Paraclius, Loew, 1864 b

Pollet, M. A. A., Brooks, S. E. & Cumming, J. M., 2004, Catalog Of The Dolichopodidae (Diptera) Of America North Of Mexico, Bulletin of the American Museum of Natural History 2004 (283), pp. 1-114 : 43-47

Genus PARACLIUS Loew, 1864b

PARACLIUS Loew, 1864b: 97 . Type species: Pelastoneurus arcuatus Loew [Neotropical], designation by Coquillett, 1910c: 583. Erroneously treated as an emendation of Paracleius Bigot in Foote et al. (1965), see comments un­ der Pelastoneurus Loew.

LEPTOCORYPHA Aldrich, 1896b: 315 . Type species: Leptocorypha pavo Aldrich , by monotypy. Synonymized by Robinson (1970b).

LEPTORHETHUM, Parent, 1934a , not Aldrich, 1893b, misid. listed by Bickel & Dyte (1989).

NOTES: Species of Paraclius are known from every zoogeographical realm, but the genus is most diverse in the Neotropics. Nearctic species are found on moist soil or sand or on low herbaceous foliage near water. Most Nearctic species are coastal ( Robinson, 1964; Foote et al., 1965).

REFERENCES: Aldrich (1904; key); Robinson (1964; key, eastern North America).

aberrans Robinson, 1964: 180 . HT in USNM.

Type locality: USA: TX, Collin Co., Plano.

DISTRIBUTION: TX; MEX. alternans (Loew), 1864a: 95 (Cent. 5, no. 91)

( Pelastoneurus ). HT in MCZ. Type locality:

USA: NY. DISTRIBUTION: ME to FL (MA, CT,

NY, MD, NC), Bermuda [ Robinson (1964);

Woodley & Hilburn (1994)]. vicinus Aldrich, 1904: 277 . ST in USNM (8).

Type locality: USA: MA, New Bedford. claviculatus Loew, 1866b: 45 (Cent. 7, no. 83).

HT in MCZ. Type locality: USA: NY, New

Rochelle. DISTRIBUTION: ON to NS, s. to NC

(NY, MA);?Neotropics [ Robinson (1964,

1970b)]. fraternus Van Duzee, 1933a: 21 . ST in AMNH,

CAS (1). Type locality: USA: NY, Long Is.,

Cold Spring Harbor. consors (Walker), 1852 –1853: 213 ( Dolichopus ).

HT in BMNH. Type locality: USA. filifer Aldrich, 1896b: 314 . ST in USNM (5),

SEMC (4), CUIC. Type locality: St. Vincent

Is. DISTRIBUTION: FL; MEX, Neotropics. flagellatus (Harmston), 1952: 293 ( Hercostomus );

comb. by Foote et al. (1965). HT in USNM.

Type locality: USA: AL, Mobile. DISTRIBU­

TION:?AL (immigrant, found in hold of ship containing cargo of bananas from Panama);

Neotropics [ Robinson (1970b)]. floridensis Robinson, 1964: 181 . HT in USNM.

Type locality: USA: FL, nr Gainsville, across from Glen Springs, on property of H.V. Weems

Jr. hybridus Melander, 1900: 141 . ST in AMNH (9).

Type locality: USA: MA, Woods Hole. DIS­

TRIBUTION: NY, ME, MA, CT, NC. magnicornis Van Duzee, 1927a: 146 . HT in CAS.

Type locality: USA: ID, Lewiston. minutus Van Duzee, 1921b: 127 . HT in CAS.

Type localities: USA: FL, Bradentown ; Anna

Maria Key. nigrocaudatus Van Duzee, 1918: 47 . HT in

SEMC. Type locality: USA: MT, Beaver

Creek, 6300 ft. ovatus Van Duzee, 1914c: 436 . HT in CUIC.

Type locality: USA: GA, Decatur Co. DISTRI­

BUTION: MI to NY, s. to FL (IN, OH, MD, NC,

SC, GA);?Neotropics [ Robinson (1964)]. propinquus Wheeler, 1899: 18 . ST in AMNH.

Type localities: USA: FL, Charlotte Harbor ;

Ormund. DISTRIBUTION: FL, MA;?Neotropics

[ Robinson (1970b)]. pumilio Loew, 1872: 90 (Cent. 10, no. 63). ST in

MCZ. Type locality: USA: TX. DISTRIBUTION:

TX, NE, IA, MI, IN, TN, MD, VA; MEX

[ Robinson (1964); Harmston & Rapp (1983)]. quadrinotatus Aldrich, 1902: 81 . ST in USNM

(8). Type locality: Grenada Is. DISTRIBUTION:

TX, FL; Neotropics.

