Europiella consors (Uhler)

SCHUH, RANDALL T., 2004, Revision of Europiella Reuter in North America, with the Description of a New Genus (Heteroptera: Miridae: Phylinae), American Museum Novitates 3463, pp. 1-59 : 21-23

Europiella consors (Uhler)

Figures 1 , 3 , 8 , 9

Maurodactylus consors Uhler, 1895: 53 (n. sp., descr.); Schuh et al., 1995: 387 (n. syn., disc.).

Europiella fuscicornis Knight, 1969: 82 (n. sp.; syn. by Schuh et al., 1995: 387).

Europiella basicornis Knight, 1970: 230 (n. sp.; syn. by Schuh et al., 1995: 387).

Europiella monticola Knight, 1970: 230 (n. sp., descr.). NEW SYNONYMY.

Europiella yampae: Kelton, 1980: 341 (misidentification).

Megalopsallus monticola Schuh et al., 1995: 389 (n. comb.).

DIAGNOSIS: Recognized by the relatively large size, elongate, nearly parallel­sided, broad body form, general coloration of dorsum being either dark or light (figs. 1, 8), and the vesica with a long, flat, decurved, anterior apical spine and shorter posterior spine (fig. 3), a condition shared with E. umbrina and E. stigmosa . Most similar in size, general appearance, and coloration to E. angulata , and apparently not separable from it with certainty except by dissection of male genitalia. Sexual dimorphism pronounced, females being much smaller than males and conspicuously ovoid, in contrast to the nearly monomorphic situation found in E. stigmosa and E. umbrina .

REDESCRIPTION: Male: Relatively large, broad­bodied species, total length 3.22–3.68, width pronotum 1.00–1.15. COLORATION (figs. 1, 8): General coloration of dorsum either largely dark or mostly pale; antennal segments 1–4 black; labium mostly black in dark specimens, pale basally in pale specimens; much of venter dark in dark specimens, largely pale in pale specimens; coxae, trochanters, and femora mostly dark in dark specimens, largely pale in pale specimens, femora when pale with dark spots or blotches; tibiae with pale background coloration with black spines with black bases. SUR­ FACE AND VESTITURE (fig. 9C): Surface and vestiture as in generic diagnosis. STRUCTURE (fig. 9): Elongate nearly parallel­sided, corial margin only weakly convex. GENITALIA (fig. 3): Pygophore moderately large relative to size of abdomen; vesica with anterior spine long and decurved, posterior spine shorter and nearly erect, similar to condition found in E. stigmosa and E. umbrina .

Female: Coloration usually light, seldom as dark as in males. Females much shorter than males, ovoid. Total length 2.79–3.26, width pronotum 0.98–1.09.

HOST: Known from various Artemisia spp. , although recorded primarily from A. filifolia and A. tridentata . Also known to breed on Chrysothamnus spp. Records from Cowania and Crotalaria are almost certainly not breeding records.

DISTRIBUTION: Ranging from British Columbia to Saskatchewan in the North , south through the western plains to Texas , and west to California.

DISCUSSION: Examination of topotypic

2000 m.

paratypes of E. monticola Knight suggests that this nominal species is the same as E. consors , and I am therefore treating the former as a junior synonym of the latter new synonymy. Both specimens examined were teneral, and the male genitalia were therefore of no value; nonetheless, judging from the light coloration, which is much more common in E. consors than in E. angulata , the only other species with which monticola could be easily confused, the synonymy with E. consors would appear to be reasonably secure. This taxon was placed in Megalopsallus in error by Schuh et al. (1995).

