Orictites mjoebergi LouWERENS, 1964

Balkenohl, Michael, 2017, Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini), Contributions to Natural History 35, pp. 1-68 : 25-29

Orictites mjoebergi LouWERENS, 1964

Catalogue

Orictites mjoebergi ; Louwerens, 1964: 172;

Orictites mjoebergi LouWERENS ; Lorenz 1998: 135;

Orictites mjoebergi LouWERENS ; Balkenohl 2001: 18.

Material examined: Type material: Lectotype (by present designation). ♂, with labels and data: white, handwritten: “Type Orictites mjÖbergi sp.n.” and printed “det. C.J.Louwerens ” / blue: “3491 E91 +” / 2 nd pin with original small paper card: small, white, “ Pajau River ” / white, “O. Borneo MjÖberg” / small blue: “3492 E91 +” / big, white, “ ORICTITES Andr. ” / big, white: “MjÖbergi Louw.” ( NHRS).

Remarks: C.J. Louwerens (1964) based the description on two specimens from Pajan River. One of them is located in the NHRS (see material examined), the other one could not be located and is obviously lost. The available specimen isdesignated here as lectotype in the interst of the stability. In the specimen the right wing of the clypeus is slightly more angled (dorsal view) and more fused with the median part. I recognise this character as intraindividual variation. Exactly this asymmetric character is described by Louwerens (1964). However, careful investigation revealed that this specimen is a male and not a female as indicated by Louwerens. The inner structures were slightly damaged, obviously by a former mounting attempt. The left antennae misses segments 8–11. It should be mentioned that the sketch provided by Louwerens does not display a realistic picture of the species. In addition, Louwerens (1964: 173) describes the presence of five setigerous punctures on interval 3 of the elytra. However, the type specimen possesses four, respectively five setigerous punctures (right/left side). The additional specimens all exhibit four setigerous punctures on either side.

Additional material: 1 ♂, NORD-SUMATRA: Umg. Siulakosa 5.8.1991, ERBER / Samosir, ob. Tomok 1500 m, Kiefernwald , unter Rinde ( SMNS) ; 1 ♂, Filippine Mindanao III.1995 M. Malindang North Sambuanga ( CBP) ; 1 ♀, PHILIPPINES, N Luzon , Ifugao , Banaue , XI 2014. local collector leg. ( CBB) ; 1 specimen: head and pronotum lost, F. C. DRESCHER G. Tangkoeban Prahoe 4000 – 5000 Voet Praeanger, Java VI. 1933 / Ex coll. Bänninger / H.E. Andrewes Coll. B.M.1945–97. ( BMNH) .

Redescription ( Figs 3 , 21 , 33 ; pp. 8, 24, 35)

Measurements: Lectotype: Body length 5.4 mm, width 1.52 mm; ratio length/ width of pronotum 1.07; ratio length/width of elytra 1.95.

Additional material (n = 3): Body length 4.93–5.9 mm ( = 5.4 mm), width 1.37–1.65 mm ( = 1.53 mm); ratio length/width of pronotum 0.96–1.06 ( = 1.0); ratio length/width of elytra 1.78–1.97 ( = 1.89).

Colour: Shiny. Black. Elytra red-brown, pronotum darker. Wings of clypeus and supraantennal plates transparent red-brown. Antennae, mouthparts, middle and hind legs yellowish brown, front legs dark red-brown.

Head: A quarter narrower than pronotum. Clypeus limited laterally by small obtuse angled tooth, anterior margin straight, with blunt raised knob between the two clypeal setae, posterior part of knob covered with medium sized punctures, anterior part smooth; wings obtuse-angular, rounded at tip, hollowed out, divided from middle part of clypeus by slight incisions and carinae, divided from supraantennal plates by distinct obtusely angled incisions and indistinct carina, clypeus and wings reflexed margined, supraantennal plates vaulted, elongated as supraorbital carina, smooth, with reflexed rounded margin, frons separated from clypeus by flat transverse depression and a group of punctures formed like a letter V, with nearly parallel running costate frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina. Frons moderately convex, nearly smooth, with small but distinct longitudinal impression at middle. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from a tubercle. Eyes of moderate size, moderately convex, genae enlarged, regularly rounded, not as high as and a third shorter than eyes, enclosing eyes posteriorly. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked as distinct step, with band of big punctures, neck laterally densely covered with medium sized punctures. Labrum indistinctly trilobed, appearing straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles half as long as head, flattened, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled and somewhat hollowed out dorsally. Maxilla nearly straight, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, longer than 2 nd segment, 2 nd segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck covered with medium sized punctures and irregular reticulation. Gula smooth. Submentum and mentum distinctly separated; submentum with two pairs of setae; mentum keeled at middle, nearly smooth, with fine and complete reflexed margin, lateral lobes projecting, thought obtusely angled at tips, with two setae near base of each lobe, median tooth forming broad obtuse triangle, bisetose, less projecting than lateral lobes. Paragenae sharp, carina-like, without tooth. Antennae of medium length, reaching over middle of pronotum, scapus carinate, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 transverse (L/ W 0.80), antennomeres subglobose, nearly dull, segments 4–11 densely pubescent, with a few additional long setae.