Genus PELASTONEURUS Loew, 1861a

PARACLEIUS Bigot, 1859: 215 , 227. Type species: Dolichopus heteronevrus Macquart , by monotypy. Erroneously listed as a senior synonym of Paraclius Loew in Foote et al. (1965); see comments under Paraclius Loew.

PELASTONEURUS Loew, 1861a: 36 . Type species: Pelastoneurus vagans Loew [Nearctic], designation by Coquillett, 1910c: 586. Application to the I.C.Z.N. has been made for the suppression of Paracleius Bigot ( Brooks et al., 2002) .

METAPELASTONEURUS Aldrich, 1894: 152 . Type species: Metapelastoneurus kansensis Aldrich , by monotypy.

PARACLIUS Kertész, 1909: 230 . Type species: Dolichopus heteronevrus Macquart , automatic. Unjustified emendation.

PALASTONEURUS , error by Parent (1933d: 249).

PELASTRONEURUS , error by Robinson (1964: 177).

NOTES: Pelastoneurus has been recorded from the Nearctic, Neotropical, Afrotropical,

and Oriental realms, as well as the Hawaiian

Islands. Most species have been described from the New World and many more await discovery, especially in the Neotropics. Species of Pelastoneurus are found in association with freshwater seepages and mudflats,

and on wet soil or on low herbaceous vegetation near water. The immatures of P. vagans Loew inhabit mud substrates in various aquatic and semiaquatic habitats ( Corpus,

1986a).

REFERENCES: Van Duzee (1923b, rev.);

Robinson (1964; key, eastern U.S.).

abbreviatus Loew, 1864a: 94 (Cent. 5, no. 89). ST in MCZ. Type locality: USA: NY, New Rochelle. DISTRIBUTION: ON to NS, s. to TX and FL (IL, QC, NY, NC),?MT [ Robinson (1964)].

aldrichi Van Duzee, 1923b: 36 . HT in USNM. Type locality: USA: UT, Brigham.

angulatus Robinson, 1964: 178 . HT in USNM. Type locality: USA: NC, Beaufort, Duke University Marine Biological Station.

asciaeformis Becker, 1922a: 63 . HT in ZMHB. Type locality: USA: GA.

aurifacies Van Duzee, 1923b: 35 . HT in CAS. Type locality: USA: FL, Bradentown. DISTRIBUTION: AL, FL.

minutus Van Duzee, 1933a: 27 . HT in AMNH. Type locality: USA: AL, Mobile Co., Baraland.

barri Harmston, 1972: 153 . HT in CAS. Type locality: USA: CA, Inyo Co., Tecopa Hot Springs.

bifrons (Walker), 1852 –1853: 212 ( Dolichopus ). ST in BMNH. Type locality: USA.

brevis Robinson, 1964: 179 . HT in USNM. Type locality: USA: GA, Montgomery Co., Ailey. DISTRIBUTION: AL, GA, FL, NC.

cognatus Loew, 1861a: 40 . HT in MCZ. Type locality: USA: ‘‘ Middle States’ ’. DISTRIBUTION: TX, IL, FL; MEX.

comatus Robinson, 1992: 576 . HT in USNM. Type locality: USA: DC, Roosevelt Is. DISTRIBUTION: DC, NC [Robinson (personal commun.)]

cyaneus Wheeler, 1899: 17 . ST in AMNH. Type localities: USA: CA, on road from Monterey to Del Monte; WY, Lusk. DISTRIBUTION: OR to SD, s. to CA (NV, AZ, NE); MEX [ Robinson (1964); Harmston & Rapp (1983)].

dissimilipes Wheeler, 1899: 16 . ST in AMNH. Type locality: USA: CA, Monterey. DISTRIBUTION: CA, AZ.

nigrescens Wheeler, 1899: 78 , 80; alternate orig. spelling. Type locality: USA: CA, Monterey .

floridanus Wheeler, 1899: 13 . ST in AMNH. Type locality: USA: FL, St. Angustine. DISTRIBUTION: TX, FL, NC; Neotropics [ Robinson (1970b)].

furcifer Loew, 1872: 91 (Cent. 10, no. 64). ST in MCZ. Type locality: USA: TX.