SPECIMENS EXAMINED: Holotype: 3, Colo., 1744; P. R. Uhler Collection (USNM). Holotypes of synonyms: Europiella fuscicornis Knight : 3, Hudson, Colo., Aug. 25, 1925, H. H. Knight, Artem. filifolia (USNM) . Europiella basicornis Knight : 3, Coalville, UTAH, July 13, 1965, H. H. Knight (USNM). Additional specimens: CAN­ ADA : Alberta: Irvine, June 11, 1952, L. A. Konotopetz, 33 (CNC). British Columbia: Mt. Kobau, 11 km NW of Osoyoos Summit, 1834 m, August 30, 1993, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 383 (CNC). Osoyoos, Mount Kobau Road, August 18, 1996, J. Jarrett (UBC). Osoyoos, Mount Kobau Road, August 18, 1997, J. Jarrett (UBC). Osoyoos, Mount Kobau, 1865 m, August 27, 1988, G.G.E. Scudder (UBC). Osoyoos, Mount Kobau, 1885 m, August 26, 1986, G.G.E. Scudder, Artemisia tridentata (Asteraceae) (UBC). Osoyoos, Mount Kobau, 1885 m, August 27, 1986, G.G.E. Scudder, Artemisia tridentata (Asteraceae) (UBC). Osoyoos, Mount Kobau, May 22, 1996, J. Jarrett (UBC). Osoyoos, Mount Kobau, September 22, 1997, J. Jarrett (UBC). Osoyoos, Mt. Kobau, 1885 m, August 26, 1986 – August 30, 1993, G.G.E. Scudder, Artemisia tridentata (Asteraceae) , 453, 45♀ (UBC, AMNH). Osoyoos, Mt. Kobau, September 22, 1997 – October 25, 1997, J. Jarrett, 33, 3♀ (UBC). Manitoba: Russell, July 17, 1954, Brooks and Wallis, 63, 8♀ (CNC). Saskatchewan: Elbow, June 20, 1960, A. R. Brooks, 1♀ (CNC). Willow Bunch, July 27, 1955, A. R. Brooks, 63, 5♀ (CNC). USA: Arizona: Cochise Co. : 1 mi E of Wilcox, October 20, 1979, J. D. Pinto, 53 (UCR). 4 mi E of Wilcox, May 10, 1987, J. G. Rosen, 13 (AMNH). Coconino Co.: Grand Canyon, 7000 ft, June 22, 1925, A. A. Nichol, 13 (CNC). Grand Canyon, 7000 ft, June 22, 1925, A. A. Nichol, 13 (USNM). Rt 64 just SE of Grand Canyon Natl. Park in Kaibab National Forest, June 26, 1980, K. and R. Schmidt, Artemisia sp. (Asteraceae) , 203 (AMNH). Navajo Co.: Holbrook, June 25, 1980, J. D. Pinto, 13 (UCR). Pinal Co.: Superior, 2400 ft, April 16, 1928, A. A. Nichol, 23 (USNM). Yavapai Co.: near Congress, Aguila, May 20, 1960, B. P. Bliven, 13 (CAS). California: Inyo Co. : Mammoth Lakes, June 29, 1940, L. C. Kuitert, 13 (KU). Kern Co.: 11 mi NW of Tehachapi on Rt 58, April 25, 1983, G. M. Stonedahl, Chrysothamnus nauseosus (Asteraceae) , 213 (AMNH). Los Angeles Co.: 1 mi S of Gorman, June 13, 1964, A. G. Raske, 13 (UCB). Mono Co.: Mammoth Camp, 33 (LACM). San Diego Co.: Morena Dam, May 26, 1929, C. C. Sarl, 13 (SDNM). Siskiyou Co.: 5 mi N of Gazelle, June 15, 1959, Kelton and Madge, Crotalaria sp. (Fabaceae) , 93, 18♀ (CNC). Tulare Co.: 21 mi W of Rt 395 toward Kennedy Meadows, 1875 m, July 1, 1980, R. T. Schuh, Chrysothamnus parryi (Asteraceae) , 43 (AMNH). Ventura Co.: Ventura, January 23, 1919, E. O. Essig, Artemisia californica (Asteraceae) , 13 (UCB). Colorado: Archuleta Co.: Piedra, San Juan National Forest, 6300 ft, July 22, 1968, L. A. Kelton ( Asteraceae ), 63, 8♀ (CNC). Baca Co. : 3 mi S of Walsh, May 23, 1947, M. T. James, 153, 5♀ (WSU). Clear Creek Co.: Berthoud Pass, July 12, 