Crocidura Wagler 1832

Wilson, Don E. & Reeder, DeeAnn, 2005, Order Soricomorpha, Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1, Baltimore: The Johns Hopkins University Press, pp. 220-311 : 224

Crocidura Wagler 1832

Crocidura Wagler 1832 , Isis: 275.

Type Species: Sorex leucodon Herman 1780

Synonyms: Afrosorex Hutterer 1986 ; Heliosorex Heller 1910 ; Leucodon Fatio 1869 ; Paurodus Schulze 1897 ; Praesorex Thomas 1913 ; Rhinomus Murray 1860 .

Species and subspecies: 172 species with 56 subspecies:

Species Crocidura aleksandrisi Vesmanis 1977

Species Crocidura allex Osgood 1910

Species Crocidura andamanensis Miller 1902

Species Crocidura ansellorum Hutterer and Dippenaar 1987

Species Crocidura arabica Hutterer and Harrison 1988

Species Crocidura arispa Spitzenberger 1971

Species Crocidura armenica Gureev 1963

Species Crocidura attenuata Milne-Edwards 1872

Species Crocidura attila Dollman 1915

Species Crocidura baileyi Osgood 1936

Species Crocidura baluensis Thomas 1898

Species Crocidura batesi Dollman 1915

Species Crocidura beatus Miller 1910

Species Crocidura beccarii Dobson 1887

Species Crocidura bottegi Thomas 1898

Species Crocidura bottegoides Hutterer and Yalden 1990

Species Crocidura brunnea Jentink 1888

Subspecies Crocidura brunnea subsp. brunnea Jentink 1888

Subspecies Crocidura brunnea subsp. pudjonica Sody 1936

Species Crocidura buettikoferi Jentink 1888

Species Crocidura caliginea Hollister 1916

Species Crocidura canariensis Hutterer, Lopez-Jurado and Vogel 1987

Species Crocidura caspica Thomas 1907

Species Crocidura cinderella Thomas 1911

Species Crocidura congobelgica Hollister 1916

Species Crocidura crenata Brosset, Dubost, and Heim de Balsac 1965

Species Crocidura crossei Thomas 1895

Species Crocidura cyanea Duvernoy 1838

Species Crocidura denti Dollman 1915

Species Crocidura desperata Hutterer, Jenkins and Verheyen 1991

Species Crocidura dhofarensis Hutterer and Harrison 1988

Species Crocidura dolichura Peters 1876

Species Crocidura douceti Heim de Balsac 1958

Species Crocidura dsinezumi Temminck 1842

Species Crocidura eisentrauti Heim de Balsac 1957

Species Crocidura elgonius Osgood 1910

Species Crocidura elongata Miller and Hollister 1921

Species Crocidura erica Dollman 1915

Species Crocidura fischeri Pagenstecher 1885

Species Crocidura flavescens (I. Geoffroy 1827)

Species Crocidura floweri Dollman 1915

Species Crocidura foetida Peters 1870

Subspecies Crocidura foetida subsp. foetida Peters 1870

Subspecies Crocidura foetida subsp. doriae Peters 1870

Subspecies Crocidura foetida subsp. kelabit Medway 1965

Species Crocidura foxi Dollman 1915

Species Crocidura fuliginosa Blyth 1855

Subspecies Crocidura fuliginosa subsp. fuliginosa Blyth 1855

Subspecies Crocidura fuliginosa subsp. dracula Thomas 1912

Species Crocidura fulvastra Sundevall 1842

Species Crocidura fumosa Thomas 1904

Species Crocidura fuscomurina Heuglin 1865

Species Crocidura glassi Heim de Balsac 1966

Species Crocidura gmelini ( Pallas 1811)

Species Crocidura goliath Thomas 1906

Subspecies Crocidura goliath subsp. goliath Thomas 1906

Subspecies Crocidura goliath subsp. nimbasilvanus Hutterer 2003

Species Crocidura gracilipes Peters 1870

Species Crocidura grandiceps Hutterer 1983

Species Crocidura grandis Miller 1911

Species Crocidura grassei Brosset, Dubost, and Heim de Balsac 1965

Species Crocidura grayi Dobson 1890

Species Crocidura greenwoodi Heim de Balsac 1966

Species Crocidura harenna Hutterer and Yalden 1990

Species Crocidura hildegardeae Thomas 1904

Species Crocidura hilliana Jenkins and Smith 1995

Species Crocidura hirta Peters 1852

Species Crocidura hispida Thomas 1913

Species Crocidura horsfieldii Tomes 1856

Species Crocidura hutanis Ruedi and Vogel 1995

Species Crocidura ichnusae Festa 1912

Species Crocidura indochinensis Robinson and Kloss 1922

Species Crocidura jacksoni Thomas 1904

Species Crocidura jenkinsi Chakraborty 1978

Species Crocidura jouvenetae Heim de Balsac 1958

Species Crocidura katinka Bate 1937

Species Crocidura kivuana Heim de Balsac 1968

Species Crocidura lamottei Heim de Balsac 1968

Subspecies Crocidura lamottei subsp. lamottei Heim de Balsac 1968

Subspecies Crocidura lamottei subsp. elegans Hutterer 1986

Species Crocidura lanosa Heim de Balsac 1968

Species Crocidura lasiura Dobson 1890

Species Crocidura latona Hollister 1916

Species Crocidura lea Miller and Hollister 1921

Species Crocidura lepidura Lyon 1908

Species Crocidura leucodon (Hermann 1780)

