Orictites charleshuberi, Balkenohl, 2017

Balkenohl, Michael, 2017, Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini), Contributions to Natural History 35, pp. 1-68 : 57-60

Orictites charleshuberi sp. nov.

Type material: Holotype: 1 ♀, labels: beige, black printed and handwritten: “ Karapin Near Ari Formosa . 10–V 1938 Coll. Yoshio Yano” / beige, handwritten in black ink: “Karapin Mt. Ari (950 m.)” / beige, black printed: “ H.E. Andrewes Coll. B.M.1945—97.” / backside, handwritten in black ink: “2541” ( BMNH).

Paratype: 1 ♀, labels: white, printed “TAIWAN P.o.CHINA PULI – YÜCHIH SUN MOON LAIE 16.6.–26 .6.1993 J. DALIHOD lgt.“ ( CBP) .

Remark: In the paratype the right antennae is missing from joint two onwards.

Description ( Figs 13 , 40 ; pp. 11, 37)

Measurements (n = 2): Body length 5.8/ 5.8 mm, width 1.65/ 1.68 mm; ratio length/width of pronotum 0.98/0.95; ratio length/width of elytra 1.84/1.86.

Colour: Shiny. Black to dark-brown. Wings of clypeus, supraantennal plates and labrum dark-brown, slightly transparent at margins, antennae brown, palpi yellowish brown, legs red-brown.

Head: A quarter narrower than pronotum. Clypeus nearly fused with wings, with small rounded tooth laterally, anterior margin slightly excised, with blunt raised knob between the two clypeal seatae, margin of wings convex, slightly hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to mid-eye level, shiny but with indistinct irregular reticulation, with reflexed rounded margin, elongated as supraorbital carina, frons separated from clypeus by flat transverse depression, depression with broad band of big punctures, elongated V-like on frons, with costate, nearly parallel running frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina. Frons moderately convex, with very few punctures. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes of moderate size, flattened but still convex, genae enlarged, regularly rounded, not as high as and shorter than eyes, enclosing eyes posteriorly, forming rectangular angle at neck. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked as distinct step, with broader band of big punctures, interrupted at middle. Neck laterally covered with reticulation and densely scattered with punctures of medium size. Labrum indistinctly trilobed, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, shorter than head, flattened, stout, irregularly reticulated, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2 nd segment, 2 nd segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck covered with bigger sized punctures and transverse reticulation, Submentum and mentum distinctly separated; submentum with four bigger, about equally spaced setigerous punctures; mentum at base bilaterally with broad and rounded tubercle, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming broad obtuse triangle, bisetose, not projecting as far as lateral lobes. Paragenae sharp, carina-like, with distinct tooth anteriorly, rounded at base. Antennae relatively long, reaching up to beginning of posterior third of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with isodiametric reticulation, segments 5–10 moniliform, antennomeres somewhat flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with a few additional long setae.

Pronotum: Square, nearly as long as wide, sides slightly convex at middle, rounded in anterior tenth to anterior angles. Anterior margin nearly straight. Reflexed lateral margin distinctly crenulated. Margin from posterior angle to base running as straight line, with small bilateral tubercle at base. Lateral channel broad between the two lateral setigerous punctures, with isodiametric reticulation and some punctures. Anterior setigerous puncture located at the end of anterior quarter adjoining convexity of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angles slightly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as distinct tooth, projecting antero-laterally. Anterior transverse line relatively fine, formed by partly connected punctures. Median line moderately deep, reticulated, surpassing anterior transverse line without joining, smaller at base, joining base. Surface with smaller and bigger punctures, with isodiametric reticulation laterally and at base, basal impression indistinct, basal channel distinct, broad and deep. Flange raised keel-like (lateral view).

Elytra: Subcylindrical. Sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles. Reflexed margin with distinct crenulation in anterior half, margin thickened at humerus, bending with channel over rounded humerus up to 5 th stria. Humeral tooth big, sharp. Basal tubercle distinct, with setigerous puncture, situated at declivity of 1 st stria. Striae 1–4 free at base, striae deep, punctate-striate, striae 1–2, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura somewhat raised, reticulated. Intervals moderately convex, 6 th to 8 th carinate, 3 rd and4 th slightly raised at base, both with tubercle at base. Intervals shiny, with isodiametric reticulation laterally and at base. Interval 3 with four setigerous punctures adjoining stria 3.

Hind wings: Fully developed.

Ventral surface: Epipleuron shiny, covered with big, scattered punctures. submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle like an inverted V, mesosternum smooth. Sternites with transverse to irregular reticulation, each with bigger punctures laterally, 3 rd to 5 th with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.

Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, with indistinct reticulation, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.

Male genitalia: Unknown.

Female genitalia ( Fig. 40 ): Coxostylus slightly sigmoid, gently curved to apex, apex broad and with a pair of setae close together, with five longer nematiform and one slender ensiform seta at middle.

Variation: In the paratype, striae 1 and 2 adjoining indistinctly at apex.

Diagnosis: A medium-sized nearly black species with distinct knob on the clypeus and distinct transverse depression with punctures behind in form of the letter V, with distinct and sharp humeral tooth, with tubercles at the base of intervals 3 and 4 of the elytra, and four setigerous punctures on interval 3. Distinguished from all other species by the crenulation of the lateral margin of the pronotum, and of the elytra in the anterior half.

Distribution: Known from Taiwan only. The paratype was collected at an altitude of 950 m.

Etymology: The name is a patronym of Dr. Charles Huber, former curator at the Naturhistorisches Museum der Burgergemeinde Bern ( Switzerland) and specialist in Nebriinae who supported my work significantly over the last decade.

Figures

Figs 13–14: Orictites spp., habitus. 13: O. charleshuberi sp. nov., holotype, female; 14: O. anteriorlatus sp. nov., holotype, male.

Figs 39–40: Orictites spp., coxostylus. 39: O. tubercucollis sp. nov., scale bar 0.1 mm; 40: O. charleshuberi sp. nov., scale bar 0.1 mm.