1949, J. R. White , 13 (KU). Conejos Co.: Elk Creek Recreation Area, August 22, 1969, J. C. Schaffner, 73 (TAMU). Douglas Co.: Roxborough Road near Chatfield Park, May 28, 1978, J. T. Polhemus, 13 (JTP). Eagle Co.: 1 mi N of Minturn, Grouse Creek, 7840 ft, August 2, 1986 – August 11, 1987, T. J. Henry, J. T. and D. A. Polhemus, Artemisia tridentata (Asteraceae) , 143, 8♀ (USNM, JTP). McCoy, August 22, 1968, L. A. Kelton, 103, 8♀ (CNC). N of Minturn, Grouse Creek Trail, 7840 ft, August 11, 1986, J. T. and D. A. Polhemus, Artemisia tridentata (Asteraceae) , 43 (JTP). Sloss, August 17, 1929, L. D. Anderson, R. H. Beamer, 43 (KU). Wolcott, August 1, 1932 – August 24, 1935, G. F. Knowlton, Vasco Tanner, 23 (USU, BYU). Garfield Co.: W Evacuation Creek, 11 mi SE state line on Rt 45, 6800 ft, July 9, 1981, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 23 (AMNH). Grand Co.: Granby, August 19, 1968, L. A. Kelton, 23 (CNC). Parshall, August 17, 1973, W. J. Hanson, G. F. Knowlton, 13 (USU). Gunnison Co.: 35 mi E of Gunnison on Rt 50, 8500 ft, July 17, 1986, R. T. Schuh, Artemisia tridentata (Asteraceae) , 33 (AMNH). 5 mi NE of Almont, along Taylor River, 8400 ft, August 13, 1987, J. T. and D. A. Polhemus, T. J. Henry, Artemisia tridentata (Asteraceae) , 303, 15♀ (JTP, USNM). 5 mi NE of Almont, Taylor River Bridge, 8400 ft, August 13, 1987, T. J. Henry, Artemisia tridentata (Asteraceae) , 253, 10♀ (USNM). Black Mesa , 10.7 mi W of Sapinero, August 19, 1966, T. C. Emmel, 13 (LACM). Gunnison, August 17, 1925, H. H. Knight, Chrysothamnus sp. (Asteraceae) , 153, 15♀ (USNM). Gunnison, August 17, 1925, H. H. Knight, Chrysothamnus sp. (Asteraceae) , 23 (CNC). Parlin, August 27, 1968, L. A. Kelton, 163, 5♀ (CNC). Huerfano Co.: Veta Pass, August 9, 1925, H. H. Knight, 33 (USNM). Jackson Co.: 1 mi E of Gould, August 13, 1969, J. C. Schaffner, 413 (TAMU). 7 mi NE of Rand, August 14, 1969, J. C. Schaffner, 603 (TAMU). Cowdrey, 8000 ft, August 15, 1968, L. A. Kelton ( Asteraceae ), 503, 25♀ (CNC). Gould, 9200 ft, August 12, 1968, L. A. Kelton, Artemisia sp. (Asteraceae) , 53, 5♀ (CNC). Muddy Pass, Routt National Forest, 8600 ft, August 19, 1968, L. A. Kelton, 163, 2♀ (CNC). Northgate, August 20, 1931, R. H. Beamer, 13 (KU). Rabbit Ears Pass, 9680 ft, July 11, 1964, H. H. Knight, 23 (USNM). Rabbit Ears Pass, July 11, 1964, H. H. Knight, 13 (CNC); paratype of Europiella monticola Knight. Walden , August 20, 1931, R. H. Beamer, 23 (KU). Jefferson Co.: Golden, 6000 ft, August 6, 1919, 13 (AMNH). Lake Co.: Leadville, 10,300 ft, August 3, 1919, 83 (AMNH). Larimer Co.: 3 mi S of Estes Park, August 2, 1997 – August 4, 1997, J. C. Schaffner, 13 (TAMU). Estes Park, 8800 ft, August 27, 1920, H. C. Severin, 53 (USNM). Estes Park, August 24, 1925, H. H. Knight, 13 (USNM); paratype of Europiella fuscicornis Knight ; Hudson, August 25, 1925, H. H. Knight, 23 (USNM). Rocky Mountain National Park, August 5, 1947, R. H. and L. D. Beamer, 23 (KU). Rocky Mountain National Park, Grand Lake Entrance, August 18, 1968, L. A. Kelton, 13 (CNC). Rustic, Poudre Canyon, September 6, 1972, L. A. Kelton, 33, 1♀ (CNC). Logan Co.: Sterling, August 25, 1925, H. H. Knight, 23 (USNM). Moffat Co.: Craig, July 14, 1949, R. H. Beamer, 13 (KU). Montezuma Co. : Dolores, August 15, 1925, H. H. Knight, 53, 5♀ (USNM). Montrose Co.: Black Canyon, August 15, 1973, J. C. Schaffner, 13 (TAMU). Park Co.: Grant, Geneva Park , 9500 ft, July 22, 1916, L. O. Jackson, 13 (USNM). Pitkin Co.: Aspen, White River National Forest, August 24, 1968, L. A. Kelton, 253, 12♀ (CNC). Pueblo Co.: Avondale, May 10, 1981, J. T. Polhemus, 513 (JTP). Rio Blanco Co. : 30 mi N of Rifle on Rt 13, 6800 ft, August 14, 1986, R. T. Schuh, Artemisia tridentata (Asteraceae) , 113 (AMNH). Routt Co.: Steamboat Springs, July 15, 1964, H. H. Knight, 23 (USNM). Steamboat Springs, September 1, 1943, O. B., 13 (KU). Saguache Co.: 1 mi E of North Pass, August 19, 1969, J. C. Schaffner, 483 (TAMU). 23 mi E of Gunnison on Rt 50, 8100 ft, July 17, 1986, R. T. Schuh, Artemisia tridentata (Asteraceae) , 443 (AMNH). Valley View Springs, ca. 7 mi E Mineral Hot Springs, 8500 ft, August 14, 1965, H. B. Leech, 13 (CAS). Summit Co.: Dillon, August 29, 1972, L. A. Kelton, Artemisia sp. (Asteraceae) , 53, 3♀ (CNC). Sutter Co.: Nicolaus, June 27, 1935, R. H. Beamer, 13 (KU). Weld Co.: 10 mi E of Roggen, July 16, 1983, D. A. Polhemus, 13 (JTP). 10 mi N of Hodson, June 11, 1979, D. A. Polhemus, 13 (JTP). 2 mi SE of Masters, June 4, 1979, D. A. Polhemus, 23 (JTP). 24 mi N of Stoneham along Hwy 71, June 5, 1979, D. A. Polhemus, 23 (JTP). 5 mi NW of Roggen, July 15, 1983, D. A. and J. T. Polhemus, 33 (JTP). 5 mi W of Roggen, 5200 ft, July 16, 1983, R. T. Schuh, D. A. and J. T. Polhemus, Artemisia filifolia (Asteraceae) , 93 (AMNH). 7 mi E of Nunn, August 15, 1968, Oman, 13 (OSU). Riverside Reservoir, June 4, 1979, D. A. Polhemus, 23 (JTP). Idaho: Franklin Co.: Cub River Canyon, August 13, 1974, G. F. Knowlton, 13 (OSU). Montana: Deer Lodge Co.: Silver Lake, 14 mi W of Anaconda on Rt 10A, 6400 ft, August 9, 1986, Schuh, Schwartz, Stonedahl, Artemisia ludoviciana (Asteraceae) , 663 (AMNH). Fergus Co.: 6 mi S of Grass Range, July 6, 1972, G. C. Gaumer, 23 (TAMU). Gallatin Co.: Targhee Pass on Rt 191, 7072 ft, August 10, 1986, Schuh, Schwartz, Stonedahl, Artemisia tridentata (Asteraceae) , 523 (AMNH). Nevada: Carson City Co.: Carson City, Summit Clear Creek Grade, July 10, 1934, E. P. Van Duzee, 13 (CAS). Elko Co.: Ruby Mts., Lomoille Canyon, 6200 ft, June 16, 1983, R. T. Schuh, M. D. Schwartz, Artemisia dracunculus (Asteraceae) , 233 (AMNH). Esmeralda Co. : 2 mi W of Lida, 6500 ft, August 6, 1966, W. C. Gagne, 13 (UCB). 