Angustopila apiostoma Pall-Gergely & Vermeulen, 2023
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publication ID |
https://dx.doi.org/10.3897/zookeys.1147.93824 |
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publication LSID |
lsid:zoobank.org:pub:9BB9881B-0076-473D-8E53-155D37CA1F50 |
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persistent identifier |
https://treatment.plazi.org/id/7369BED7-8447-443D-8FC7-627FEB3B1CBB |
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taxon LSID |
lsid:zoobank.org:act:7369BED7-8447-443D-8FC7-627FEB3B1CBB |
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treatment provided by |
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scientific name |
Angustopila apiostoma Pall-Gergely & Vermeulen |
| status |
sp. nov. |
Angustopila apiostoma Pall-Gergely & Vermeulen sp. nov.
Figs 9 View Figure 9 , 10 View Figure 10
Type material.
Holotype: Vietnam • 1 empty shell (H: 0.84 mm, D. 0.71 mm); Quang Ninh Province, Halong Bay area, Thay Cave on Congfo Island, collected inside cave (locality code: WMVT.0327); 20°52.07'N, 107°12.06'E; 6 Sep. 2003; W.J.M. Maassen leg.; RMNH 5006718.
Paratypes: Vietnam • 2 figured shells; same data as for holotype; RMNH 5006722 • 5 shells; same data as for holotype; HNHM 100441 • 5 shells; same data as for holotype; NMBE 550640 • 28 shells; same data as for holotype; coll. PGB (ex coll. W. Maassen) • 987 shells; same data as for holotype; RMNH 347758 • 10 shells; same data as for holotype; coll. HA • 53 shells; Quang Ninh Province, Halong Bay area, unnamed island in Dau Moi Temple area (locality code: WMVT.0328); 20°55.69'N, 107°09.40'E; 13 Sep. 2003; W.J.M. Maassen leg.; RMNH 347759 • 41 shells (one of them is figured: Fig. 99D View Figure 99 ); Haiphong Province, Cat Ba Island, Cave Qua Vang, inside cave, large, ecologically intact active cave with speleothems; 20°48.64'N, 107°04.64'E; 60 m a.s.l.; 6 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16614 • 1 shell; Haiphong Province, Cat Ba Island, Cave Qua Vang, around cave entrance, rocky limestone slope with low, rather mature forest; 6 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16604 • 397 shells; Haiphong Province, Cat Ba Island, large, somewhat ecologically-disturbed, active cave with speleothems; 20°50.06'N, 106°55.91'E; 0-50 m a.s.l.; 7 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16605 • 343 shells; Haiphong Province, Cat Ba Island, Cave Hoa Cuong, inside cave, polluted cave disturbed by tourism, with concrete paths; 20°50.41'N, 106°59.15'E; 50 m a.s.l.; 5 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16609 • 13 shells; Haiphong Province, Cat Ba Island, Cave Xa Bac, inside cave; 20°50.07'N, 106°58.61'E; 30 m a.s.l.; 9 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16615 • 29 shells; Quang Ninh Province, Halong Bay area, Dao Bo Hon, Song Sot Cave, drift material washed over sinkhole in cave; 20°50.83'N, 107°05.67'E; 2 Oct. 1998; J.J. Vermeulen & A.J. Whitten leg.; JJV 6268 • 248 shells; Quang Ninh Province, Halong Bay area, Dao Bo Hon, Song Sot Cave, guano-enriched sediments in cave; 20°50.83'N, 107°05.67'E; 2 Oct. 1998; J.J. Vermeulen & A.J. Whitten leg.; JJV 6269 • 4 shells; Haiphong Province, Cat Ba Island, Cave Xa Bac, around cave entrance; 20°50.07'N, 106°58.61'E; 30 m a.s.l.; 9 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16616 • 6 shells; Haiphong Province, Cat Ba Island, cave Trung Trang, inside cave, cave disturbed by tourism and concrete paths; 20°47.30'N, 106°59.84'E; 50 m a.s.l.; 5 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16602.
Diagnosis.
A small to medium-sized, toothless Angustopila species with a very narrow umbilicus, conspicuously protruding aperture, and a characteristically shaped aperture (kidney-shaped with narrow sinulus).
Description.
Shell small to medium-sized for the genus, shell height variable, but always higher than wide; pale grey, conical, last whorl widest from standard apertural view; protoconch consists of 1.5 whorls, microstructure finely pitted and granular with a powdery superficial texture, spiral striation not discernible; teleoconch finely ornamented with irregularly spaced radial growth lines crossed by fine rows of equidistantly-spaced microscopic spiral threads (ca. 18-24 on last whorl in apertural view); on frontal and ventral surface of body whorl spiral and radial lines prominent; whorls 4.25-5, shouldered; aperture slightly oblique to shell axis from lateral view; umbilicus deep, very narrow; ventral side characteristic due to comparatively long and straight tuba; aperture kidney-shaped with narrow sinulus; peristome slightly expanded, not reflected; parietal callus slightly protruding, aperture detached from penultimate whorl; aperture toothless or with a vestigial, deeply-situated parietal tooth.
Measurements (in mm).
H = 0.73-0.93, D = 0.67-0.85, H/D*100 = 101.2-128.2 (n = 14), RUD = 19.2-25.7 (n = 6).
Differential diagnosis.
Angustopila apiostoma sp. nov. can be distinguished from A. elevata , by the very narrow umbilicus, the protruding aperture (visible in ventral view) and the kidney-shaped aperture. See also under A. fratermajor sp. nov. and remarks.
Etymology.
The name Angustopila apiostoma derives from the Greek ἄπιον (= a pear) and στόμα (= mouth) referring to the pear-shaped aperture (to be used as a noun in apposition).
Distribution.
This species is known from two nearby localities in the Halong Bay Area, Vietnam (Fig. 11 View Figure 11 ).
Remarks.
The ventral side and the shape of the aperture of A. apiostoma sp. nov. resembles those of Dentisphaera maxema Páll-Gergely & Jochum, 2017, which has a more protruding aperture (i.e., a longer tuba) and six apertural teeth. However, the two prominent teeth on the parietal side of Dentisphaera distinguishes this genus from Angustopila .
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