hebes (Walker), 1852 –1853: 213 ( Dolichopus ). HT in BMNH. Type locality: USA.

heteronevrus (Macquart), 1850: 432 (1850: 128) ( Dolichopus ). HT in? MNHN. Type locality: North America (listed under Paracleius by Foote et al. (1965)).

heteroneurus ; error or unjustified emendation listed by Kertész (1909) and Van Duzee (1923b).

heteropterus; error listed by Foote et al. (1965).

ineptus (Walker), 1852 –1853: 214 ( Dolichopus ). HT in BMNH. Type locality: USA.

irrasus (Walker), 1848 –1849: 667 ( Dolichopus ). HT in BMNH. Type locality: USA: FL. DISTRIBUTION: FL;?Neotropics [ Robinson (1970b)].

kansensis (Aldrich), 1894: 153 ( Metapelastoneurus ). ST in SEMC (2). Type locality: USA: KS. DISTRIBUTION: SD, NE, KS, IL, MI, IN [ Harmston & Rapp (1983)].

laetus Loew, 1861a: 38 . ST in MCZ. Type locality: USA: GA. DISTRIBUTION: NE and MI to QC, s. to LA and FL (IN, OH, NY, MD, DC, VA, NC, GA) [ Robinson (1964); Harmston & Rapp (1983)].

falcatus Aldrich, 1904: 277 . ST in USNM (3). Type locality: CAN: QC, Montreal Is.

ramosus Van Duzee, 1923b: 33 . HT in CAS. Type locality: USA: VA, Chesterfield Co., Petersburg.

lamellatus Loew, 1864a: 95 (Cent. 5, no. 90). HT in MCZ. Type locality: USA: NY. DISTRIBUTION: IA to ME, s. to TX, MS, and FL (NY, NH, MA, NJ, MD, NC) [ Robinson (1964)].

cristatus Van Duzee, 1924a: 105 . HT in MCZ. Type locality: USA: MA, Eastham.

latifacies Van Duzee, 1930d: 123 . HT in CAS. Type locality: USA: CA, Lake Co., Clear Lake, Konocti Bay.

longicauda Loew, 1861a: 37 . HT in MCZ. Type locality: USA: NY. DISTRIBUTION:?CA, NY, NC, FL.

quadricincta Van Duzee, 1928d: 89 . HT in USNM. Type locality: USA: NC, Willard.

lugubris Loew, 1861a: 38 . HT in MCZ. Type locality: USA: NY, Trenton Falls. DISTRIBUTION: QC, MI, VT to TX, LA, and FL (IL, IN, NY, MA); MEX, Hawaiian Is. [ Robinson (1964); Evenhuis (1996); Beaulieu & Wheeler (2001)].

maculipes (Walker), 1852 –1853: 214 ( Dolichopus ). HT in BMNH. Type locality: USA.

neglectus Wheeler, 1899: 12 . ST in AMNH. Type localities: USA: WI, Milwaukee ; IL, Chicago. DISTRIBUTION: WI to NH, s. to LA and GA (TX, MO, IL, TN, IN, MI, NY, VA, SC) [ Robinson (1964)].

nigricornis Van Duzee, 1923b: 39 . HT in USNM. Type locality: USA: MO, Atherton. DISTRIBUTION: TX, LA, MO, TN.

penicillatus Parent, 1929a: 181 . HT in MLUH. Type locality: USA: TX.

occidentalis Wheeler, 1899: 13 . ST in AMNH. Type locality: USA: CA, Pacific Grove. DISTRIBUTION: CA, OR,?VA.

parvus Aldrich, 1904: 276 . HT in USNM. Type locality: USA: LA, Opelousas. DISTRIBUTION: LA, GA, FL.

potomacus Robinson, 1992: 578 . HT in USNM. Type locality: USA: DC, Roosevelt Is.

proximus Aldrich, 1904: 278 . ST in CNC (1). Type locality: USA: LA, Opelousas. DISTRIBUTION: TX, LA, TN, IN, MI, NC.

arboreus Van Duzee, 1923b: 34 . HT in USNM. Type locality: USA: IN, Lafayette.

scutatus Aldrich, 1904: 276 . HT in USNM. Type locality: FL, Biscayne Bay.

semiplumatus Becker, 1922a: 69 . HT in DEI. Type locality: America. Possibly not Nearctic.

seticauda Van Duzee, 1930b: 85 . HT in SEMC. Type locality: USA: MO, Atherton. DISTRIBUTION: MO, TN.

