Speocera babau, Brescovit & Zampaulo & Cizauskas & Pedroso, 2022

Brescovit, Antonio Domingos, Zampaulo, Robson De Almeida, Cizauskas, Igor & Pedroso, Leandro, 2022, Three new subterranean species of the genus Speocera (Araneae, Ochyroceratidae) from caves of the Carajás area in the state of Pará, Brazil, Zootaxa 5169 (2), pp. 101-135 : 109-120

publication ID	https://doi.org/ 10.11646/zootaxa.5169.2.1
publication LSID	lsid:zoobank.org:pub:F4E7B475-B12C-4281-932C-212AE51B62AC
DOI	https://doi.org/10.5281/zenodo.6929150
persistent identifier	https://treatment.plazi.org/id/213A87E1-FFD4-FFEC-26C4-FF09632AE843
treatment provided by	Plazi
scientific name	Speocera babau
status	sp. nov.

Treatment

Speocera babau sp. nov.

Figures 7–11 View FIGURE 7 View FIGURE 8 View FIGURE 9 View FIGURE 10 View FIGURE 11 , 19B, D View FIGURE 19 , 20 A–D View FIGURE 20

Type material. Male holotype and female paratype from Cave N 1_0024 ( GEM –1220) (6°01’53”S 50°16’20”W), Serra dos Carajás, Parauapebas, Pará, Brazil, 11/VI – 02/VII/2014, Equipe Carste et al. col., deposited in IBSP 207354; GoogleMaps Paratypes: male and female from Cave N 1_0056 (GEM-1254) (6°01’11”S 50°16’44”W), Serra dos Carajás, Parauapebas, Pará, Brazil, 07 – 28/I/2015, Equipe Carste et al. col. (ex IBSP 207537 View Materials ), deposited in MPEG 37251; GoogleMaps two males and two females from Cave N 1_0226 (6°02’16”S 50°16’03”W), Serra dos Carajás, Parauapebas , Pará , Brazil, 04/IX – 06/X/2014, Equipe Carste et al. col. (ex IBSP 207471 View Materials ), deposited in ISLA GoogleMaps .

Material examined. Brazil. Pará: Parauapebas, Serra dos Carajás, Cave N1_0008 (GEM-1204) (6°02’20”S 50°16’13”W), 11/VI – 02/VII/2014, 1♀ ( IBSP 207343); Cave N1_0015 (GEM-1211) (6°02’02”S 50°16’16”W), 2♂ ( IBSP 2073440; 1♀ ( IBSP 207345); 24/II – 13/III/2015, 1♀ ( IBSP 208795); 1♀ ( IBSP 208844); Cave N1_0018 (GEM-1214) (6°02’01”S 50°16’17”W), 3♀ ( IBSP 207346-207347); 07 – 28/I/2015, 2♀ ( IBSP 207513); 07 – 28/ I/2015, 2♀ ( IBSP 207514); Cave N1_0019 (GEM-1215) (6°02’01”S 50°16’17”W), 1♂ ( IBSP 207515); Cave N1_ 0020 (GEM-1216) (6°01’58”S 50°16’18”W), 3♂ 9♀ ( IBSP 207516-207521); 16/VII-06/VIII/2014, 1♂ 4♀ ( IBSP 208469-208473); Cave N1_0021 (GEM-1217) (6°01’57”S 50°16’19”W), 2♀ ( IBSP 207522-207523); Cave N1_ 0022 (GEM-1218) (6°01’57”S 50°16’19”W), 07-28/I/2015, 1♂ 1♀ ( IBSP 207524); Cave