Melaenini

Ball, George E. & Shpeley, Danny, 2005, Taxonomic review of the Tribe Melaenini (Coleoptera: Carabidae), with observations on morphological, ecological and chorological evolution, Zootaxa 1099 (1), pp. 1-120 : 1-120

publication ID

https://doi.org/ 10.11646/zootaxa.1099.1.1

publication LSID

lsid:zoobank.org:pub:CA0A0ED3-D49D-4198-A409-407EF86A164E

persistent identifier

https://treatment.plazi.org/id/227887BE-FFFF-FFAF-250D-F7CCFA88FBC4

treatment provided by

Felipe

scientific name

Melaenini
status

 

Key to the genera, subgenera and species of Melaenini

1. Head dorsally with single supraorbital seta over each eye. Dorsal surface of body with fixed setae only, without general setose vestiture. Antennomeres 1–4 with setae near distal margin of each article. Elytron with striae distinctly impressed, intervals convex ................................. Melaenus Dejean View in CoL ................ 2

– Head dorsally with two supraorbital setae over each eye. Dorsal surface of body with general setose vestiture. Antennomeres 1–4 with setae over surface of each article. Elytron with striae shallow (stria 1), or indistinct; intervals flat............... ......................................................................... Cymbionotum Baudi di Selve View in CoL ... 3

2 (1). Pronotum each side with two posteriolateral projections. Specimen from locality in Africa ( Fig. 10 View FIGURE 10 )......................................................... M. elegans Dejean View in CoL , p. 39

– Pronotum each side with one posteriolateral projection. Specimen from locality in India ( Fig. 10 View FIGURE 10 ) ........................................................... M. piger Fabricius View in CoL , p. 38

3 (1) Antennomere 2 ( Fig. 11E, A View FIGURE 11 2 View FIGURE 2 ) globose, spherical in outline, not pyriform; antennomeres 5–10 ( Fig. 11G, A View FIGURE 11 6 View FIGURE 6 , A 7 View FIGURE 7 ) markedly asymmetric ............................ ................................................................................ semelederi View in CoL species group ... 4

– Antennomere 2 ( Fig. 11D, A View FIGURE 11 2 View FIGURE 2 ) pyriform, margins markedly constricted proximad base; antennomeres 5–10 ( Fig. 11F, A View FIGURE 11 6 View FIGURE 6 , A 7 View FIGURE 7 ) moderately asymmetric........ 6

4 (3). Elytral vestiture denser ( Figs. 2C View FIGURE 2 , 3B View FIGURE 3 ) ................. C. semelederi Chaudoir View in CoL , p. 68

– Elytral vestiture sparser ( Fig. 2D View FIGURE 2 , 3A View FIGURE 3 ) .............................................................. 5

5 (4). Phallus in left lateral aspect markedly curved preapically and endophallus with two microtrichial fields (Fig. 17J). Specimen from locality in or near Afghanistan (Fig. 18) ................................................................. C. striatum Reitter View in CoL , p. 73

– Phallus in left lateral aspect gradually curved preapically and endophallus with single microtrichial field (Fig. 17K). Specimen from locality in or near Iran (Fig. 18) ........................................................................ C. mandli Jedlička View in CoL , p. 74

6 (3). Dorsal surface uniformly dark rufous. Pronotum ( Fig. 14A View FIGURE 14 ) with posteriolateral angles distinctly anteriad posterior margin, proepipleuron not visible. Specimen from locality in Western Hemisphere (northern South America— Fig. 16 View FIGURE 16 )........... ............................................................... Procoscinia, new subgenus............... 7

Dorsal surface various in color: uniformly dark concolorous to uniformly pale concolorous, to bicolored (Figs. 17A–C, 17F­H, 19A­B, 19E­F, 19I­J, 19L­N, 22A­C). Pronotum ( Fig. 14B View FIGURE 14 ) with posteriolateral angles near posterior margin, proepipleuron visible. Specimen from locality in Eastern Hemisphere (Afrotropical, southern Palaearctic , Oriental Regions­ Figs. 16 View FIGURE 16 , 20, 21, 23) .................... ...................................................... Subgenus Cymbionotum (sensu stricto) View in CoL ...... 8

7 (6). Dorsal surface with shallow microlines, mesh pattern isodiametric. Specimen from locality in northern Colombia........................... C. fernandezi View in CoL , n. sp., p. 56

– Dorsal surface without microlines, smooth. Specimen from locality in northern Venezuela....................................................................... C. negrei Perrault View in CoL , p. 58

8 (6). Body size large (SBL more than 5.0 mm, EW more than 1.5 mm). Elytron bicolored (Figs. 17A–C), pale anteriorly, dark posteriorly. Femora and tibiae piceous to rufous ...................................................................... basale View in CoL species group......9

– Body size smaller (SBL less than 4.5 mm, EW less than 1.5 mm). Elytron concolorous or bicolored. Femora and tibiae rufous to testaceous ......................... 10

9 (8). Dorsal vestiture ( Fig. 2D View FIGURE 2 ) of elytron relatively long setae. Specimen from locality in Palaearctic southwestern Asia or in the Afrotropical Region west to Niger ( Fig. 16 View FIGURE 16 ) .............................................................. C. semirubricum Reitter View in CoL , p. 61

– Dorsal vestiture of elytron relatively shorter setae. Specimen from locality in northwestern Afrotropical Region ( Mali and Sénégal, Fig. 16 View FIGURE 16 )............................ ......................................................................................... C. basale Dejean View in CoL , p. 65