stentorius Harmston, 1971: 88 . HT in FSCA. Type locality: USA: GA, Oatland Is.

taeniatus Becker, 1922a: 70 . HT in ZMHB. Type locality: Virgin Is.: St. Jean. DISTRIBUTION: FL; Neotropics ( Dominica). NEW RECORD.

tibialis Van Duzee, 1923b: 41 . ST in? Type locality: USA: NM, Jemezsprings Mts. DISTRIBUTION: UT, NM [ Knowlton & Harmston (1937)].

umbripictus Becker, 1922a: 73 . ST in?ZMHB. Type localities: USA: Carolina ; Colombia: Baranquilla, Bogota. DISTRIBUTION: TX, MI, NC, FL ; MEX, Neotropics [ Robinson (1964)].

vagans Loew, 1861a: 39 . ST in MCZ, CNC (1). Type locality: USA: ‘‘Middle States’’. DISTRIBUTION: AK, OR, UT to QC, s. to NM, TX, and GA, Bermuda; MEX,?Neotropics [ Knowlton & Harmston (1937); Robinson (1964, 1970b); Woodley & Hilburn (1994)].

vagrans; error by Snodgrass (1904: 290).

longilamellatus Parent, 1929a: 180 . ST in MLUH. Type localities: USA: TX; GA.

varius (Walker), 1852 –1853: 215 ( Dolichopus ). HT in BMNH. Type locality: USA. DISTRIBUTION: GA, FL.

pictipennis Wheeler, 1899: 14 . ST in AMNH. Type localities: USA: FL, St. Augustine ; Gotha.

punctipennis (not Say); error by Parent (1929a: 170).

wheelerii Melander, 1900: 140. HT in?AMNH. Type locality: USA: TX, along Colorado River S of Austin. DISTRIBUTION: CA, TX, MI, TN, NC, SC, GA.

wheeleri ; error by Foote et al. (1965: 502).

Genus TACHYTRECHUS Stannius, 1831

AMMOBATES Stannius, 1831: 33 . Incorrect original spelling by revision of Haliday (1851: 173); preocc. by Ammobates Latreille, 1809 .

TACHYTRECHUS Stannius, 1831: 261 . Erroneously treated as nomen nudum by Sabrosky (1999: 297–298) and previous regional catalogs. Type species: Ammobates notatus Stannius , designation by Rondani, 1856: 143 from species first included by Stannius (1831: 268– 270).

HAMMOBATES , subsequent misspelling by Rondani, 1856: 143.

STANNIA Rondani, 1857: 14 . Type species: Ammobates notatus Stannius , automatic. Unnecessary n. name for Ammobates Stannius, 1831 .

GONGOPHORA Philippi, 1875: 86 . Type species: Gongophora medinae Philippi , by monotypy. Synonymized by Robinson (1970b).

CONGOPHORA Philippi, 1875: 86 . Incorrect original spelling by present revision.

POLYMEDON Osten Sacken, 1877: 317 . Type species: Polymedon flabellifer Osten Sacken , by monotypy. Synonymized by Robinson (1970b).

MACELLOCERUS Mik, 1878b: 5 . Type species: Tachytrechus moechus Loew , by original designation.

PSILISCHIUM Becker, 1922a: 93 . Type species: Psilischium laevigatum Becker , by monotypy. Synonymized by Robinson (1970b).

TETRECHUS , error by Van Duzee (1924d: 43).

GONGROPHORA , subsequent misspelling by Porter (1929: 230), repeated by Robinson (1970b: 53).

TACHYTERECHUS , error by Dyte (1975: 238).

NOTES: Tachytrechus is known from every realm; however, the Neotropics has the majority of the species. Tachytrechus species are hygrophilous and riparian, and mainly occur on open, well­lit, wet, sandy soils or rocky substrates near or in open water. Larval and pupal stages have been discovered along temporal pools ( Dyte, 1959) and in freshet seeps and mud flats ( Corpus, 1988).

REFERENCES: Greene (1922; key); Van Duzee (1927d; key, as Polymedon ); Harmston

& Knowlton (1940a; key to males); Robinson (1964; key, eastern U.S.).

albonotatus (Loew), 1864b: 102 ( Paraclius ). ST in?MCZ. Type locality: USA: LA, New Orleans. DISTRIBUTION: UT to VT, s. to LA and FL; Neotropics.

angulatus (Van Duzee), 1914c: 436 ( Paraclius ). HT in CAS. Type locality: USA: NY, Erie Co., Colden. DISTRIBUTION: TN, NY, NC.

angustipennis Loew, 1862: 213 (Cent. 2, no. 64). HT in MCZ. Type locality: USA: DC. DISTRIBUTION: CA and UT to DC, s. to FL (NV, AZ, WY, NM, TX, NC, GA); MEX, Neotropics, Hawaiian Is. [ Knowlton & Harmston (1937); Robinson (1964, 1970b); Evenhuis (1996, 1997a); Vockeroth (personal commun.)].

argyropus Becker, 1922a: 79 . HT in NMW. Type locality: America. Possibly not Nearctic.

auratus (Aldrich), 1896a: 83 ( Macellocerus ). ST in USNM (3). Type locality: USA: ID, Moscow. DISTRIBUTION: WA, CA, NV, ID, UT, AZ, CO [ Corpus (1988)].

binodatus Loew, 1866b: 46 (Cent. 7, no. 84). HT in MCZ. Type locality: USA: NY, Saratoga. DISTRIBUTION: AK, ON, QC, and NS, s. to NE and TN (MI, IN, OH, NY, NJ) [ Robinson (1964); Harmston & Rapp (1983); Bissett (1987)].

boharti Harmston, 1968: 15 . HT in UCDC. Type locality: USA: CA, Sierra Co., Yuba Pass.

californicus (Harmston & Knowlton), 1943a: 105 ( Polymedon ). HT in CAS. Type locality: USA: CA, Sonora.

canacolli Brooks , NEW NAME for Paraclius utahensis Harmston & Knowlton, 1946b proposed herein by second author in recognition of the contribution of the CanaColl Foundation to the production of this catalog. Type locality: USA: UT, St. George.

utahensis (Harmston & Knowlton), 1946b: 23 ( Paraclius ); preocc. by Tachytrechus utahensis Harmston & Knowlton, 1940a . HT in USNM. Type locality: USA: UT, St. George. NEW COM­ BINATION .

castus (Wheeler), 1899: 6 ( Polymedon ). HT in AMNH. Type locality: USA: AZ, Grand Canon. DISTRIBUTION: AZ, UT.

castor; error listed by Foote et al. (1965).

dilaticosta (Van Duzee), 1927d: 124 ( Polymedon ). HT in CNC. Type locality: USA: AZ, Grand Canon, Bright Angel Trail.

duplicatus Harmston, 1972: 157 . HT in CAS. Type locality: USA: OR, Baker.

flabellifer (Osten Sacken), 1877: 317 ( Polymedon ). ST in MCZ. Type locality: USA: CA, Sonoma Co., Los Guilucos. DISTRIBUTION: WA, OR, CA, ID, UT.

flavitibialis (Van Duzee), 1930c: 71 ( Polymedon ). HT in SEMC. Type locality: USA: southern AZ.

floridensis Aldrich, 1896a: 82 . ST in USNM (4). Type locality: USA: FL.

granditarsis Greene, 1922: 17 . HT in USNM. Type locality: USA: CA, Jacumba Spring. DISTRIBUTION: CA, UT, AZ, WY [Vockeroth (personal commun.)].

greenei Foote et al., 1965: 503 ; n. name for Tachytrechus bipunctatus Greene. Type locality: USA: ID, St. Anthony. DISTRIBUTION: ID, UT, AB .

bipunctatus Greene, 1922: 9 ; preocc. by Dolichopus bipunctatus Macquart, 1842 , now Tachytrechus . HT in USNM. Type locality: USA: ID, St. Anthony.

harmstoni Meuffels & Grootaert, 1999: 292 ; n. name for Tachytrechus californicus Harmston & Knowlton, 1963 . Type locality: USA: CA, Trinity Co., Trinity River Camp .

californicus Harmston & Knowlton, 1963: 234 ; preocc. by Tachytrechus californicus (Harmston & Knowlton), 1943a , previously in Polymedon . HT in CAS. Type locality: USA: CA, Trinity Co., Trinity River Camp.

indianus (Harmston & Knowlton), 1946b: 24 ( Paraclius ). HT in?USNM. Type locality: USA: IN, Indianapolis. DISTRIBUTION: TN, IN, OH, NC, SC, FL.

keiferi (Van Duzee), 1927a: 146 ( Paraclius ). HT in CAS. Type locality: MEX: Baja California, Cedros Is. DISTRIBUTION: OR, CA, ID, AZ; MEX. NEW COMBINATION, NEW RECORD.

