Semictites, Balkenoh, 2017

Balkenohl, Michael, 2017, Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini), Contributions to Natural History 35, pp. 1-68 : 7-13

Subgenus Semictites subgen. nov. ( Figs 2 , 16 , 18 ; pp. 8, 15, 19)

Type species Clivina costulipennis BATES, 1892: 281 , by present designation.

Diagnosis: Black or brown. General appearance like a Clivina . Anterior margin of clypeus straight or excised, without teeth, clypeal wings small, obtuse angled or rounded. Mandibles distinctly shorter than head; dorsal margin of scrobe of both mandibles at base without elevated tooth, right mandible with indistinct tooth at base, left mandible with small but distinct tooth slightly anterior to base. Apical maxillary palpomere securiform, apical labial palpomere nearly straight, fusiform, slightly bottle-like. Pronotum about as long as wide, almost flattened on disc, anterior angles angulate, more or less projecting. Aedeagus with spatula or stick-like. Parameres asetose or with up to two apical setae. Coxostylus with one or two subapical setae. Body length smaller than 7.4 mm.

Etymology: The name refers to the numerous similarities shared with the subgenus Orictites .

Key to the species

1 Frontal margin of clypeus with four distinctly projecting teeth; clypeal wings tooth-like projecting anteriorly, sharply angled (distinctly less than 90°); mandibles as long as head; dorsal margin of scrobe of both mandibles at base with carina-like tooth; pronotum distinctly broader than long; length 10–10.3 mm ............................ Subgenus Orictites ANDREWES, 1931 One species ....................................................... minotaur ANDREWES, 1931

– Frontal margin of clypeus without teeth; clypeal wings small, obtusely angled or rounded; mandibles distinctly shorter than head; dorsal margin of scrobe of both mandibles at base without elevated tooth; pronotum about as long as wide (ratio length/width 0.9–1.1); length shorter than 7.4 mm ............................................... 2: Subgenus Semictites subgen. nov.

2 Intervals of elytra without any setigerous punctures; length 4.5 mm. ............................................................................. desuntsetosus sp. nov.

– Interval 3, or 3, 5 and 7 of elytra with setigerous punctures ..................... 3

3 Intervals 3, 5, and 7 of elytra with series of setigerous punctures; clypeus deeply excised anteriorly; length 3.8 mm ................... plurisetosus sp. nov.

– Setigerous punctures on interval 3 of elytra, only; clypeus straight or slightly excised anteriorly .......................................................................... 4

4 With up to seven setigerous punctures on interval 3 of elytra .................. 5

– With four setigerous punctures on interval 3 of elytra .............................. 9

5 Black; with five setigerous punctures on interval 3 of elytra; knob on clypeus indistinct, developed as elevation; interval 3 and 4 of elytra at base with a tubercle each; length 4.8 mm ........................ anteriortenuis sp. nov.

– With more than five setigerous punctures on interval 3 of elytra; brown or black species ............................................................................................. 6

6 Brown; with up to seven setigerous punctures on interval 3 of elytra (four to seven punctures, variable, see tab. under description); interval 3 and 4 at base with tubercle; knob on clypeus indistinct, developed as elevation; punctures of neck constriction of head visible laterally, no constriction at middle; lateral margin of pronotum straight, anterior angles not projecting; length 3.4–4.0 mm ................................................. omnipunctatus sp. nov.

– Black; with six or seven setigerous punctures on interval 3 of elytra; only interval 3 at base with tubercle; knob on clypeus distinct, neck constriction of head with big punctures; length 4.8–6.6 mm ........................................ 7

7 Lateral margin of pronotum straight, anterior angles distinctly projecting; median line of pronotum conspicuously broad, interval 3 of elytra with six to seven setigerous punctures; length 6.6 mm ....... ampliosetosus sp. nov.

– Lateral margin of pronotum convex, anterior angles indistinctly projecting; median line of pronotum moderately narrow ............................................ 8

8 Median line of pronotum finely adjoining basal constriction; without central puncture on frons; interval 3 of elytra with seven setigerous punctures; length 4.8 mm ......................................................... barclayi sp. nov.

– Median line of pronotum not joining basal constriction; with central puncture on frons; interval 3 of elytra with six setigerous punctures; length 4.25 mm ........................................................ barclayi minorfrater ssp. nov.

9 Elytra without tooth at humerus .............................................................. 12

– Elytra with distinct tooth at humerus ...................................................... 10

10 Bigger and more massive species; pronotum distinctly transverse, disc distinctly convex, surface sparsely covered with small punctures; anterior transverse line of pronotum indistinct; preapical spine of mesotibia nearly at apex of tibia, length 7.4 mm ....................................... brancuccii sp. nov.

– Smaller and slender species; pronotum as long as wide or slightly broader than long, disc somewhat flattened, surface densely covered with big punctures; anterior transverse line of pronotum distinct; preapical spine of mesotibia located distinctly preapically, length up to 5.9 mm ................. 11

11 Elytra with tooth at humerus of moderate size, tooth rounded; reflexed margin of pronotum at base tooth-like raised bilaterally; lateral margin of pronotum with few fine scars anteriorly; length 5.8 mm .............................. ............................................................................... tubercucollis sp. nov.

– Elytra with tooth at humerus big, sharp; reflexed margin of pronotum at base in total of about same appearance; lateral margin of pronotum crenulated; length 5.8 mm ................................................ charleshuberi sp. nov.

12 Disc of pronotum with basal part bilaterally elongated posteriorly as toothlike vault slightly hanging over basal channel; lateral margin of pronotum with traces of scars .................................................................................. 13

– Disc of pronotum in basal part regularly developed; lateral margin of pronotum smooth; pronotum as long as wide; length 5.6 mm .......................... ...................................................................... costulipennis BATES, 1892

13 Clypeus without distinct knob, developed as broad elevation without punctures; intervals 3 and 4 at base of elytra with tubercles; interval 3 of elytra with four setigerous punctures; length 5.8 mm ................................. .............................................................................. anteriorlatus sp. nov.

– Clypeus with distinct knob, with punctures posterior to knob in form of a V; only interval 3 at base of elytra with tubercle; interval 3 of elytra with four/ five setigerous punctures; length 5.4 mm ..... mjoebergi LouWERENS, 1964

Figures

Figs 1–4: Orictites spp., habitus. 1: O. minotaur ANDREWES, 1931, lectotype, male; 2: O. costulipennis BATES, 1892, holotype, male; 3: O. mjoebergi LouWERENS, 1964, lectotype, male; 4: O. desuntsetosus sp. nov., holotype, male.

Figs 15–16: Orictites ssp., head and pronotum, lateral view; 15: Orictites (subgenus Orictites) minotaur ANDREWES, 1931,; 16: Orictites (subgenus Semictites) costulipennis BATES, 1892.

Figs 17–18: Orictites spp., Maxilla, maxillary and labial palpomere. 17: O. minotaur ANDREWES, 1931 (subgenus Orictites); 18: O. costulipennis BATES, 1892 (subgenus Semictites subg. nov.).