Species Crocidura levicula Miller and Hollister 1921

Species Crocidura littoralis Heller 1910

Species Crocidura longipes Hutterer and Happold 1983

Species Crocidura lucina Dippenaar 1980

Species Crocidura ludia Hollister 1916

Species Crocidura luna Dollman 1910

Species Crocidura lusitania Dollman 1915

Species Crocidura macarthuri St. Leger 1934

Species Crocidura macmillani Dollman 1915

Species Crocidura macowi Dollman 1915

Species Crocidura malayana Robinson and Kloss 1911

Species Crocidura manengubae Hutterer 1982

Species Crocidura maquassiensis Roberts 1946

Species Crocidura mariquensis A. Smith 1844

Subspecies Crocidura mariquensis subsp. mariquensis A. Smith 1844

Subspecies Crocidura mariquensis subsp. neavei Wroughton 1907

Subspecies Crocidura mariquensis subsp. shortridgei St. Leger 1932

Species Crocidura maurisca Thomas 1904

Species Crocidura maxi Sody 1936

Species Crocidura mindorus Miller 1910

Species Crocidura miya Phillips 1929

Species Crocidura monax Thomas 1910

Species Crocidura monticola Peters 1870

Species Crocidura montis Thomas 1906

Species Crocidura muricauda Miller 1900

Species Crocidura musseri Ruedi and Vogel 1995

Species Crocidura mutesae Heller 1910

Species Crocidura nana Dobson 1890

Species Crocidura nanilla Thomas 1909

Species Crocidura negligens Robinson and Kloss 1914

Species Crocidura negrina Rabor 1952

Species Crocidura nicobarica Miller 1902

Species Crocidura nigeriae Dollman 1915

Species Crocidura nigricans Bocage 1889

Species Crocidura nigripes Miller and Hollister 1921

Subspecies Crocidura nigripes subsp. nigripes Miller and Hollister 1921

Subspecies Crocidura nigripes subsp. lipara Miller and Hollister 1921

Species Crocidura nigrofusca Matschie 1895

Species Crocidura nimbae Heim de Balsac 1956

Species Crocidura niobe Thomas 1906

Species Crocidura obscurior Heim de Balsac 1958

Species Crocidura olivieri (Lesson 1827)