3 mi W of Lida, 6800 ft, July 7, 1966, C. W. O’Brien, 13 (UCB). Lander Co.: Austin Summit, July 21, 1968, J. Scott, 13 (UCB). Washoe Co.: Reno, June 27, 1927, E. P. Van Duzee, 13 (CAS). Reno, Sky Ranch, June 16, 1952, E. I. Schlinger, 13 (UCB). White Pine Co. : 2.3 mi N of Hwy 50 on Steptoe Creek Rd, 6800 ft, July 19, 1980, G. M. Stonedahl, Chrysothamnus viscidiflorus (Asteraceae) , 183 (AMNH). Wheel­ er Peak Drive, Humboldt National Forest, 7000– 10,000 ft, August 6, 1982, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 13 (AMNH). New Mexico: Bernalillo Co.: Albuquerque, June 1, 1983, J. T. Polhemus, 93 (JTP). Tijeras, along I­40, June 1, 1983, J. T. Polhemus, 63 (JTP). Dona Ana Co.: 4 mi W of Las Cruces, April 10, 1976, Oman, 33 (OSU). Eddy Co.: Site 1, May 23, 1979, Delorme and Carrola, 73 (TAMU). Site 11, April 24, 1979, Burke, Delorme, Schaffner, 1613 (TAMU). Site 13, April 24, 1979, Burke, Delorme, Schaffner, 93 (TAMU). Site 7, September 23, 1979 – June 2, 1979, Murray, Schaffner, Burke, Delorme, Carrola, Friedlander, Artemisia filifolia (Asteraceae) , 273 (TAMU). Hidalgo Co.: Lordsburg, May 10, 1939, G. Willette, 13 (LACM). Otero Co.: Cloudcroft, June 27, 1940 – June 17, 1976, Lipovsky, Wickham, Oman, 93 (KU, CAS,OSU). Taos Co.: Taos, August 20, 1927, P. A. Readio, 13 (KU). North Dakota: Ramsey Co.: Roosevelt Natl. Mem. Park, N. Unit, June 30, 1973, L. A. Kelton, Artemisia sp. (Asteraceae) , 33, 6♀ (CNC). Oregon: Crook Co.: Ochoco National Forest, July 29, 1967, P. Oman, 23 (OSU). SE of Prineville near Eagle Rock, July 2, 1968, P. Oman, 23 (OSU). Deschutes Co.: 0.5 mi S of Millican, June 29, 1978, Lattin, Artemisia tridentata (Asteraceae) , 23 (OSU). 1 mi S of Millican, 4180 ft, June 21, 1979, R. T. Schuh, Artemisia tridentata (Asteraceae) , 13 (AMNH). 3 mi W of Millican, sand dunes, July 6, 1977, J. D. Lattin ( Asteraceae ), 113 (OSU). Grant Co. : 1 mi S of Silvies, Malheur National Forest, T18S, R32E, Sec. 7, 4580 ft, July 23, 1979, M. D. Schwartz, Chrysothamnus nauseosus (Asteraceae) , 283 (AMNH). Harney Co.: Fish Lake, Steens Mts., 7200 ft, August 14, 1958 – August 18, 1958, K.M. Fender, 283 (OSU). T36S, R35E, Sec. 8, dunes, June 27, 1979, Lightfood and Cobb, 13 (OSU). Klamath Co.: 28 mi SE Jct 97 31, July 25, 1979, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 13 (AMNH). Lake Co.: 11 mi NW of Silver Lake on Hwy 31, 4500 ft, June 25, 1979, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 63 (AMNH). 5.5 mi W of Silver Lake, 4955 ft, July 13, 1978, Oman, 43 (OSU). Wallowa Co.: Joseph, July 10, 1969, P. Oman, 53 (OSU). Texas: Ward Co.: Monahans Sandhills St. Pk., April 12, 1971, R. R. Murray, 103 (TAMU). Utah: Box Elder Co. : Devil’s Gate, July 28, 1969, T. L. Whitworth, 23 (USU). South Curlew Valley, June 18, 1969, G. F. Knowlton, 13 (USU). Cache Co.: 0.4 mi off Rt 189 around Franklin Basin Road, 6750 ft, July 18, 2001, T. J. Henry, Artemisia tridentata (Asteraceae) , 133, 3♀ (USNM). Beaver Mountain, August 31, 1982, G. F. Knowlton, 13 (USU). Elk Valley, July 29, 1976, Hanson, 13 (USU). Franklin Basin, August 13, 1974, Hanson, Knowlton, 13 (USU). Tony Grove Lake, Campground, 8400 ft, August 17, 1976 – September 7, 1981, M. D. Schwartz, G. F. Knowlton, Artemisia tridentata (Asteraceae) , 403 (UCD, USU, AMNH). Daggett Co.: Red Springs Camp, Ashley National Forest, August 25, 1966, C. W. O’Brien, 13 (UCB). Duchesne Co.: 10 mi W of Duchesne, July 20, 