SYMPYCNINAE ALDRICH, 1905: 292

Pollet, M. A. A., Brooks, S. E. & Cumming, J. M., 2004, Catalog Of The Dolichopodidae (Diptera) Of America North Of Mexico, Bulletin of the American Museum of Natural History 2004 (283), pp. 1-114 : 73-76

SUBFAMILY SYMPYCNINAE ALDRICH, 1905: 292

Genus CALYXOCHAETUS Bigot, 1888a

CALYXOCHAETUS Bigot, 1888a : xxiv. Type species: Sympycnus nodatus Loew [Nearctic], by original designation. [As ‘‘ notatus ’’, error].

Treated as subgenus of Sympycnus by Van Duzee (1930e). Listed as synonym of Sympycnus by Dyte & Smith (1980), but treated as valid genus by Robinson & Vockeroth (1981).

NOTHOSYMPYCNUS Wheeler, 1899: 51 . Type species: Nothosympycnus vegetus Wheeler , designation by Coquillett, 1910c: 576.

NOTOSYMPYCNUS, error or unjustified emendation listed in Foote et al. (1965).

NOTES: Calyxochaetus is a New World genus and is most diverse in the Nearctic. Adults are common in shady areas and frequently hover over moist rocks and falling water.

REFERENCES: Van Duzee (1930e; rev.); Robinson (1964; key, eastern U.S.).

arizonicus (Harmston), 1968: 24 ( Sympycnus ). HT in CAS. Type locality: USA: AZ, Santa Catalina Mts. NEW COMBINATION.

binodatus (Harmston & Knowlton), 1940c: 397 ( Sympycnus ). HT in USNM. Type locality: USA: UT, Leeds. NEW COMBINATION.

cilifemoratus (Van Duzee), 1924f: 12 ( Nothosympycnus ). HT in USNM. Type locality: USA: AK, Anchorage. DISTRIBUTION: AK, UT, CO.

fuscitibialis (Harmston & Knowlton), 1940c: 400 ( Sympycnus ); synonymized by Foote et al. (1965) after Harmston (in litt.). HT in USNM. Type locality: USA: UT, Duchesne Co., Currant Creek .

clavicornis (Van Duzee), 1930e: 55 ( Sympycnus ). HT in USNM. Type locality: USA: CA, Mono Lake.

distortus (Van Duzee), 1930e: 55 ( Sympycnus ). HT in USNM. Type locality: USA: UT, Soldier Summit, 7454 ft.

fortunatus (Wheeler), 1899: 52 ( Nothosympycnus ). HT in AMNH. Type locality: USA: PA, Natrona. DISTRIBUTION: ON to NS and ME, s. to NC (NY, NJ, PA, New England) [ Robinson (1964); Bissett (1987)].

frontalis (Loew), 1861a: 67 ( Sympycnus ). ST in MCZ. Type locality: USA: PA. DISTRIBUTION: IA to NS and ME, s. to TN and SC (MO, IN, NY, PA, MA, DC, VA, NC) [ Robinson (1964); Bissett (1987); Vockeroth (personal commun.)].

hardyi (Harmston & Knowlton), 1940c: 397 ( Sympycnus ). HT in USNM. Type locality: USA: UT, Hooper. DISTRIBUTION: UT, NE [ Harmston & Rapp (1983)]. NEW COMBINATION.

hastatus (Van Duzee), 1930e: 57 ( Sympycnus ). HT in CAS. Type locality: USA: CA, San Diego Co., on edge of Colorado Desert. DISTRIBUTION: CA; MEX.

inornatus (Van Duzee), 1917b: 340 ( Nothosympycnus ). ST in CUIC, CAS (1). Type locality: USA: CA, Contra Costa Co., San Pablo, Wild Cat Canyon.

insolitus (Van Duzee), 1932b: 20 ( Sympycnus ). HT in AMNH. Type locality: USA: CO, Aspen, at about 8000 ft. DISTRIBUTION: ID, UT, CO [ Turner (1987)].

isoaristus (Harmston & Knowlton), 1940c: 402 ( Sympycnus ). HT in USNM. Type locality: USA: UT, Cedar Breaks. DISTRIBUTION: UT, NE [ Harmston & Rapp (1983)]. NEW COMBINATION.

luteipes (Van Duzee), 1923d: 63 ( Nothosympycnus ). ST in MCZ. Type locality: USA: ME, Bar Harbor. DISTRIBUTION: ON, QC, NY, ME [Vockeroth (personal commun.)].