10 (8). Elytron with punctation in straight lines, stria­like. Dorsal integument uniformly rufotestaceous. Clypeus (cf. Fig. 11B View FIGURE 11 ) without dentiform projection .................. ...................................................................................... C. namwala View in CoL , n. sp., p. 99

– Elytron with punctation scattered over surface (if lines of punctation apparent, punctures also in between). Dorsal integument various in color. Clypeus with ( Fig. 11C View FIGURE 11 ) or without ( Fig. 11B View FIGURE 11 ) dentiform projection ..................................... 11

11 (10). Eyes somewhat flattened, only slightly projected laterad dorsolateral carina of head ( Fig. 11B View FIGURE 11 ). Values for VW/HW 0.87 or more. Dorsal integument concolorous, uniformly pale .......................................................................................... 12

– Eyes rounded, moderately projected laterad dorsolateral carina of head ( Fig. 11C View FIGURE 11 ). Values for VW/HW 0.87 or less. Dorsal integument various: distinctly or indistinctly bicolored, or uniformly pale........................................................... 15

12 (11). Clypeus anteriomedially with dentiform projection ( Fig. 11C View FIGURE 11 , ct) ................. 13

– Clypeus anteriomedially without dentiform projection ( Fig. 11B View FIGURE 11 ) ................. 14

13 (12). Scutellum with microsculpture mesh pattern isodiametric, sculpticells convex .. ............................................................................. C. candidum Andrewes View in CoL , p. 100

– Scutellum smooth, without microlines .................. C. subcaecum View in CoL , n. sp., p. 101

14 (12). Scutellum with slightly transverse mesh pattern over much of surface, smooth apically. Specimen from locality in southwestern Asia, west to eastern Palaearctic Africa (Fig. 23) ........................................ C. transcaspicum Semenov View in CoL p. 102

– Scutellum with slightly transverse mesh pattern near anterior margin, smooth apically. Specimen from locality in Afrotropical Region, east to Anatolian peninsula in Palaearctic Region (Fig. 23) ...... C. microphthalmum Chaudoir View in CoL , p. 104

15 (11). Specimen from locality in South Africa or Botswana (Fig. 20). Male genitalia as in Fig. 19D.............................................................. C. capicola Péringuey View in CoL , p. 83

– Specimen from locality farther north in Africa, and eastward in Palaearctic southwestern Asia, to Vietnam, in the Oriental Region. Male genitalia otherwise (Figs. 19C, 19G–H, 19K, 19O and 22D)........................................................... 16

16 (15). Head with postocular transverse impression with microsculpture mesh pattern ( Figs. 11B–C View FIGURE 11 ) isodiametric to slightly transverse ............................................ 17

– Head with postocular transverse impression smooth, without microlines, or microlines not joined in form of a network...................................................... 20

17 (16). Specimen from locality in Palaearctic southwestern Asia, or in eastern Afrotropical Region, north of South Africa (Fig. 21); elytron bicolored (Figs. 19L–N; cf. Fig. 19F) ................................................................... C. pictulum Bates View in CoL , p. 93

– Specimen from locality in Oriental Region (Fig. 21) ...................................... 18

18 (17). Elytron concolorous, rufous, or bicolored, with piceous or black mark small, confined to disc (Figs. 19E–F). Specimen from locality in Burma, or farther east (Fig. 21) .................... .................................................. C. helferi Chaudoir View in CoL , p. 88

– Elytron bicolored, dark marks more extensive, extended to lateral margin. Specimen from locality in India, Sri Lanka, or Nepal............................................ 19

19 (18). Phallus in left lateral aspect gradually curved preapically (Fig. 19K). Specimen from locality in southern India (Calcutta area southward) or Sri Lanka.............. ................................................................................... C. fluviale Andrewes View in CoL , p. 92

– Phallus in left lateral aspect markedly curved preapically (Fig. 19H). Specimen from locality in northern India or Nepal................... C. fasciger Chaudoir View in CoL , p. 91

20 (16). Elytron concolorous, pale ................................................................................ 21

– Elytron bicolored (Figs. 19A–B, 22A–C) ........................................................ 24

21 (20). Scutellum entirely smooth or surface wrinkled basally, without microlines ........ ........................................................................ C. rufotestaceum Fairmaire View in CoL , p. 81

– Scutellum basally with microlines in form of mesh pattern ............................ 22

22 (21). Dissected specimen female (one of two species) ................................................. ..................... C. schueppelii Dejean View in CoL (in part), p. 78, C. fasciatum View in CoL (in part), p. 95

– Dissected specimen male ................................................................................. 23

23 (22). Male genitalia (Fig. 19C) with microtrichial field (mf) of endophallus extensive; phallus with preapical portion shorter, curved ventrad rather sharply ................. .................................................................... C. schueppelii Dejean View in CoL (in part), p. 78

– Male genitalia (Fig. 22D) with microtrichial field (mf) of endophallus very small; phallus with preapical portion longer, curved ventrad more gradually ..... ...................................................................... C. fasciatum Dejean View in CoL (in part), p. 98

24 (20). Elytron bicolored, with dark marking (Figs. 19A–B) extended to entire posterior margin ...................................................................... C. schueppelii Dejean View in CoL , p. 78

– Elytron bicolored, fasciate (Figs. 22A–C), fascia narrow or broad, but complete posterior margin not dark, at most bicolored............... C. fasciatum Dejean View in CoL , p. 95

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Carabidae

Darwin Core Archive (for parent article) View in SIBiLS Plain XML RDF