Riculorampha, Rota & Rota & Brown, 2009

Rota, Jadranka, Rota, Jadranka & Brown, John, 2009, A new genus and species of Grapholitini (Lepidoptera, Tortricidae) from Florida, U. S. A., ZooKeys 23 (23), pp. 39-46 : 41-42

Riculorampha , gen. n.

urn:lsid:zoobank.org:act:F399FA93-767D-4CA3-BC2D-2B75CCF168E3

Type species: Riculorampha ancyloides , sp. n.; designated by monotypy

Figs 1–6

Riculorampha is provisionally assigned to the Dichrorampha -group of genera on the basis of the following putative synapomorphies: forewing termen with a row of black dots, forewing termen with a shallow subapical notch, hindwing veins R s and M 1 separate and parallel or subparallel, female frenulum with two bristles, and corpus bursae with a single signum. The significance of these characters is discussed below.

Diagnosis. Superficially, Riculorampha is similar to nearly all other genera in the Dichrorampha -group, with a subapically “notched” termen of the forewing, a row of black dots along the forewing termen, and a two-bristled frenulum in the female. It shares with Dichrorampha , Goditha Heinrich , Ricula Heinrich , and Riculoides the loss of one of the two signa in the corpus bursae in the female genitalia. The male genitalia of Riculorampha can be separated from those of Ricula and Riculoides by the reduction of socii (long and extremely slender in Ricula and Riculoides ), the broad, rounded dorsum of the tegumen (narrow and attenuate dorsally in Ricula and Riculoides ), and the large, triangular sacculus (smooth and rounded in Ricula and Riculoides ). The last character also serves to distinguish Riculorampha from Goditha and Dichrorampha and all other members of the Dichrorampha -group.

Description. Head: Vertex with large scale patches overlapping mediodorsally (Fig. 2); upper frons with overhanging tuft of relatively smooth scales; lower frons sparsely covered with appressed scales; ocellus relatively large, ca. 0.2× diameter of compound eye. Antenna slightly shorter than 0.5 length of forewing costa, somewhat thickened, with fine, short, dense sensory setae in both sexes. Length of labial palpus subequal to diameter of compound eye, upcurved, smooth scaled. Maxillary palpus rudimentary. Thorax: Dorsum smooth scaled, without posterior tuft. Legs unmodified in male. Forewing length (Fig. 1) ca. 2.0× width, forewing termen broadly bilobed, notched at vein M 1; row of black dots along termen; discal cell length ca. 0.6× wing length; all veins present and separate (Fig. 3); chorda present, weakly defined basally. M vein well developed throughout discal cell. M 2, M 3, and CuA 1 parallel beyond discal cell and remote at termen. CuA 2 originating from ca. 0.66× length of discal cell. Hindwing length ca. 1.75× width; cubital pecten absent in both sexes; female frenulum with two bristles; R s and M 1 separate, but relatively close together and parallel in basal 0.25; M 3 and CuA 1 connate. Abdomen: Male with internal glands on segment three; sternite of segment eight with Y-shaped sclerotization medially. Male genitalia (Fig. 4) with tegumen nearly parallel-sided, ca. 0.6× length of valve, rounded dorsally, expanded medially into triangular flap; no trace of uncus; socius present as short, subtriangular, hairless pad (in slide-mounted preparation of holotype, one socius hidden beneath triangular flap of tegumen); valva elongate, upcurved, constricted at neck, narrowed apically, rounded at apex, with dense setae and spines on cucullus; basal cavity large, extending distally to neck; sacculus strongly produced apically, triangular, smooth with exception of scattered setae; microtrichiae present on outer surface of valva. Phallus curved at 0.2 from base, then straight and weakly attenuate in distal 0.8; vesica with ca. 5 slender, elongate spindle-shaped cornuti. Female genitalia (Fig. 6) with papilla analis unmodified; apophysis anterioris and posterioris subequal in length and slender, anterioris extending slightly anterad of ostium; sternite of segment eight wrinkled anterad of ostium; ductus bursae ca. 2.0× as long as abdominal segment eight, with posterior 0.5 strongly sclerotized, slightly narrowed from ostium to end of sclerotization, with rough surface texture; ductus seminalis originating at about 0.5 length of ductus bursae; ductus bursase about 1.5× wider at ostium than at junction with corpus, at junction with corpus ca. 0.25× width of corpus. Corpus bursae almost round, with rough surface texture and a single spine-shaped signum [assumed to be unmated].

Pupa (Based on 3 exuvia). Typically olethreutine (Fig. 5); head without apical projection; thorax and abdomen without conspicuous sculpturing; abdomen with row of small spines dorsally near middle of segments A2–A6, extending over nearly entire dorsum; slightly shorter row of larger spines near anterior edge of segments A3–A8; cremaster absent; 4 pairs of long hook-tipped setae at posterior end; pair of spines absent from anal rise.

Etymology. The name is a combination of Ricula and Dichrorampha , to which the genus appears to be related; it is interpreted as feminine in gender.