tarsalis (Curran), 1924a: 108 ( Nothosympycnus ). HT in CNC. Type locality: CAN: ON, Orillia.

millardi Meuffels & Grootaert, 1999: 290 ; n. name for Calyxochaetus monticola (Van Duzee) . Type locality: USA: CO, La Plata Co., Electric Lake , at about 8400 ft.

monticola (Van Duzee), 1932b: 20 ( Sympycnus ); preocc. by Sympycnus monticolus Becker, 1919 . HT in AMNH. Type locality: USA: CO, La Plata Co., Electric Lake , at about 8400 ft.

nodatus (Loew), 1862: 215 (Cent. 2, no. 68) ( Sympycnus ). ST in MCZ. Type locality: USA: IL. DISTRIBUTION: MB to NY, s. to TX (NE, KS, MN, IA, MO, AR, MS, ON, WI, IL, IN, OH) [ Robinson (1964); Harmston & Rapp (1983)].

notatus ; error by Bigot (1888a: xxiv).

abbreviatus (Van Duzee), 1917b: 341 ( Nothosympycnus ); synonymized by Harmston & Miller (1966). HT in SEMC. Type locality: USA: KS, Douglas Co .

oreas (Wheeler), 1899: 55 ( Nothosympycnus ). HT in AMNH. Type locality: USA: WY, Buck Creek. DISTRIBUTION: CA, UT, WY.

spatulatus (Harmston & Knowlton), 1940c: 398 ( Sympycnus ). HT in USNM. Type locality: USA: UT, Hanksville.

pennarista (Harmston & Knowlton), 1940c: 399 ( Sympycnus ). HT in USNM. Type locality: USA: UT, Manila. NEW COMBINATION.

pictipes (Harmston & Knowlton), 1940c: 401 ( Sympycnus ). HT in USNM. Type locality: USA: UT, Kanosh Canyon. NEW COMBINATION.

sobrinus (Wheeler), 1899: 54 ( Nothosympycnus ). HT in AMNH. Type locality: USA: ID, Lewiston. DISTRIBUTION: CA, ID.

tripilus (Van Duzee), 1930e: 52 ( Sympycnus ). HT in CAS. Type locality: USA: CA, San Diego Co., on edge of Colorado Desert. DISTRIBUTION: CA, UT. NEW COMBINATION.

vegetus (Wheeler), 1899: 53 ( Nothosympycnus ). ST in AMNH, CNC (1). Type locality: USA:

CA, Monterey Co. DISTRIBUTION: OR, CA, ID, UT, NE,?NS [ Harmston & Rapp (1983)].

Genus CAMPSICNEMUS Haliday, 1851

CAMPTOSCELES Haliday, 1832: 357 (subgenus of Medeterus ; suppressed by I.C.Z.N. 1958: 349).

LEPTOPEZINA Macquart, 1835: 554 . Type species: Diastata gracilis Meigen, 1820 , by monotypy. Nomen oblitum (see Evenhuis, 2003).

CAMPSICNEMUS Haliday, 1851: 187 . Type species: Dolichopus scambus Fallén [Palearctic]. By validation of I.C.Z.N. (1958: 351) (Opinion 531) [Name No. 1316 on Official list of Generic Names in Zoology]. Nomen protectum (see Evenhuis, 2003).

ECTOMUS Mik, 1878b: 8 . Type species: Medeterus alpinus Haliday , by original designation.

EMPEROPTERA Grimshaw in Grimshaw & Speiser, 1902: 81. Type species: Emperoptera mirabilis Grimshaw , by monotypy.

CAMPTOSCELUS Kertész, 1909: 306 . Type species: Dolichopus scambus Fallén , automatic. Unjustified emendation of Camptosceles Haliday, 1832 .

EMPEROTERA , error by Hardy & Kohn (1964: 98).

NOTES: Campsicnemus has been recorded from all realms except the Neotropical and has undergone a huge species radiation on the Hawaiian Islands with more than 130 described species. Elsewhere it is relatively poor in species. Species of Campsicnemus are generally hygrophilous and soil­dwelling.

They mainly occur near open water in forests, marshlands, and on banks of all kinds of water bodies while a number of species are restricted to humid heathland. Like Hydrophorus , they have been observed skating on the water surface and some species are active during winter.

