Orictites minotaur ANDREWES, 1931

Balkenohl, Michael, 2017, Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini), Contributions to Natural History 35, pp. 1-68 : 14-18

Orictites minotaur ANDREWES, 1931

Catalogue

Orictites minotaur ; Andrewes 1931: 440;

Orictites minotaur ANDREWES ; Csiki 1933: 638;

Orictites minotaur ANDREWES ; Louwerens 1964: 174;

Orictites minotaur ANDREWES ; Lorenz 1998: 135;

Orictites minotaur ANDREWES ; Balkenohl 2001: 18.

Type material: Lectotype (by present designation): ♂, with labels and data: round, white, with red border, “Type” / white and yellow: “Mt. Kinabalu, Brit.N.Borneo. B.M.1931–386.” / white, handwritten: “ Orictites minotaur Type Andr.” and printed “H.E. Andrewes det.” / light red: “B.N. BORNEO. Mt. Kinabalu, Lumu Lumu, 5580 ft. April 12 th, 1929 ”; with backside: “ H. M. Pendlebury coll. F.M.S. Museums.” ( BMNH).

Additional material: 1 ♀, Malaysia-Borneo Sabah 21.3.–20 .4.1996 GUNUNG EMAS 1700m lgt. J. Kadleg, ( CBB) .

Remarks: Andrewes (1931) based his description on two specimens from Lumu Lumu. One of them is deposited in the BMNH and is labelled as “Type”. The specimen was pinned in the middle between the two elytra so that the elytra are opened. It has been mounted on a paper card and the original pin has been used for that card. The second specimen is not anymore in the BMNH, could not be located, and is obviously lost. In the interst of the stability the lectotype is designated here.

Redescription ( Figs 1 , 15 , 17 , 19 , 31 ; pp. 8, 15, 19, 23, 35)

Measurements: Lectotype. Body length 10.1 mm, width 2.7 mm; ratio length/ width of pronotum 0.74; ratio length/width of elytra 1.85.

Additional material: Body length 10.3 mm, width 2.8 mm; ratio length/ width of pronotum 0.76; ratio length/width of elytra 1.82.

Colour: Shiny. Black. Wings of clypeus and supraantennal plates transparent red-brown, antennae, mouthparts, tarsalia medium brown, legs dark redbrown.

Head: Relatively wide, 15% narrower than pronotum. Clypeus with four anteriorly projecting teeth, with blunt raised knob between the two clypeal setae, transversally hollowed out between anterior margin and knob, wings of clypeus sharply angled (<90°), projecting, divided from supraantennal plates by distinct obtusely angled incision and from middle part of clypeus by broad rounded incision. Supraantennal plates vaulted, with reflexed rounded margin, hollowed out anteriorly; frons separated from clypeus by band of big punctures, with obtuse nearly parallel running carinae at each side posteriorly, carinae with furrows medially and laterally, carinae enclosing the slightly convex frons, frons with irregular big and small punctures, with longitudinal impression at middle. Supraorbital carina flat, with two setae, posterior one arising from big tubercle. Eyes not big, projecting, tempora carinate, genae distinctly vaulted, nearly as high and as long as eyes, enclosing eyes posteriorly, forming distinct angle at neck. Eyes and tempora separated from middle part of head by distinct supraorbital carinae and deep broad furrows. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction formed by doubled irregular row of big punctures, neck laterally densely covered with medium sized punctures. Labrum formed by two broad lobes, with slight emargination at middle, 7-setose (in the lectotype one seta is missing), ciliate laterally, with isodiametric reticulation. Mandibles robust, nearly as long as head, relatively slender, acute at apex, upper margin of scrobe of both mandibles at base with carina-like tooth, right one with distinct tooth at middle, left one with knob-like tooth at base visible in opened position, both mandibles hollowed out at base; scrobe rounded dorsally and ventrally in apical half, with no carinae, appearing rounded at sides. Maxilla bent sigmoidally, acutely hooked at apex. Apical maxillary palpomere slender, straight. Apical labial palpomere slender, straight, more than twice as long as 2 nd segment, 2 nd segment bisetose. Ligula bilobed at apex, with long seta; paraglossae slender. Ventral surface of neck covered with big punctures, with transverse wrinkles near gula. Gula smooth. Submentum and mentum distinctly separated; submentum with two pairs of setae; mentum keeled at middle, nearly smooth, with fine and complete reflexed margin, lateral lobes projecting, thought obtusely angled at tips, with two setae near base of each lobe, median tooth forming broad obtuse triangle, bisetose, projecting as far as lateral lobes. Paragena sharp, carina-like, without tooth. Antennae short, reaching up to middle of pronotum, scapus carinate, with one seta dorsoapically, scapus and pedicellus longitudinally reticulated, segments 5–10 broader than long (L/ W 0.90), antennomeres flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with some additional longer setae.