Subspecies Crocidura olivieri subsp. olivieri Lesson 1827

Subspecies Crocidura olivieri subsp. anchietae Bocage 1889

Subspecies Crocidura olivieri subsp. bueae Heim de Balsac and Barloy 1966

Subspecies Crocidura olivieri subsp. cara Dollmann 1915

Subspecies Crocidura olivieri subsp. cinereoaenea Rüppell 1842

Subspecies Crocidura olivieri subsp. darfurea Thomas and Hinton 1923

Subspecies Crocidura olivieri subsp. giffardi de Winton 1898

Subspecies Crocidura olivieri subsp. guineensis Cabrera 1903

Subspecies Crocidura olivieri subsp. hansruppi Hutterer 1981

Subspecies Crocidura olivieri subsp. hedenborgiana Sundevall 1843

Subspecies Crocidura olivieri subsp. kivu Osgood 1910

Subspecies Crocidura olivieri subsp. manni Peters 1878

Subspecies Crocidura olivieri subsp. martiensseni Neumann 1900

Subspecies Crocidura olivieri subsp. nyansae Neumann 1900

Subspecies Crocidura olivieri subsp. occidentalis Pucheran 1855

Subspecies Crocidura olivieri subsp. odorata Leconte 1857

Subspecies Crocidura olivieri subsp. spurelli Thomas 1910

Subspecies Crocidura olivieri subsp. sururae Heller 1910

Subspecies Crocidura olivieri subsp. toritensis Setzer 1956

Subspecies Crocidura olivieri subsp. zuleika Dollman 1915

Species Crocidura orientalis Jentink in Weber 1890

Subspecies Crocidura orientalis subsp. orientalis Jentink in Weber 1890

Subspecies Crocidura orientalis subsp. lawuana Sody 1936

Species Crocidura orii Kuroda 1924

Species Crocidura palawanensis Taylor 1934

Species Crocidura paradoxura Dobson 1886

Species Crocidura parvipes Osgood 1910

Species Crocidura pasha Dollman 1915

Species Crocidura pergrisea Miller 1913

Species Crocidura phaeura Osgood 1936

Species Crocidura picea Sanderson 1940

Species Crocidura pitmani Barclay 1932

Species Crocidura planiceps Heller 1910

Species Crocidura poensis Fraser 1842

Species Crocidura polia Hollister 1916

Species Crocidura pullata Miller 1911

Species Crocidura raineyi Heller 1912

Species Crocidura ramona Ivanitskaya, Shenbrot, and Nevo 1996

Species Crocidura rapax G. Allen 1923

Subspecies Crocidura rapax subsp. rapax G. Allen 1923

Subspecies Crocidura rapax subsp. kurodai Jameson and Jones 1977

Subspecies Crocidura rapax subsp. lutaoensis Fang and Lee 2002

Subspecies Crocidura rapax subsp. tadae Tokuda and Kano 1936

Species Crocidura religiosa (I. Geoffroy 1827)

Species Crocidura rhoditis Miller and Hollister 1921

Species Crocidura roosevelti Heller 1910

Species Crocidura russula (Hermann 1780)

Subspecies Crocidura russula subsp. russula Hermann 1780

Subspecies Crocidura russula subsp. cintrae Miller 1907

Subspecies Crocidura russula subsp. osorio Molina and Hutterer 1989

Subspecies Crocidura russula subsp. peta Montagu and Pickford 1923

Subspecies Crocidura russula subsp. pulchra Cabrera 1907

Subspecies Crocidura russula subsp. yebalensis Cabrera 1913

Species Crocidura selina Dollman 1915

Species Crocidura serezkyensis Laptev 1929

Species Crocidura shantungensis Miller 1901

Subspecies Crocidura shantungensis subsp. shantungensis Miller 1901

Subspecies Crocidura shantungensis subsp. quelpartis Kuroda 1934

Species Crocidura sibirica Dukelsky 1930

Species Crocidura sicula Miller 1900

Subspecies Crocidura sicula subsp. sicula Miller 1900

Subspecies Crocidura sicula subsp. aegatensis Hutterer 1991

Subspecies Crocidura sicula subsp. calypso Hutterer 1991

Subspecies Crocidura sicula subsp. esuae Kotsakis 1984

Species Crocidura silacea Thomas 1895

Species Crocidura smithii Thomas 1895

Subspecies Crocidura smithii subsp. smithii Thomas 1895

Subspecies Crocidura smithii subsp. debalsaci Hutterer 1981

Species Crocidura somalica Thomas 1895

Species Crocidura stenocephala Heim de Balsac 1979

Species Crocidura suaveolens ( Pallas 1811)

Species Crocidura susiana Redding and Lay 1978

Species Crocidura tanakae Kuroda 1938

Species Crocidura tansaniana Hutterer 1986

Species Crocidura tarella Dollman 1915

Species Crocidura tarfayensis Vesmanis and Vesmanis 1980

Species Crocidura telfordi Hutterer 1986

Species Crocidura tenuis Müller 1840

Species Crocidura thalia Dippenaar 1980

Species Crocidura theresae Heim de Balsac 1968

Species Crocidura thomensis Bocage 1887

Species Crocidura trichura Dobson in Thomas 1888

Species Crocidura turba Dollman 1910

Species Crocidura ultima Dollman 1915

Species Crocidura usambarae Dippenaar 1980

Species Crocidura viaria I. Geoffroy 1834

Species Crocidura virgata Sanderson 1940

Species Crocidura voi Osgood 1910

Species Crocidura vorax G. Allen 1923

Species Crocidura vosmaeri Jentink 1888

Species Crocidura watasei Kuroda 1924

Species Crocidura whitakeri de Winton 1897

Species Crocidura wimmeri Heim de Balsac and Aellen 1958

Species Crocidura wuchihensis Wang 1966

Species Crocidura xantippe Osgood 1910

Species Crocidura yankariensis Hutterer and Jenkins 1980

Species Crocidura zaphiri Dollman 1915

Species Crocidura zarudnyi Ognev 1928

Species Crocidura zimmeri Osgood 1936

Species Crocidura zimmermanni Wettstein 1953

Discussion: Eurasian species revised by Jenkins (1976), Indomalayan and Philippine species by Heaney and Ruedi (1994) and Ruedi (1995), Chinese species by Jiang and Hoffmann (2001). Phenetic and Phylogenetic relationships of African and Palearctic species studied by Butler et al. (1989), Maddalena (1990) and McLellan (1994), and of Asian and Indomalayan species by Heaney and Ruedi (1994), Ruedi (1996), and Ruedi et al. (1998). Karyotypes of SE Asian species described by Ruedi and Vogel (1995), those of Mediterranean isls shrews by Vogel et al. (1990), and of African shrews by Schlitter et al. (1999). No formal subgenera are recognized here, although the type species of proposed subgenera are indicated. For synonyms see Allen (1939), Heim de Balsac and Meester (1977) and Hutterer (1986a).