N1_0024 (GEM-1220) (6°01’53”S 50°16’20”W), 11/VI-02/VII/2014, 3♂ 15♀ ( IBSP 207349-207358; IBSP 207563-IBSP 207568; IBSP 207570; 2♀ ( IBSP 207569; SEM); 1♀ ( IBSP 207571; SEM); Cave N1_0038 (GEM-1234) (6°01’49”S 50°16’17”W), 04/IX-06/X/2014, 1♀ ( IBSP 207419); 02-29/IV/2015, 7♀ ( IBSP 207572-207575); Cave N1_0039 (GEM-1235) (6°01’46”S 50°16’13”W), 1♀ ( IBSP 207576); Cave N1_0054 (GEM-1251) (6°01’14”S 50°16’38”W), 11/VI-02/ VII/2014, 2♀ ( IBSP 207359); 07-28/I/2015, 1♂ 1♀ ( IBSP 207525); Cave N1_0055 (GEM-1253) (6°01’12”S 50°16’43”W), 2♂ 4♀ ( IBSP 207526-207530); 16/VII-06/VIII/2014, 3♀ ( IBSP 208474); Cave N1_0056 (GEM-1254) (6°01’11”S 50°16’44”W), 07-28/I/2015, 8♂ 16♀ ( IBSP 207531-207538); Cave N1_0057 (GEM-1255) (6°01’10”S 50°16’44”W), 02-29/IV/2015, 1♂ ( IBSP 207577); Cave N1_0060 (GEM-1258) (6°01’12”S 50°16’41”W), 11/VI-02/VII/2014, 4♂ 32♀ ( IBSP 207360-207368; IBSP 207539-207545; IBSP 207560); Cave N1_0070 (GEM-1268) (6°01’14”S 50°17’18”W), 1♀ ( IBSP 207369); 24/II-13/III/2015, 1♀ ( IBSP 208802); Cave N1_0071 (GEM-1269) (6°01’14”S 50°17’17”W), 1♀ ( IBSP 208803); Cave N1_0072 (GEM-1270) (6°01’13”S 50°17’17”W), 11/VI-02/VII/2014, 1♂ 1♀ ( IBSP 207370); Cave N1_0073 (GEM-1271) (6°01’13”S 50°17’17”W), 02-29/IV/2015, 17♂ 47♀ ( IBSP 207584-207603; IBSP 208475- 208486; Cave N1_0085 (GEM-1283) (6°01’07”S 50°17’01”W), 11/VI-02/VII/2014, 1♀ ( IBSP 207371); Cave N1_0096 (GEM-1294) (6°01’09”S 50°16’59”W), 2♂ 4♀ ( IBSP 207372-207374); Cave N1_0098 (GEM-1296) (6°01’08”S 50°17’05”W), 24/II-13/III/2015, 1♀ ( IBSP 208804); Cave N1_0124 (GEM-1322) (6°00’14”S 50°17’15”W), 11/VI-02/VII/2014, 1♀ ( IBSP 207375); Cave N1_0124 (GEM-1322) (6°00’14”S 50°17’15”W), 07-28/I/2015, 2♀ ( IBSP 207547-207548); Cave N1_0125 (GEM-1323) (6°00’15”S 50°17’15”W), 11/VI-02/VII/2014, 1♀ ( IBSP 207376); Cave N1_0128 (GEM-1326) (6°00’28”S 50°17’29”W), 1♂ ( IBSP 207377); 02-29/IV/2015, 1♀ ( IBSP 207609); Cave N1_0129 (GEM-1327) (6°00’36”S 50°17’28”W), 2♀ ( IBSP 207610; Cave N1_0130 (GEM-1328) (6°00’35”S 50°17’29”W), 1♂ 3♀ ( IBSP 207378- 207379); Cave N1_0130 (GEM-1328) (6°00’35”S 50°17’29”W), 02-29/IV/2015, 1♂ 3♀ ( IBSP 207611-207612); Cave N1_0147 (GEM-1347) (6°02’32”S 50°16’26”W), 2♀ ( IBSP 207613); Cave N1_0152 (GEM-1352) (6°02’45”S 50°16’14”W), 1♀ ( IBSP 207614); Cave N1_0153 (GEM-1353) (6°02’43”S 50°16’19”W), 1♂ ( IBSP 207615); Cave N1_0155 (GEM-1355) (6°02’41”S 50°16’22”W), 11/VI-02/VII/2014, 1♀ ( IBSP 207380); Cave N1_0156 (GEM-1356) (6°02’41”S 50°16’22”W), 1♂ ( IBSP 207381); 03-17/XII/2014, 1♂ ( IBSP 207488); Cave N1_0158 (GEM-1358) (6°02’38”S 50°16’23”W), 11/VI-02/VII/2014, 1♂ 1♀ ( IBSP 207382), 03-17/XII/2014, 1♂ 3♀ ( IBSP 207489-207490); 2♂ 4♀ ( IBSP 207491; SEM); Cave N1_0073 (GEM-1271) (6°01’13”S 50°17’17”W), 16/VII-06/VIII/2014, 6♀ ( IBSP 207390-207392); Cave N1_0160 (GEM-1360) (6°02’34”S 50°16’24”W), 02-29/IV/2015, 3♀ ( IBSP 207616-207617); Cave N1_0168 (GEM-1368) (6°01’16”S 50°18’05”W), 16/VII-06/VIII/2014,4 ♂ 3♀ ( IBSP 207393; IBSP 207492- 207495); 16/VII-06/VIII/2014, 2♂ 1♀ ( IBSP 208487); Cave N1_0170 (GEM-1370) (6°01’23”S 50°17’58”W), 3♀ ( IBSP 207394); 3♀ ( IBSP 207496); 1♂ 2♀ ( IBSP 208488-208489); Cave N1_0111 (GEM-1309) (6°00’13”S 50°17’28”W), 1♀ ( IBSP 207546); Cave N1_0171 (GEM-1371) (6°01’18”S 50°18’04”W), 7♂ 11♀ ( IBSP 207395; IBSP 207497-207500; IBSP 208490- IBSP 208492); Cave N1_0173 (GEM-1373) (6°01’27”S 50°17’55”W), 3♀ ( IBSP 207396-207397); 07-28/I/2015, 1♂ 3♀ ( IBSP 207549); 207551); Cave N1_0174 (GEM-1374) (6°01’28”S 50°17’53”W), 4♂ 13♀ ( IBSP 207398; IBSP 207552-207556; IBSP 208493-208496); Cave N1_ 0175 (GEM-1375) (6°01’28”S 50°17’53”W), 1♀ ( IBSP 207399); 07-28/I/2015, 1♀ ( IBSP 207557); Cave N1_0176 (GEM-1376) (6°01’38”S 50°17’51”W), 1♀ ( IBSP 207558); Cave N1_0187 (GEM-1388) (6°02’38”S 50°16’33”W), 1♀ ( IBSP 208497); Cave N1_0010 (GEM-1206) (6°01’10”S 50°16’42”W), 02-29/IV/2015, 1♀ ( IBSP 207561); 2♀ ( IBSP 208468); Cave N1_0016 (GEM-1212) (6°01’10”S 50°16’41”W), 04/IX-06/X/2014, 1♂ 1♀ ( IBSP 207410); 1♀ ( IBSP 207411); 02-29/IV/2015, 3♀ ( IBSP 207562); Cave N1_0038 (GEM-1234) (6°01’49”S 50°16’17”W), 04/IX-06/X/2014, 1♀ ( IBSP 207412); 2♀ ( IBSP 207413); 2♀ ( IBSP 207414); 2♂ ( IBSP 207415); 1♀ ( IBSP 207416); 2♀ ( IBSP 207418); 1♀ ( IBSP 207420); Cave N1_0041 (GEM-1237) (6°01’44”S 50°16’09”W), 1♀ ( IBSP 207421); Cave N1_0044 (GEM-1240) (6°01’13”S 50°16’40”W), 11♂ 14♀ ( IBSP 207422-207427; IBSP 208796- 208800; IBSP 208845); Cave N1_0045 (GEM-1241) (6°01’43”S 50°16’31”W), 2♂ 1♀ ( IBSP 207428); Cave N1_ 0045 (GEM-1241) (6°01’43”S 50°16’31”W), 3♀ ( IBSP 208846); Cave N1_0046 (GEM-1242) (6°01’43”S 50°16’31”W), 3♀ ( IBSP 208801); 1♀ ( IBSP 208847); Cave N1_0049 (GEM-1245) (6°01’40”S 50°16’32”W), 04/ IX-06/X/2014, 2♀ ( IBSP 207430); Cave N1_0062 (GEM-1260) (6°01’10”S 50°16’44”W), 7♂ 14♀ ( IBSP 207431- 207435; IBSP 207579-207583); Cave N1_0065 (GEM-1263) (6°01’05”S 50°16’48”W), 04/IX-06/X/2014, 2♀ ( IBSP 207436); 02-29/IV/2015, 1♂ 1♀ ( IBSP 207578); Cave N1_0067 (GEM-1265) (6°01’13”S 50°17’20”W), 1♀ ( IBSP 207437); Cave N1_0088 (GEM-1286) (6°01’02”S 50°17’04”W), 1♂ 4♀ ( IBSP 207438-207439); 02-29/ IV/2015, 2♂ 10♀ ( IBSP 207604-207606); Cave N1_0102 (GEM-1300) (6°00’51”S 50°17’27”W), 04/IX-06/ X/2014, 2♂ 3♀ ( IBSP 207440-207442); 1♂ ( IBSP 207443; SEM); 02-29/IV/2015, 1♀ ( IBSP 207607); Cave N1_ 0112 (GEM-1310) (6°00’27”S 50°17’29”W), 1♀ ( IBSP 207608); Cave N1_0121 (GEM-1319) (6°02’03”S 50°17’56”W), 04/IX-06/X/2014, 1♀ ( IBSP 207444); Cave N1_0134 (GEM-1332) (6°01’03”S 50°17’28”W), 1♀ ( IBSP 207445); Cave N1_0137 (GEM-1335) (6°01’32”S 50°16’29”W), 1♀ ( IBSP 207446); Cave N1_0148 (GEM-1348) (6°02’32”S 50°16’25”W), 1♀ ( IBSP 207447); Cave N1_0165 (GEM-1365) (6°01’08”S 50°18’27”W), 2♀ ( IBSP 207448); 1♀ ( IBSP 207449); 1♂ ( IBSP 207450); 1♀ ( IBSP 207451); 02-29/IV/2015, 2♀ ( IBSP 207618- 207619); Cave N1_0177 (GEM-1377) (6°01’38”S 50°17’51”W), 2♀ ( IBSP 207452); 1♀ ( IBSP 207453); 1♀ ( IBSP 207620); Cave N1_0179 (GEM-1380) (6°02’47”S 50°17’25”W), 1♀ ( IBSP 207454); 1♀ ( IBSP 207455); 24/II-13/ III/2015, 1♀ ( IBSP 208805); 4♂ 3♀ ( IBSP 208848); Cave N1_0180 (GEM-1381) (6°02’33”S 50°16’25”W), 02- 29/IV/2015, 1♀ ( IBSP 207621); Cave N1_0181 (GEM-1382) (6°02’33”S 50°16’26”W), 04/IX-06/X/2014, 1♀ ( IBSP 207456); 1♀ ( IBSP 207622); Cave N1_0191 (GEM-1392) (6°02’46”S 50°16’39”W), 1♀ ( IBSP 207623); Cave N1_0193 (GEM-1394) (6°02’53”S 50°17’29”W), 04/IX-06/X/2014, 1♀ ( IBSP 207457); Cave N1_0195 (GEM-1396) (6°02’22”S 50°17’26”W), 24/II-13/III/2015, 1♀ ( IBSP 208849); Cave N1_0202 (GEM-1403) (6°02’40”S 50°16’33”W), 2♀ ( IBSP 208850-208851); Cave N1_0203 (GEM-1404) (6°02’42”S 50°16’42”W), 04/ IX-06/X/2014, 1♀ ( IBSP 207458); Cave N1_0205 (GEM-1406) (6°02’42”S 50°16’36”W), 1♀ ( IBSP 207459); Cave N1_0206 (GEM-1407) (6°03’29”S 50°17’04”W), 1♀ ( IBSP 208852); Cave N1_0207 (GEM-1408) (6°03’27”S 50°17’10”W), 1♀ ( IBSP 207460); Cave N1_0214 (GEM-1416) (6°02’08”S 50°17’47”W), 2♀ ( IBSP 207461); Cave N1_0216 (6°03’20”S 50°17’36”W), 1♀ ( IBSP 207462); 13/III/2015, 1♀ ( IBSP 208853); Cave N1_0219 (6°02’23”S 50°17’32”W), 04/IX-06/X/2014, 2♀ ( IBSP 207463-207464); Cave N1_0222 (6°00’35”S 