Enicospilus unctus Shimizu, 2020

Shimizu, So, Broad, Gavin R. & Maeto, Kaoru, 2020, Integrative taxonomy and analysis of species richness patterns of nocturnal Darwin wasps of the genus Enicospilus Stephens (Hymenoptera, Ichneumonidae, Ophioninae) in Japan, ZooKeys 990, pp. 1-144 : 1

publication ID

https://dx.doi.org/10.3897/zookeys.990.55542

publication LSID

lsid:zoobank.org:pub:7B73642C-278D-40F8-9091-B26213C9A704

persistent identifier

https://treatment.plazi.org/id/71C94F28-C439-4F37-86F8-DDEE53B9BD67

taxon LSID

lsid:zoobank.org:act:71C94F28-C439-4F37-86F8-DDEE53B9BD67

treatment provided by

ZooKeys by Pensoft

scientific name

Enicospilus unctus Shimizu
status

sp. nov.

Enicospilus unctus Shimizu sp. nov. Figure 50 View Figure 50

Etymology.

The specific name is derived from the Latin unctus , meaning polished, referring to the posterior part of the propodeum.

Type series.

The holotype female only.

HT: ♀, Matsuyama, Ehime, Shikoku, JAPAN, 26.X.1954, S. Ueda leg. (EUM).

Distribution.

Eastern Palaearctic region.

JAPAN: [Shikoku] Ehime.

Bionomics.

Unknown.

Differential diagnosis.

Some characters (e.g., wide face, long and slender mandible, shape of sclerites) suggest that E. unctus sp. nov. is related to E. shikokuensis . However, E. unctus sp. nov. can easily be distinguished from all other Japanese Enicospilus by the uniformly punctate and shiny posterior area of the propodeum.

Description.

Female (HT). Body length ca. 21.0 mm.

Head with GOI = 2.0 (Fig. 50D View Figure 50 ). Lower face 0.9 × as wide as high, strongly shiny, moderately punctate with setae (Fig. 50B, D View Figure 50 ). Clypeus 2.1 × as wide as high, moderately punctate with setae, convex in profile, and ventral margin impressed (Fig. 50B, D View Figure 50 ). Malar space 0.4 × as long as basal mandibular width (Fig. 50B, D View Figure 50 ). Mandible moderately twisted by 20°, very long, proximally narrowed, distally parallel sided, outer surface with diagonal setose groove (Fig. 50B, D View Figure 50 ). Upper tooth of mandible 2.1 × as long as lower. Frons, vertex and gena strongly shiny with fine setae (Fig. 50B-D View Figure 50 ). Posterior ocellus distinctly separated from eye (Fig. 50B-D View Figure 50 ). Ventral end of occipital carina joining oral carina. Antennae with 54 flagellomeres; first flagellomere 1.5 × as long as second; 20th flagellomere 1.9 × as long as wide.

Mesosoma entirely strongly shiny with fine setae (Fig. 50E View Figure 50 ). Pronotum diagonally finely punctostrigose (Fig. 50E View Figure 50 ). Mesoscutum 1.3 × as long as maximum width, finely punctate to smooth and evenly rounded in profile (Fig. 50E View Figure 50 ). Notauli absent (Fig. 50E View Figure 50 ). Scutellum moderately convex, finely punctate to smooth, with lateral longitudinal carinae along entire length of scutellum. Epicnemium moderately punctate with setae. Epicnemial carina weak, evenly curved to anterior, dorsal end not reaching anterior margin of mesopleuron (Fig. 50E View Figure 50 ). Meso- and metapleuron evenly moderately punctate (Fig. 50E View Figure 50 ). Submetapleural carina broadened anteriorly (Fig. 50E View Figure 50 ). Propodeum evenly rounded in profile; anterior transverse carina complete centrally and not joining pleural carina laterally; anterior area longitudinally striate; spiracular area smooth; posterior area entirely moderately punctate; propodeal spiracle elliptical and not joining pleural carina by ridge (Fig. 50E View Figure 50 ).

Wings (Fig. 50F View Figure 50 ). Fore wing length ca. 14.5 mm with AI = 0.6, CI = 0.3, DI = 0.4, ICI = 0.4, SDI = 1.2, SI = 0.1, SRI = 0.4; vein 1m-cu&M slightly sinuous; vein 2r&RS slightly sinuous; vein RS evenly curved; fenestra and sclerites of discosubmarginal cell as in Fig. 50F View Figure 50 ; proximal sclerite rounded triangular, not confluent with distal sclerite, strongly pigmented; central sclerite suboval, pigmented, positioned in anterodistal part of fenestra; distal sclerite vestigial; proximal corner of marginal cell evenly setose; posterodistal corner of second discal cell ca. 80° and of subbasal cell ca. 80°; vein 1cu-a antefurcal to M&RS by less than 0.1 × length of 1cu-a. Hind wing with NI = 1.0; vein RS almost straight; vein RA with 6 rather stout uniform hamuli.

Legs. Hind leg with coxa in profile 1.9 × as long as deep; basitarsus 2.1 × as long as second tarsomere; fourth tarsomere 3.7 × as long as wide; tarsal claw simply pectinate.

Metasoma with DMI = 1.2, PI = 3.1, THI = 1.7; thyridium oval; ovipositor sheath not longer than posterior depth of metasoma (Fig. 50A View Figure 50 ).

Colour (Fig. 50 View Figure 50 ). Entirely red-brown except for head yellow-brown and apex of mandible black. Wings hyaline. Fore wing sclerites pigmented and amber. Wing veins red-brown to amber.

Male. Unknown.