Orictites anteriorlatus, Balkenohl, 2017

Balkenohl, Michael, 2017, Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini), Contributions to Natural History 35, pp. 1-68 : 60-62

Orictites anteriorlatus sp. nov.

Type material: Holotype: ♂, labels: white, black printed: “Mt. Lompo Battang S.C. Sulawesi 25–XII–1999 M. ANDO” / blue, black printed: “K. ANDO Collection” ( CBM-ZSM).

Remark: In the holotype the left hind tibia is missing.

Description ( Figs 14 , 30 ; pp. 11, 25)

Measurements: Body length 5.8 mm, width 1.6 mm; ratio length/width of pronotum 0.99; ratio length/width of elytra 1.94.

Colour: Shiny. Black to dark-brown. Wings of clypeus and supraantennal plates brown and slightly transparent, antennae, labrum and mandibles brown, palpi yellowish, legs red-brown.

Head: Wide, a sixth narrower than pronotum. Clypeus with small rounded indistinctly projecting tooth laterally, separated from wings, anterior margin excised, with blunt raised broad knob between the two clypeal setae, margin of wings convex, hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to mid-eye level, smooth, with reflexed rounded margin, elongated as supraorbital carina. Frons separated from clypeus by transverse furrow interrupted at middle, sharp furrow between frons and frontal carina, frons moderately convex. Frons and clypeus smooth. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle behind genae level. Eyes well developed, convex, genae distinct, regularly rounded, not as high as eyes, enclosing eyes posteriorly by one third, forming nearly right angle at neck. Grooves for reception of antennae of moderate length, 0.5 of eye diameter. Neck constriction distinct, with band of big punctures. Neck laterally covered with reticulation and some medium sized punctures. Labrum straight, 7-setose, ciliate laterally, with fine isodiametric reticulation. Mandibles robust, length three quarters of head, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles rounded towards base. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2 nd segment, 2 nd segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck covered with punctures and reticulation. Submentum and mentum separated; submentum with four, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae of moderate length, reaching up to middle of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine isodiametric reticulation, segments 5–10 transverse (L/ W 0.73), antennomeres slightly flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with a few additional long setae.

Pronotum: Square, wider than long, outline reminescent of the inverted frustum of a pyramid, sides straight, convex in anterior quarter for a short distance to anterior angles. Anterior margin distinctly bisinuate. Reflexed lateral margin with traces of scars (120x). Margin from posterior angle to base running as a straight line. Lateral channel disctinctly broadened between the two lateral setigerous punctures, with isodiametric reticulation and few punctures. Anterior setigerous puncture located at the end of anterior fifth adjoining convexity of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angles distinctly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as blunt tooth. Anterior transverse line narrow, sharp, formed by punctures. Median line deep, relatively narrow, rugose, surpassing anterior transverse line but not joining, joining base. Surface with a few punctures and transverse rugae, basal impression indistinct. Basal channel moderately broad, deep. Basal part of disc elongated posteriorly and bilaterally at declivity as tooth-like vault slightly hanging over basal channel (lateral view). Flange raised keel-like (lateral view).

Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles and with a second row of punctures. Reflexed margin with a few indistinct scars posterior to humerus (120x). Margin bending with channel over rounded humerus up to 5 th stria. Humerus without tooth. Basal tubercle distinct, with setigerous puncture, situated at declivity of 1 st stria. Striae 1–4 free at base, all striae deep, indistinctly punctate, striae 1–2, 3–4, and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct. Intervals distinctly convex, 3 rd and4 th distinctly raised at base, both with tubercle at base, 8 th carinate. All intervals shiny on disc. Interval 3 with four setigerous punctures adjoining stria 3.

Hind wings: Fully developed.

Ventral surface: Epipleuron shiny, with some punctures, submarginal furrow distinct, with transverse rugae. Proepisternum and episternum with a few punctures, prosternite nearly smooth, keeled at middle, mesosternum smooth. Sternites with few punctures laterally, 3 rd to 5 th with paralateral ambulatory setae at each side, ventral strigae distinct, with transverse reticulation, sternite 6 smooth at middle, with two widely separated apical setae at each side.

Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.

Male genitalia ( Fig. 30 ): Relatively long, distinctly sclerotized. Median lobe slender, moderately arcuate, slightly twisted apically. Endophallus strongly folded, teeth not visible. Parameres of about equal length, both slender, slightly twisted, asetose.

Female genitalia: Unknown.

Variation: No bilateral variation observed.

Diagnosis: A medium-sized almost black species, with blunt broad knob on the clypeus, humerus without tooth, and four setigerous punctures on interval 3 of the elytra. Only in this species and in O. mjoebergi the basal part of the disc of the pronotum is posteriorly elongated bilaterally at the declivity as a tooth-like vault slightly hanging over the basal channel. In contrast to O. mjoebergi , O. anteriorlatus sp. nov. exhibits tubercles at the base of intervals 3 and 4 of the elytra, the furrow between clypeus and frons is interrupted at middle, and clypeus and frons are impunctured.

Distribution: Known from the South of Sulawesi.

Etymology: The name refers to the anteriorly widened pronotum.

Figures

Figs 13–14: Orictites spp., habitus. 13: O. charleshuberi sp. nov., holotype, female; 14: O. anteriorlatus sp. nov., holotype, male.

Figs 27–30: Orictites spp., aedeagus, ventral view. 27: O. brancuccii sp. nov., holotype, scale bar 0.2 mm; 28: O. barclayi minorfrater ssp. nov., holotype, scale bar 0.2 mm; 29: O. tubercucollis sp. nov., holotype, scale bar 0.2 mm; 30: O. anteriorlatus sp. nov., holotype, scale bar 0.2 mm.