Pronotum: Square (L/ W 1.07), sides straight but slightly diverging to base, gently rounded before posterior angle. Anterior margin slightly bisinuate. Reflexed lateral margin nearly smooth, with indistinct reticulation (100x), from posterior angle to base running as smooth, slightly convex line. Lateral channel distinctly broadened between the two lateral setigerous punctures, with fine irregular reticulation, with a few medium-sized punctures situated in broadened part of channel, basal channel broad, deep, foveolate. Anterior setigerous puncture located at end of anterior quarter, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angle slightly but distinctly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as large rounded tooth, projecting laterally. Anterior transverse line sharp, punctured, not adjoining lateral channel. Median line deep, broad, thought with indistinct blunt carina-like vault bilaterally, surpassing level of anterior transverse line but not joining, smaller at base, joining base. Surface with two longitudinal groups of paramedian and paralateral big punctures and with a few fine, irregularly situated punctures, with irregular reticulation laterally, shiny on disc; basal impression developed as small flat furrow; base of disc with indistinct reticulation. Basal part of disc elongated posteriorly and bilaterally at declivity, as tooth-like vault slightly hanging over basal channel (lateral view). Flange raised keel-like (lateral view).

Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, not completely visible from above due to convexity of intervals 7 and 8, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of punctures laterally. Reflexed margin with fine crenulation in anterior quarter, channel broadened at humerus, bending with margin over rounded toothless humerus up to 5 th stria. Basal tubercle big, with setigerous puncture, situated at declivity of first stria. Striae 1–4 free at base, all striae deep, broad, distinctly punctate-striate, striae 1–2, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura costiform, raised, basal area around striole moderately impressed. Intervals shiny, distinctly convex, 3 rd broader, strikingly raised and with large tubercle at base, 4 th broader and raised at base but not as strongly as 3 rd, 7 th running to apex as obtuse carina. 8 th small, completely carinate. Interval 3 with four/five setigerous punctures adjoining stria 3 (see variation).

Hind wings: Fully developed.

Ventral surface: Proepisternum with complete submarginal ridge, with some punctures, with irregular reticulation laterally. Proepipleuron small, with punctures and indistinct reticulation. Epipleuron broadened and with big punctures in basal quarter. Proepisternum somewhat swollen at level of posterior setigerous puncture, prosternite smooth, double keeled at middle, mesosternum smooth. Sternites smooth at middle, each with big punctures at base laterally, 3 rd to 5 th with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.

Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral and lateral, no dorsal furrow, basal tarsomere elongated, longer than the following four together. Mesotibia with apical spine, furnished with strong seta. All tarsomeres relatively broad.

Male genitalia ( Fig. 21 ): Relatively short, median lobe broad, regularly arcuate at middle, apex twisted apically. Endophallus with bristles laterally. Both parameres slender, slightly twisted, asetose.

Female genitalia ( Fig. 33 ): Coxostylus of moderate width, distinctly curved to apex, with two preapical setae close together, dorsally with some minute scars in apical half, with obtuse knob-like tooth dorsally at base, with two short ensiform setae at middle, with four longer nematiform setae.

Variation: Bilateral variation is observed in the right clypeal wing, mentioned under remarks. In addition, in the lectotype the left elytra bears five setigerous punctures and the right one four. The other specimens all have four setigerous punctures on each side.

Diagnosis: A medium-sized almost black species with red-brownish elytra, with distinct knob on the clypeus, humerus without tooth, and four setigerous punctures on interval 3 (but compare remarks). Only in this species and in O. anteriorlatus is the basal part of the disc of the pronotum posteriorly elongated bilaterally at the declivity as a tooth-like vault slightly hanging over the basal channel. By contrast to O. anteriorlatus , O. mjoebergi exhibits a tubercle at the base of interval 3 of the elytra, only, and the furrow between clypeus and frons consists of punctures in form of the letter V.

Figures

Figs 1–4: Orictites spp., habitus. 1: O. minotaur ANDREWES, 1931, lectotype, male; 2: O. costulipennis BATES, 1892, holotype, male; 3: O. mjoebergi LouWERENS, 1964, lectotype, male; 4: O. desuntsetosus sp. nov., holotype, male.

Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm.

Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm.