Namadytes maculiventris (Hesse, 1969)

Dikow, Torsten & Leon, Stephanie, 2014, Review of the genus Namadytes Hesse, 1969 (Insecta: Diptera: Mydidae: Syllegomydinae), Biodiversity Data Journal 2, pp. 1071-1071 : 1071

Namadytes maculiventris (Hesse, 1969)

Namamydas maculiventris Hesse, 1969: 285.

Namadytes maculiventris (Hesse, 1969) new combination sensu Hesse 1972: 142.

Namadytes pallidus Hesse, 1972: 146 syn. nov. (ZooBank LSID). Type locality of Namadytes pallidus : Namibia: Karas: Keetmanshoop, 48 km SE (26°53'00''S 018°26'00''E).

Materials

Type status: Holotype. Occurrence: catalogNumber: SAM-DIP-A007147 ; recordedBy: SAM Museum Staff; sex: 1 male; lifeStage: Adult; previousIdentifications: Namamydas maculiventris by A. Hesse in 1969; Taxon: scientificNameID: urn:lsid:zoobank.org:act:78BC184A-C090-4A75-9DDC-EF4CA95CB254; scientificName: Namadytesmaculiventris Hesse, 1969; family: Mydidae; genus: Namadytes; specificEpithet: maculiventris; scientificNameAuthorship: Hesse, 1969; Location: country: South Africa; stateProvince: Northern Cape; locality: Vioolsdrift ; verbatimCoordinates: 28°46'10''S 017°37'37''E; decimalLatitude: -28.76944; decimalLongitude: 17.62694; Identification: identifiedBy: A. Hesse; dateIdentified: 1972; Event: eventDate: 1935-03-00; Record Level: institutionCode: SAMC; collectionCode: Insects; basisOfRecord: PreservedSpecimen Type status: Holotype. Occurrence: catalogNumber: SAM-DIP-A007148 ; recordedBy: J. Rozen E. Martinez; sex: 1 male; lifeStage: Adult; otherCatalogNumbers: AAM-000454; previousIdentifications: Namadytes pallidus by A. Hesse in 1972; Taxon: scientificNameID: urn:lsid:zoobank.org:act:78BC184A-C090-4A75-9DDC-EF4CA95CB254; scientificName: Namadytesmaculiventris Hesse, 1969; family: Mydidae; genus: Namadytes; specificEpithet: maculiventris; scientificNameAuthorship: Hesse, 1969; Location: country: Namibia; stateProvince: Karas; locality: Keetmanshoop, 48 km SE ; verbatimCoordinates: 26°46'47''S 018°32'15''E; decimalLatitude: -26.77972; decimalLongitude: 18.5375; Identification: identifiedBy: T. Dikow S. Leon; dateIdentified: 2013; Event: eventDate: 1968-10-30; Record Level: institutionCode: SAMC; collectionCode: Insects; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: SAM-DIP-A007148 ; recordedBy: J. Rozen E. Martinez; sex: 1 male; lifeStage: Adult; otherCatalogNumbers: AAM-000455; previousIdentifications: Namadytes pallidus by A. Hesse in 1972; Taxon: scientificNameID: urn:lsid:zoobank.org:act:78BC184A-C090-4A75-9DDC-EF4CA95CB254; scientificName: Namadytesmaculiventris Hesse, 1969; family: Mydidae; genus: Namadytes; specificEpithet: maculiventris; scientificNameAuthorship: Hesse, 1969; Location: country: Namibia; stateProvince: Karas; locality: Keetmanshoop, 48 km SE ; verbatimCoordinates: 26°46'47''S 018°32'15''E; decimalLatitude: -26.77972; decimalLongitude: 18.5375; Identification: identifiedBy: T. Dikow S. Leon; dateIdentified: 2013; Event: eventDate: 1968-10-30; Record Level: institutionCode: SAMC; collectionCode: Insects; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: USNMENT00779997 ; recordedBy: M. Schwarz; sex: 1 male; lifeStage: Adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:78BC184A-C090-4A75-9DDC-EF4CA95CB254; scientificName: Namadytesmaculiventris Hesse, 1969; family: Mydidae; genus: Namadytes; specificEpithet: maculiventris; scientificNameAuthorship: Hesse, 1969; Location: country: Namibia; stateProvince: Karas; locality: Aus ; verbatimCoordinates: 26°40'00''S 016°16'00''E; decimalLatitude: -26.66667; decimalLongitude: 16.26667; Identification: identifiedBy: T. Dikow S. Leon; dateIdentified: 2013; Event: eventDate: 1990-02-11; Record Level: institutionCode: USNM; collectionCode: Entomology; basisOfRecord: PreservedSpecimen

Description

Male: Fig. 7.

Head: brown, facial gibbosity light brown, in general densely grey pubescent; width distinctly greater than thorax, interocular distance on vertex larger than at ventral eye margin, vertex between compound eyes ± horizontally straight, medially only slightly below dorsal eye margin, parafacial area about as wide as ½ the width of central facial gibbosity; facial gibbosity distinct, well-developed and discernible in lateral view; mystax white, densely covering entire facial gibbosity; frons not elevated, predominantly apubescent; vertex entirely white pubescent; postgena white pubescent; setation: vertex white, frons white, ocp setae white, pocl macrosetae white; ocellar triangle apubescent; proboscis yellow, very short, vestigial, knob-like; labellum small, as wide as prementum, as long as prementum, unsclerotized laterally; maxillary palpus cylindrical, yellow, minute.

Antenna: brown, scape and pedicel white setose dorsally and ventrally; postpedicel cylindrical in proximal ½, symmetrically bulbous in distal ½, ≥ 7.0 times as long as combined length of scape and pedicel, asetose; apical seta-like sensory element situated apically in cavity on postpedicel.

Thorax: brown, lightly grey pubescent; scutum predominantly black, anteriorly and laterally yellow to light brown, surface entirely smooth, lightly grey pubescent, scutal setation comprised of long white setae with distinct rows of long dorsocentral setae and dense lateral scutal setae; dc setae pre- and postsuturally white, acr setae absent, lateral scutal setae white, npl setae 0, spal setae 0, pal setae 0; antepronotum dorso-medially with V-shaped indentation; postpronotal lobe yellow, white pubescent; proepisternum, lateral postpronotum, and postpronotal lobe long white setose; scutellum apubescent, asetose, apical scutellar setae absent; mesopostnotum, anatergite, and katatergite grey pubescent, mesopostnotum asetose, anatergite asetose, katatergite long white setose; katatergite ± flat; anterior anepisternum white setose, supero-posterior anepisternum long white setose; posterior anepimeron asetose, katepimeron asetose; metanepisternum grey pubescent, asetose, metepimeron ± flat, yellow, grey pubescent, long white setose; infra-halter sclerite white setose.

Leg: yellow to light brown, setation predominantly white; pro, mes, and met coxa lightly white pubescent, long white setose; met trochanter setose medially; femur yellow to light brown, met femur ± cylindrical only slightly wider than pro and mes femur, in distal ½ macrosetose, 1 antero-ventral and 1 postero-ventral row of macrosetae, postero-ventrally long white, erect setose with setae arranged in distinct row; pro, mes, and met tibia straight, met tibia cylindrical, ventral keel absent, latero-posteriorly long white, erect setose with setae arranged in distinct row; pro and mes tarsomere 1 longer than tarsomere 2, but less than combined length of tarsomeres 2-3, met tarsomere 1 as long as combined length of tarsomeres 2-4; pulvillus well-developed, as long as well-developed claw, and as wide as base of claw; empodium absent.

Wing: length = 9.6-12.2 mm; hyaline throughout, veins light brown, microtrichia absent; cells r1, r4, r5, m3, + cup closed; C terminates at junction with R1; R4 terminates in R1; R5 terminates in R1; stump vein (R3) at base of R4 present, short not reaching R2; R4 and R5 widest apart medially; r-m distinct, R4+5 and M1 apart, connected by crossvein; M1 straight at r-m (not curving anteriorly), M1 (or M1+M2) terminates in R1; CuA1 and CuA2 split proximally to m-cu (cell m3 narrow proximally); M3+CuA1 do not terminate together in C; A1 undulating, cell a1 wide, A1 and wing margin further apart proximally than distally; alula well-developed; halter light yellow.

Abdomen: yellow to brown; setation comprised of dense white setae, surface entirely smooth; T1-2 anteriorly yellow otherwise brown, T3 antero-medially brown otherwise yellow, T4-7 yellow to light brown; T1 and anterior ½ of T2 long white setose, remaining T short white setose; T predominantly apubescent; S1-7 yellow; S1-7 short white setose; S predominantly apubescent; T2-4 parallel-sided and not constricted waist-like; bullae on T2 black, transversely elongate, surface entirely smooth, T2 surface anterior to bullae smooth.

♂ terminalia: Fig. 8a, b, c.

Female: unknown.

Diagnosis

This large species (wing length in males 9.6-12.2 mm, females unknown) is distinguished from congeners by the entirely white pubescent vertex and postgena, by the long postpedicel (about 7 times as long as combined length of scape and pedicel), the long white scutal setation, the yellow postpronotal lobes, the densely grey pubescent mesopostnotum, anatergite, and katatergite, the yellow to light brown coloration of the legs, and the long, erect white setae dorsally on the metathoracic femur.

Distribution

Namibia (Karas) and South Africa (Northern Cape) (Fig. 6).

Discussion

Hesse 1972 (p. 148) alludes to the morphological similarity of Namadytes maculiventris and Namadytes pallidus . We regard the differences between the male holotype of Namadytes maculiventris and the male holotype and paratype of Namadytes pallidus as intraspecific variation. So far, only 4 specimens of this species, all males, have been collected in southern Namibia and Vioolsdrift in South Africa.

Type locality

South Africa: Northern Cape: Vioolsdrift (28°46'10''S, 17°37'37''E) (Fig. 6).

Biodiversity hotspot

Not known to occur in any of the southern African biodiversity hotspots (Cape Floristic Region, Maputaland-Pondoland-Albany, or Succulent Karoo) (Fig. 6).