REFERENCES: Curran (1933; key); Harmston & Knowlton (1942a; rev.); Robinson

(1964; key, southeastern U.S.).

alaskensis Harmston & Miller, 1966: 88 . HT in CAS. Type locality: USA: AK, Sterling Highway, Clam Beach.

alexanderi Harmston & Miller, 1966: 89 . HT in CAS. Type locality: USA: AK, Taylor Highway, West Fork Dennisan River, Mile Post 49.

americanus Van Duzee, 1924f: 3 . HT in USNM. Type locality: USA: AK, Healy. DISTRIBUTION: AK, CO, ON, QC, MI, NY, ME, MA [ Beaulieu & Wheeler (2001)].

arcuatus Van Duzee, 1917a: 125 . HT in CAS.

Type locality: USA: CO, Victor, 9800 ft. DISTRIBUTION: CO, WY.

bryanti Malloch, 1932: 121 . HT in CAS. Type locality: CAN: MB, Fort Churchill. DISTRIBUTION: MT, SK, WY, MB.

cinctipus Harmston, 1968: 17 . HT in CAS. Type locality: CAN: NT, Aklavik.

claudicans Loew, 1864b: 194 . ST in MCZ. Type locality: USA: AK, Sitka. DISTRIBUTION: AK to CA, ID, MT, and CO.

clandicans; error by Van Duzee (1923c: 241).

coloradensis Harmston & Miller, 1966: 90 . HT in CAS. Type locality: USA: CO, Ward.

curvispina Van Duzee, 1930a: 2 ; n. name for Campsicnemus calcaratus Van Duzee. Type locality: USA: AK, Fairbanks .

calcaratus Van Duzee, 1924f: 3 ; preocc. by Campsicnemus calcaratus Grimshaw, 1901 . HT in USNM. Type locality: USA: AK, Fairbanks .

degener Wheeler, 1899: 58 . ST in AMNH. Type localities: USA: CA, Pacific Grove; ID, Vollmer. DISTRIBUTION: AK and WA to CA, CO, NE, ON, QC, NS, ne. U.S. [ Harmston & Rapp (1983); Bissett (1987)].

hirtipes Loew, 1861a: 68 . ST in MCZ. Type locality: USA: PA. DISTRIBUTION: NE and KS to MI, QC, NY, and NS, s. to VA, NC, GA, and FL [ Harmston & Rapp (1983); Bissett (1987); Vockeroth (personal commun.)].

latipes (lapsus) Curran, 1934: 218, fig. 2 [ Sabrosky (1971)].

melanus Harmston & Knowlton, 1942a: 11 . HT in USNM. Type locality: USA: UT, Torrey.

milleri Harmston, 1966b: 226 . HT in CAS. Type locality: USA: OR, Boi­Cope State Park.

montanus Harmston & Knowlton, 1942a: 14 . HT in USNM. Type locality: USA: MT, Gardiner. DISTRIBUTION: MT, WY.

nigripes Van Duzee, 1917a: 126 . HT in CAS. Type locality: USA: CA, Sacramento. DISTRIBUTION: MT to CA and CO.

oedipus Wheeler, 1899: 60 . ST in AMNH, USNM (1). Type locality: USA: WY, Two­gwo­te­ee Pass. DISTRIBUTION: OR, UT, WY, CO.

oregonensis Harmston & Miller, 1966: 88 . HT in CAS. Type locality: USA: OR, Barview State Park.

philoctetes Wheeler, 1899: 59 . ST in AMNH. Type localities: USA: WY, Lusk; Little Wind River canyons ; Hunter’s Creek ; Jackson Lake ; SD. DISTRIBUTION: WA, OR, ID, UT, WY, SD, MN.

thersites Wheeler, 1899: 61 . ST in AMNH. Type localities: USA: WY, Natrona Co .; Wind River Mts .; Two­gwo­te­ee Pass. DISTRIBUTION: MT, ID, UT, WY.

utahensis Harmston & Knowlton, 1942a: 13 . HT

in USNM. Type locality: USA: UT, Heber. DIS­

TRIBUTION: UT, NE [ Harmston & Rapp (1983)]. vanduzeei Curran, 1933: 6 . HT in AMNH. Type locality: CAN: MB, Churchill. wheeleri Van Duzee, 1923d: 64 . HT in MCZ.

Type locality: USA: ME, Machias. DISTRIBU­

TION: QC, ME, NS [ Bissett (1987)]. variabilis (Van Duzee), 1932a: 9 ( Thinophilus ).

HT in CNC. Type locality: CAN: QC, Natashquan.

GENUS EREBOMYIA Runyon &