laticrus Van Duzee, 1918: 46 . LT designated by Arnaud (1979: 292) in CAS. Type locality: USA: NJ, Manahawkin.

latitarsus (Parent), 1929a: 182 ( Polymedon ). HT in MLUH. Type locality: unknown. DISTRIBUTION: America [ Robinson (1970b)].

parenti Robinson, 1970b: 55 ; unnecessary n. name for Polymedon latitarsus Parent , not latitarsis Becker, 1922a: 81 . NEW SYNONYMY.

latitarsis ; error by Robinson (1970b: 55).

mchughi Harmston, 1972: 156 . HT in CAS. Type locality: USA: OR, Latourele Falls. DISTRIBUTION: OR, CA.

milleri Harmston, 1966b: 223 . HT in CAS. Type locality: USA: OR, Crater Lake.

moechus Loew, 1861a: 40 . ST in MCZ, CNC (1). Type locality: USA: NY, Trenton Falls. DISTRIBUTION: ON, QC, and NS, s. to NE, IA, and SC (IL, TN, IN, NY, NH, NC) [ Robinson (1964); Harmston & Rapp (1983); Bissett (1987)].

mysticus (Becker), 1922a: 52 ( Paraclius ). ST in ZMHB. Type locality: USA: GA.

nigrifemoratus (Van Duzee), 1927d: 125 ( Polymedon ). HT in CAS. Type locality: USA: AZ, Oak Creek Canon, 6000 ft. DISTRIBUTION: AZ, NM.

nimius (Aldrich), 1901: 334 ( Polymedon ). ST in USNM (2). Type localities: MEX: Guerrero, Omilteme , 8000 ft; Sierra de las Aguas Escondidas , 7000 ft. DISTRIBUTION: ID, UT, AZ ; MEX, Neotropics [ Robinson (1970b)].

nigricornis (Van Duzee), 1929: 53 ( Polymedon ). HT in USNM. Type locality: Guatemala: Alta Vera Paz, Tamau [ Robinson (1970b)].

nitidus (Van Duzee), 1927d: 125 ( Polymedon ). HT in CNC. Type locality: USA: AZ, Grand Canon, Bright Angel Trail.

olympiae (Aldrich), 1896a: 83 ( Macellocerus ). ST in USNM (4). Type locality: USA: WA, Olympia. DISTRIBUTION: WA to IL, s. to CA.

protervus Melander, 1900: 143 . ST in AMNH. Type locality: USA: NJ, Clementon. DISTRIBUTION: NH to FL (NJ, MD, NC, SC) [ Robinson (1964)].

junctus Coquillett, 1910a: 125 . HT in USNM. Type locality: USA: NJ, Manahawkin.

rotundipennis Greene, 1922: 5 . HT in USNM. Type locality: USA: DE. DISTRIBUTION: DE, NJ.

sanus Osten Sacken, 1877: 316 . ST in MCZ. Type locality: USA: CA, Sierra Nevada, Sierra Co., Webber Lake. DISTRIBUTION: WA, CA, AB, UT, CO [ Knowlton & Harmston (1937)]. spinitarsis (Van Duzee), 1924d: 43 ( Tetrechus ). HT in CAS. Type locality: USA: CA, Tulare Co., Giant Forest, 9000 ft. NEW SYNONYMY.

simulatus Greene, 1922: 4 . HT in USNM. Type locality: USA: ID, St. Anthony. DISTRIBUTION: BC, OR, ID, MT, UT.

tahoensis Harmston & Knowlton, 1940a: 114 . HT in USNM. Type locality: USA: CA, Tahoe.

tenuiseta Greene, 1922: 10 . HT in USNM. Type locality: USA: OR, Newport. DISTRIBUTION: BC, OR, CA, NM [Vockeroth (personal commun.)].

utahensis Harmston & Knowlton, 1940a: 112 . HT in USNM. Type locality: USA: UT, Torrey.

volitans Melander, 1900: 143 . ST in AMNH (2). Type locality: USA: WY, 12 mi NW of Lusk. DISTRIBUTION: WY, TX.

vorax Loew, 1861a: 41 . ST in MCZ, CNC (1). Type locality: USA: DC. DISTRIBUTION: CA to QC, s. to TX and NC (CO, SD, NE, IA, ON, MI, IN, NH, NJ, DC, VA) [ Steyskal (1947a); Robinson (1964); Harmston & Rapp (1983)].