1962, J. B. Karren, 13 (USU). Emery Co.: 2.5 mi W of Rt 24 on Goblin Valley Road, 5500 ft, June 19, 1983, R. T. Schuh and M. D. Schwartz, Artemisia filifolia (Asteraceae) , 893 (AMNH). Garfield Co.: 8.7 mi S of Rt 95 on Rt 276, Maidenwater Spring, 5000 ft, June 19, 1983, R. T. Schuh and M. D. Schwartz, Artemisia filifolia (Asteraceae) , 213 (AMNH). Bryce Canyon National Park, V. M. Tanner, 13 (USNM). Hatch, August 11, 1972, L. A. Kelton, 13, 4♀ (CNC). Jct Rts 95 and 276, 4900 ft, June 19, 1983 – June 18, 1983, R. T. Schuh and M. D. Schwartz, Artemisia filifolia (Asteraceae) , 483 (AMNH). Widtsoe, V. M. Tanner, 13 (USNM). Grand Co.: 5 mi S of Moab on US 163, 4600 ft, May 27, 1978, D. A. and J. T. Polhemus, 13 (JTP). Iron Co. : 1 mi N of Parowan, June 18, 1967, Artemisia tridentata (Asteraceae) , 13 (USNM). 7 mi S of Beryl Junction, June 15, 1967, Artemisia tridentata (Asteraceae) , 13 (USNM). Juab Co.: Nephi, June 22, 1966, Knowlton, 13 (USU). Kane Co.: Long Valley Jct, July 31, 1975, W. F. Chamberlain, 13 (TAMU). Millard Co.: 4 mi N of Beaver, July 16, 1967, H. R. Burke, 23 (TAMU). Rich Co.: Allen Canyon, August 27, 1975, G. F. Knowlton, 13 (USU). Logan Canyon Summit, 7800 ft, July 30, 1940 – August 29, 1980, G. S. Stains, D. G. Hall, G. Knowlton, 153 (OSU, USU). San Juan Co.: 27 mi E of Blanding at MP 101 on Rt 95 near Mule Tower Ruins, 6500 ft, June 12, 1982, M. D. Schwartz, Artemisia ludoviciana (Asteraceae) , 13 (AMNH). Brush Basin Rim Rd., Co. rd. no. 227 0.5 E mp 116, 5700 ft, June 12, 1982, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 173 (AMNH). Canyonlands Natl. Park, Grandview Point, 6400 ft, May 22, 1981, M. D. Schwartz, Cowania mexicana (Rosaceae) , 13 (AMNH). Grand Flat near Collins Canyon, 5600 ft, June 1, 1982, D. A. and J. T. Polhemus, 53 (JTP). Near Halls Crossing, rim of Moki Canyon, 4000 ft, May 28, 1978, D. A. and J. T. Polhemus, 393 (JTP). Sanpete Co.: Ephraim Canyon, August 13, 1975, G. F. Knowlton, 13 (USU). 13 mi E of Fairview on Rt 31, T14S, R6E, 7500 ft, August 8, 1981, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 43 (AMNH, OSU). Sevier Co.: 11 mi W of Sevier on Rt 4, 2000 m, July 15, 1980, G. Stonedahl, Artemisia tridentata (Asteraceae) , 503 (AMNH). 24.7 mi N Hwy 24 and Hwy 72, 7960 ft, July 17, 1980, G. Stonedahl, 63 (AMNH). Monroe, October 20, 1929, D. E. Fox, Artemisia tridentata (Asteraceae) , 13 (USNM). Richfield, July 8, 1930, 13 (USU). Richfield, July 8, 1930, E. W. Davis, 103 (USNM). Salina, June 25, 1954, G. Nielsen, 13 (USU). Salt Gulch, 12 mi N Hwy 24 on Hwy 72, 8060 ft, July 17, 1980, G. Stonedahl, Artemisia sp. (Asteraceae) , 113 (AMNH). Summit Co.: Kimball Junction, August 31, 1967, L. A. Kelton, 153, 10♀ (CNC). Weber Canyon Road, 4.5 mi E of Oakley, August 19, 1986, G. M. Stonedahl, Artemisia tridentata (Asteraceae) , 23 (AMNH). Uintah Co.: 20–22 mi N of Vernal, August 25, 1966, C. W. O’Brien, 23 (UCB). Kane Hollow on Red Cloud Loop, 8500 ft, September 7, 1982, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 53 (AMNH). Unknown Co.: Bone Draw, July 13, 1936, G. F. Knowlton, 13 (USNM). Current Creek, August 16, 1935, G. F. Knowlton, 13 (USU). Utah Co.: Spanish Fork Canyon, July 17, 1954, G. F. Knowlton, 13 (USNM). Wasatch Co.: Daniels Canyon summit, August 16, 1935, G. F. Knowlton, Artemisia sp. (Asteraceae) , 33 (USU). Uintah National Forest, Wolf Creek Campground, T4S, R10W, Sec. 7, 30 mi SE of Kamas on Rt 35, 9000 ft, August 15, 1986, M. D. Schwartz and G. M. Stonedahl, Artemisia tridentata (Asteraceae) , 133 (AMNH). Uintah National Forest, Buckboard Creek at Rt 35, T4S R9E Sec 17, 8000 ft, August 16, 1986, Schwartz, Stonedahl, Artemisia tridentata (Asteraceae) , 193 (AMNH). Washington Co.: 2 mi NW of Toquerville on Rt 17, 3800 ft, April 25, 1981, M. D. Schwartz, Artemisia filifolia (Asteraceae) , 203 (AMNH). Hurricane, May 18, 1960, B. P. Bliven, 93 (CAS). La Verkin, V. M. Tanner, 13 (USNM). La Verkin, Vasco Tanner, 23 (CAS). Leeds, April 27, 1971, G. F. Knowlton, 13 (USU). Rt 15 at I­15 NE of St. George, 935 m, April 18, 1978, R. T. Schuh, Chrysothamnus sp. (Asteraceae) , 1653 (AMNH). Santa Clara , April 26, 1935, G. F. Knowlton, 23 (USU). Snow Canyon State Park, T41S R16W, 4000 ft, April 22, 1981 – April 23, 1981, M. D. Schwartz, Artemisia filifolia (Asteraceae) , 413 (AMNH). St. George, June 30, 1965, H. H. Knight, 43, 5♀ (USNM). Weber Co.: 6 mi S of Monte Cristo, August 17, 1979, G. F. Knowlton (USU). Wyoming: Albany Co.: 5 mi W of Woods Landing, August 15, 1968, L. A. Kelton ( Asteraceae ), 103, 15♀ (CNC). Near Keystone, July 26, 1981, J. T. and D. A. Polhemus, 13 (JTP). Big Horn Co.: 21 mi W of Burgess Jct on Rt 14, 8500 ft, August 12, 1986 – August 13, 1986, Schuh, Schwartz, Stonedahl, Artemisia tridentata (Asteraceae) , 543 (AMNH). 27 mi W of Burgess Jct on Rt 14, 7700 ft, August 12, 1986, Schwartz, Stonedahl, Schuh, Artemisia tridentata (Asteraceae) , 863 (AMNH). Carbon Co.: 11 mi S of Red Lodge, Parkside Campground, 7250 ft, August 12, 1986, M. D. Schwartz and G. M. Stonedahl, Artemisia tridentata (Asteraceae) , 43 (AMNH). Saratoga, June 29, 1931, H. H. Knight, 33 (USNM). Fremont Co.: Wind River Mts., 2.5 mi SW Shoshone NF boundary on Rt 131, August 14, 1986, M. D. Schwartz and G. M. Stonedahl, Artemisia tridentata (Asteraceae) , 63 (AMNH). Lincoln Co.: Auburn, C. Lynn Hayward, 33 (BYU). Park Co.: Shoshone National Forest, August 7, 1927, H. H. Knight, 33 (USNM). Yellowstone National Park, August 8, 1927, H. H. Knight, 23 (USNM). Sweetwater Co.: Green Riv­ er, August 3, 1937, Drake and Andre, 13 (USNM). Teton Co.: Moran, July 19, 1936, G. F. Knowlton, 13 (USU).

Figures

Fig. 1. Habitus views of Europiella spp.: angulata–decolor. See appendix 1 for localities.

Fig. 3. Male genitalia of Europiella spp.: angulata–consors.

Fig. 8. Europiella consors (Uhler). Male: USA: Utah: Sevier Co.: 11 mi W of Sevier on Rt 4,

Fig. 9. Scanning electron micrographs of Europiella consors (3). A. Mesothoracic spiracle and metepisternal scent­efferent system, lateral view. B. Pretarsus, ventral view. C. Setae on hemelytra.