Pronotum: Subquadratic, broader than long. Sides parallel at middle but convex slightly before posterior angles. Anterior margin slightly bisinuate. Reflexed lateral margin distinctly crenulated up to posterior angle, smooth up to base, running from posterior angle to base as sigmoid curve. Lateral channel broad, subfoveolate between the two lateral setigerous punctures, paralateral part of disc slightly vaulted, basal channel broad and deep. Anterior setigerous puncture at level of anterior transverse line, the posterior one located before level of posterior angle, removed from lateral channel by diameter of pore. Anterior angles distinctly projecting, rounded, formed by reflexed lateral margin, posterior angles sharp, not projecting laterally due to convexity of lateral margin in basal part. Anterior transverse line sharp, not reaching lateral channel. Median line deep and conspicuously broad, with blunt carinae bilaterally, surpassing level of anterior transverse line but not joining, running nearly up to channel of base, not adjoining base. Surface with scattered big punctures and some irregular big wrinkles, with fine isodiametric reticulation towards base, with longitudinal vault parallel to lateral channel. Basal part of disc elongated posteriorly and bilaterally, hanging over basal channel (lateral view). Flange at base sharply raised (lateral view).

Elytra: Elongate, sides parallel at middle. Marginal channel broad, completely visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles, no fold-like apical carina visible. Reflexed margin with fine crenulation up to apex, distinct at humerus, margin and channel bending over rounded humerus up to 5 th stria, with rounded humeral tooth. Basal tubercle big, with setigerous puncture, situated at declivity of 1 st stria 3. Striae 1–4 free at base, 7 th shortened at humerus and nearly joining with stria 6, all striae deep, with indistinct row of punctures, striae 1 and 2 ending free at apex, 3–4 and 5–6 joined at apex. Striole distinct, deepened at base, with small sharp tubercle at base. Intervals 1–4 moderately convex, others distinctly convex, all flattened towards apex, 3 rd at base broader, raised, and with longitudinal tubercle, 8 th small, carinate at base and apically. Interval 3 with five robust setigerous punctures adjoining stria 3.

Hind wings: Fully developed. Ventral surface: Proepisternum with submarginal ridge. Proepipleuron small, nearly smooth, submarginal furrow distinct. Epipleuron broadened in basal third, with row of partly connected big punctures in basal quarter. Proepisternum, prosternite, episternum, and lateral parts of mesosternum covered densely and irregularly with big punctures, mesosternum smooth at middle. Sternites smooth, 3 rd to 5 th with paralateral ambulatory setae at each side, ventral strigae distinct, sternite six with flat transverse depression apically, with two widely separated apical setae at each side.

Legs: Protibia with slight dorsal furrow at base, digitation distinct, with two big and one small lateral preapical denticles, all furnished with seta close to apex, apical spine curved hook-like ventrally and laterally, movable spur moderately curved, inner denticle with corkscrew-like seta; basal tarsomere elongated, longer than the following three together. Mesotibia with big long apical spine, furnished with one seta apically and another one at middle. All tarsomeres relatively broad.

Male genitalia ( Fig. 19 ): Median lobe sclerotized, regularly and moderately arcuate in middle part, distinctly bent at apex, apex formed as slightly asymmetric rounded big spatula. Oroficium large, closing lips less sclerotized. No spines or teeth visible on endophallus (at 500x by optimized condenser and different filters). Parameres asymmetrical, slender, somewhat twisted, ventral one with four medium sized setae at apex, dorsal one with three medium sized and one very fine seta at apex.

Female genitalia ( Fig. 31 ): Coxostylus sigmoid, dorso-ventrally moderately depressed, gently curved to apex, with one seta at apex, with one longer ensiform and four nematiform setae in middle part.

Variation: The labrum is 6-setose in the lectotype and 7-setose in the additional specimen. In the additional specimen one of the ambulatory setae of the sternite is missing. The anterior transverse line of the pronotum is sharper in the lectotype.

Diagnosis: A big black species with distinct knob on the clypeus and flat broad transverse impression with band of big punctures behind knob, with rounded humeral tooth, tubercle at the base of interval 3 of the elytra, and five setigerous punctures on interval 3. Moreover, the pronotum is distinctly broader than long, and the margin of pronotum and elytra is crenulated. Distinguished from all other species by the long and slender mandibles with a scrobe that is not carinate in the apical half, and they each exhibit an elevated tooth near the base. In addition, the clypeus possesses four projecting teeth anteriorly.

Distribution: Known from the North of Borneo only. The type was collected at 5580 ft (1700 m) and the additional specimen at the same altitude.

Figures

Figs 1–4: Orictites spp., habitus. 1: O. minotaur ANDREWES, 1931, lectotype, male; 2: O. costulipennis BATES, 1892, holotype, male; 3: O. mjoebergi LouWERENS, 1964, lectotype, male; 4: O. desuntsetosus sp. nov., holotype, male.

Figs 15–16: Orictites ssp., head and pronotum, lateral view; 15: Orictites (subgenus Orictites) minotaur ANDREWES, 1931,; 16: Orictites (subgenus Semictites) costulipennis BATES, 1892.

Figs 17–18: Orictites spp., Maxilla, maxillary and labial palpomere. 17: O. minotaur ANDREWES, 1931 (subgenus Orictites); 18: O. costulipennis BATES, 1892 (subgenus Semictites subg. nov.).

Figs 19–20: Orictites spp., aedeagus, ventral view. 19: O. minotaur ANDREWES, 1931, lectotype, scale bar 1.5 mm; 20: O. costulipennis BATES, 1892, holotype, scale bar 0.2 mm.

Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm.