. H. Knight, 23, 2♀ (USNM). Carbon Co.: Rock Creek Vista Pt on Rt 212 NE of Beartooth Summit, 9100 ft, August 11, 1986, Schwartz, Stonedahl, Artemisia michauxiana (Asteraceae) , 343 (AMNH). Gallatin Co.: Willow Creek, June 17, 1921, W. C. Cook, 33, 1♀ (USNM). Nevada: Washoe Co.: Verdi, June 18, 1964 – July 9, 1967, C. N. Slobodchikoff, W. Gagne, 33 (CAS). New Mexico: Grant Co. : Silver City, 1800–1900 m, August 1, 1977, N. Krauss, 13 (AMNH). Lincoln Co.: Ruidoso, June 26, 1940, D. E. Hardy, 23 (KU). Sierra Blanca Mt. , 8600 ft, August 18, 1970, J. R. and M. H. Sweet, 13 (TAMU). Los Alamos Co.: Los Alamos Mortandad Canyon, 2190 m, September 1, 1976 – February 1, 1977, D. C. Lowrie, 403 (AMNH). Los Alamos, Mortandad Canyon, 2190 m, April 1, 1976 – June 1, 1976, D. C. Lowrie, 103 (AMNH). Otero Co.: 2 mi NW of Mayhill, September 26, 1979, R. R. Murray and J. C. Schaffner, 53, 10♀ (TAMU). 4 mi E of Cloudcroft, August 18, 1979 – September 26, 1979, J. C. Schaffner et al., 433 (TAMU). 7 mi N of Cloudcroft, August 13, 1982, J. C. Schaffner, 153, 15♀ (TAMU). 7 mi N of Cloudcroft, August 13, 1982, J. C. Schaffner, 903 (TAMU). Cloudcroft, June 27, 1940, Hardy, Beamer, Kuitert, 323 (KU). Near Cloudcroft, 30 km E of Alamogordo, September 1, 1998, M. D. Schwartz, 13, 1♀ (CNC). North Dakota: Trail Co.: No specific locality, August 19, 1923, A. A. Nichol, 53, 5♀ (USNM). Oregon: Clatsop Co.: Saddle Mountain, 2300–2700 ft, August 11, 1970, Lattin, Oman, 43 (OSU). Crook Co.: 23 mi E of Prineville, June 28, 1988, G. M. Stonedahl, Artemisia arbuscula (Asteraceae) , 563 (AMNH, OSU). Jackson Co.: Medford, TouVelle Park, May 18, 1968, Oman, 13 (OSU). Mt. Ashland, 4.5 mi below ski lodge, 5500 ft, September 24, 1968, J. D. Lattin, Artemisia douglasiana (Asteraceae) , 13 (OSU). Klamath Co.: near Pothole, July 25, 1958, Joe Schuh, Haplopappus sp. (Asteraceae) , 13 (OSU). Lake Co.: 11 mi S of Summer Lake P.O., June 27, 1980, J. D. Lattin, Artemisia ludoviciana (Asteraceae) , 133 (OSU). Linn Co.: Tombstone Prairie, 36 mi E of Sweet Home, 4200 ft, September 12, 1956 – June 27, 1980, G. Eulenson, J. Lattin, P. Oman, J.Schuh, Artemisia ludoviciana (Asteraceae) , 283 (OSU). Marion Co.: 12 mi N of Salem, September 7, 1956, J. D. Lattin, 13 (OSU). Wallowa Co.: Joseph, July 10, 1969 – July 21, 1970, Paul Oman, 33 (OSU). South Dakota: Jackson Co.: Kadoka, August 23, 1927, H. H. Knight, 43 (USNM). Utah: Emery Co.: Woodside, July 14, 1954, G. F. Knowlton, 23 (KU). Kane Co.: Mount Carmel, June 16, 1948, G. F. Knowlton, 13 (USU). Salt Lake Co.: Albion Basin near Alta, 10,000 ft, August 18, 1984, D. A. Polhemus, 63 (JTP). Wasatch Mts., Hidden Peak, August 13, 1987, D. A. Rider, 373 (DAR). Wasatch Mts., Little Cottonwood Canyon, Snowbird Ski Resort, August 19, 1986, M. D. Schwartz, Artemisia ludoviciana (Asteraceae) , 403 (AMNH). Sanpete Co.: 13 mi E of Fairview on Rt 31, T14S R6E, 7500 ft, August 8, 1981, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 23 (AMNH). Sevier Co.: Fish Lake National Forest, N end of Johnson Valley Reservoir, 2725 m, July 16, 1980, R. T. Schuh, G. M. Stonedhal, Artemisia sp. (Asteraceae) , 13 (AMNH). Summit Co.: Coalville, July 13, 1965, H. H. Knight, 53, 5♀ (USNM). Wasatch Co.: Buckboard Creek at Rt 35, Uintah National Forest, 8000 ft, August 16, 1986, M. D. Schwartz and G. M. Stonedahl, Artemisia tridentata (Asteraceae) , 13 (AMNH). Washington: Pierce Co.: DuPont, July 5, 1935, R. H. Beamer, 13 (KU). Yakima Co.: 36 mi S of Toppenish, June 26, 1969, Oman, 13 (OSU). Wisconsin: Trampealeau Co.: near Osseo on Co. Rts G & K,, 800 ft, September 13, 1986, M. D. Schwartz, Artemisia ludoviciana (Asteraceae) , 453 (AMNH). Wyoming: Crook Co.: Sundance, July 20, 1927, H. H. Knight, 33, 1♀ (USNM). Sweetwater Co.: Green River Lake, 8500 ft, August 1, 1935, H. Ruckes, 13 (USNM). Weston Co.: Newcastle, August 19, 1927, H. H. Knight, 2♀ (USNM).

Figures

Fig. 1. Habitus views of Europiella spp.: angulata–decolor. See appendix 1 for localities.

Fig. 3. Male genitalia of Europiella spp.: angulata–consors.

Fig. 6. Europiella angulata (Uhler). Male: USA: Colorado: Jefferson Co.: Red Rocks Park near Morrison, 5600 ft.

Fig. 7. Scanning electron micrographs of Europiella angulata (3). A. Mesothoracic spiracle and metepisternal scent­efferent system, lateral view. B. Pretarsus, lateroventral view. C. Setae on hemelytra. D. Setae on hemelytra, detail of microstructure.