Systenus, Loew, 1857 a

Pollet, M. A. A., Brooks, S. E. & Cumming, J. M., 2004, Catalog Of The Dolichopodidae (Diptera) Of America North Of Mexico, Bulletin of the American Museum of Natural History 2004 (283), pp. 1-114 : 58-60

Genus SYSTENUS Loew, 1857a

SYSTENUS Loew, 1857a: 34 . Type species: Rhaphium adpropinquans Loew [Palearctic], designation by Foote et al., 1965: 517. [ = pallipes (Roser), 1840 ]; listed in Rhaphiinae by Foote et al. (1965).

NOTES: Systenus is known from the Holarctic, Neotropics, and Australia. Most species have not been captured in the wild as adults, but reared from larvae found in moist tree holes, ulcerated or decaying wood, and sap runs.

REFERENCES: Wirth (1952; rev.); Steyskal (1970).

albimanus Wirth, 1952: 240 . HT in USNM. Type localities: USA: VA, Falls Church. DISTRIBUTION: TX, AL, TN, VA.

apicalis Wirth, 1952: 237 . HT in USNM. Type locality: USA: VA, Falls Church. DISTRIBUTION: AL, VA, MD.

californicus Harmston, 1968: 16 . HT in CAS. Type locality: USA: CA, Marin Co., Mill Valley.

eucercus Steyskal, 1970: 107 . HT in?USNM. Type locality: USA: CT, Manchester. DISTRIBUTION: ON, CT [Vockeroth (personal commun.)].

minutus (Van Duzee), 1913b: 60 ( Neurigona ). HT in USNM. Type locality: USA: PA, Philadelphia. DISTRIBUTION: KY, TN, AL, PA.

shannoni Wirth, 1952: 240 . HT in USNM. Type locality: USA: MD, Potomac River , Plummer’s Is. DISTRIBUTION: ON, NH, MD [Vockeroth (personal commun.)].

Genus THRYPTICUS Gerstäcker, 1864

THRYPTICUS Gerstäcker, 1864: 43 . Type species: Thrypticus smaragdinus Gerstäcker [Palearctic], by monotypy.

APHANTOTIMUS Wheeler, 1890: 375 . Type species: Aphantotimus willistoni Wheeler , designation by Coquillett, 1910c: 508.

XANTHOTRICHA Aldrich, 1896b: 339 . Type species: Xanthotricha cupulifera Aldrich , designation by Robinson, 1970b: 21.

SUBMEDETERUS Becker, 1917 –1918: 360. Type species: Submedeterus cuneatus Becker , by monotypy.

NOTES: Thrypticus occurs in all realms and is well represented in the New World. The Nearctic fauna is comprised of small species occurring on herbaceous vegetation on stream banks and margins. Larval stages of Thrypticus are phytophagous and feed as miners of Cyperaceae , Poaceae , Juncaceae , and Pontederiaceae ( Dyte, 1993) .

REFERENCES: Van Duzee (1921b; key); Parent (1929a; key); Robinson (1964; key, eastern North America).

abdominalis (Say), 1829 –1830: 169 (also in

LeConte, 1859: 362) ( Chrysotus ). ST destroyed. Type locality: USA: IN. DISTRIBUTION:

KS to IN, MI, and PA, s. to LA, and NC; Neotropics [ Robinson (1970b)]. cupulifer (Aldrich), 1896b: 339 (Xanthotrichia) .

ST in SEMC (3). Type locality: St. Vincent Is. aurinotatus Van Duzee, 1915c: 87 . HT in CUIC.

Type locality: USA: GA, Okefenokee Swamp,

Billy’s Is. comosus Van Duzee, 1915c: 86 . HT in CUIC.

Type locality: USA: NY, Ellis ; CAN: ON, Toronto. DISTRIBUTION: MI, ON, NY.

deficiens Robinson, 1980: 469 . HT in USNM. Type locality: USA: MD, Prince Georges Co., Patuxent Wildlife Refuge.

dissectus Robinson, 1964: 118 . HT in CNC. Type locality: USA: NC, Franklin, ca. 2000 ft.

fosteri Robinson, 1980: 470 . HT in USNM. Type locality: USA: MD, Prince Georges Co., Patuxent Wildlife Refuge.

fraterculus (Wheeler), 1890: 376 ( Aphantotimus ). ST in AMNH. Type locality: USA: WI, Milwaukee Co. DISTRIBUTION: AK, MN to ON and MA, s. to CA, TN, and FL (UT, WY, CO, AZ, NE, TX, IA, WI, VA, NC, NS), Bermuda; MEX, Neotropics, Palearctic (Siberia) [ Robinson (1970b); Harmston & Rapp (1983); Bissett (1987); Woodley & Hilburn (1994); Bickel (personal commun.); Robinson (personal commun.)].

grogani Robinson, 1980: 472 . HT in USNM. Type locality: USA: MD, Prince Georges Co., Patuxent Wildlife Refuge. DISTRIBUTION: MD, VA [Robinson (personal commun.)].

longicauda Van Duzee, 1921b: 126 . HT in CAS. Type locality: USA: CA, San Francisco.

minutus Parent, 1929a: 186 . HT in MLUH. Type locality: USA: TX. DISTRIBUTION: CA, TX, PA; MEX, Neotropics (West Indies) [ Robinson (1964; 1975); Dyte (1993)].

insulanus Van Duzee, 1933b: 14 ; synonymized by Dyte (1993). HT in AMNH. Type locality: St. Lucia, Windward Is .

insularis ; error by Bennet (1968).

muhlenbergiae Johannsen & Crosby, 1913: 164 . ST in CUIC, USNM (1). Type locality: USA: NY, Ithaca, Cornell University Campus, Cascadilla Creek. DISTRIBUTION: CO, TX, MO, TN, ON, OH, NY, QC, MD, NC, GA [Bickel (personal commun.)].

nigripes Van Duzee, 1921b: 125 . HT in CAS. Type locality: USA: NV, Wells. DISTRIBUTION: CA, NV, UT, CO [Bickel (personal commun.)].

squamiciliatus Harmston & Rapp, 1968: 250 . HT in CAS. Type locality: USA: NE, Crawford.

tectus Van Duzee, 1915c: 87 . HT in CAS. Type locality: USA: NY, Erie Co., East Aurora. DISTRIBUTION: IA, ON, QC to CT, s. to GA (IL, IN, KY, NY, PA, DC, VA, NC) [ Robinson (1964); Bickel (personal commun.)].

vietus Van Duzee, 1915c: 86 . HT in CAS. Type locality: USA: FL, Bradentown.

violaceus Van Duzee, 1927e: 5 . HT in AMNH. Type locality: Puerto Rico: Arecibo. DISTRIBUTION: TX, NC, FL; Neotropics ( West Indies ) [ Robinson (1970b); Bickel (personal commun.)] .

setosus Robinson, 1964: 118 ; listed as synonym by Robinson (1970b). HT in FSCA. Type locality: USA: FL, Gainesville [ Robinson (1964, 1970b)].

willistoni (Wheeler), 1890: 376 ( Aphantotimus ). ST in AMNH. Type locality: USA: WI, Milwaukee Co. DISTRIBUTION: BC, WY, MB, IA, WI, ON, QC, PA, NB, MA, and NS, s. to TN, MO, and NC;?Neotropics [ Robinson (1970b); Bickel (personal commun.)].