50°17’28”W), 2♀ ( IBSP 207465); Cave N1_0225 (6°02’17”S 50°16’02”W), 1♀ ( IBSP 207466); Cave N1_0226 (6°02’16”S 50°16’03”W), 2♂ 6♀ ( IBSP 207467-207471); Cave N1_0228 (6°01’10”S 50°17’01”W), 11/VI-02/VII/2014, 1♂ 2♀ ( IBSP 207383); Cave N1_0229 (6°01’21”S 50°18’44”W), 02-29/IV/2015, 1♀ ( IBSP 207624); Cave N1_0231 (6°01’14”S 50°16’21”W), 4♀ ( IBSP 207384-207385; 3♂ 6♀ ( IBSP 207501-207502); Cave N1_0233 (6°02’23”S 50°16’23”W), 04/IX-06/X/2014, 1♀ ( IBSP 207472); 02-29/IV/2015, 1♀ ( IBSP 207625); 1♀ ( IBSP 207626); Cave N1‗0235 (6°01’16”S 50°16’24”W), 1♀ ( IBSP 207627); Cave N1_0236 (6°01’15”S 50°16’25”W), 11/VI-02/ VII/2014, 1♀ ( IBSP 207386); 03-17/XII/2014, 1♀ ( IBSP 207503); Cave N1_0237 (6°01’15”S 50°16’25”W), 1♂ ( IBSP 207504); 03-17/XII/2014, 1♂ 5♀ ( IBSP 207505-207508); Cave N1_0238 (6°01’15”S 50°16’25”W), 1♀ ( IBSP 207509); 1♀ ( IBSP 207511); 11/VI-02/VII/2014, 1♂ 1♀ ( IBSP 207387-207388); 03-17/XII/2014, 1♀ ( IBSP 207510); Cave N1_0240 (6°01’19”S 50°16’26”W), 04/IX-06/X/2014, 2♀ ( IBSP 207473-207474; 6♀ ( IBSP 207475-207478); 02-29/IV/2015, 2♂ 8♀ ( IBSP 207628-207632); Cave N1_0247 (6°01’14”S 50°16’23”W), 11/VI-02/VII/2014, 1♀ ( IBSP 207389); 03-17/XII/2014, 1♂ ( IBSP 207512), all collected by Equipe Carste et al.; Cave N6_0005 (6°07’22”S 50°10’28”W), 3♀ ( IBSP 207633-207634), all collected in 02-29/IV/2015 by Equipe Carste et al. col.; Cave N8_0002 (GEM-1803) (6°10’04”S 50°09’33”W), 16/VII-06/VIII/2014, 1♀ ( IBSP 207401; 2♀ ( IBSP 208500-208501); Cave N8_0003 (GEM-1804) (6°10’05”S 50°09’30”W), 04/IX-06/X/2014, 6♀ ( IBSP 207479- 207481); 02-29/IV/2015, 6♀ ( IBSP 207636-207639); Cave N8_0004 (GEM-1805) (6°10’06”S 50°09’27”W), 24/ II-13/III/2015, 1♂ ( IBSP 208855); Cave N8_0006 (GEM-1807) (6°10’03”S 50°09’36”W), 16/VII-06/VIII/2014, 4♀ ( IBSP 207402); 02-29/IV/2015,1 ♀ ( IBSP 207640); 02-29/IV/2015, 1♀ ( IBSP 207641); 16/VII-06/VIII/2014, 1♂ ( IBSP 208502); Cave N8_0008 (GEM-1809) (6°10’05”S 50°09’32”W), 02-29/IV/2015, 2♀ ( IBSP 207642- 207643); Cave N8_0009 (GEM-1810) (6°10’06”S 50°09’35”W), 3♀ ( IBSP 208503); Cave N8_0011 (GEM-1812) (6°10’07”S 50°09’28”W), 1♀ ( IBSP 207644); Cave N8_0012 (GEM-1813) (6°10’05”S 50°09’28”W), 1♀ ( IBSP 208504); 24/II-13/III/2015, 1♀ ( IBSP 208857); Cave N8_0013 (GEM-1814) (6°10’04”S 50°09’29”W), 02-29/ IV/2015, 1♂ ( IBSP 207645); Cave N8_0017 (GEM-1818) (6°10’06”S 50°09’26”W), 04/IX-06/X/2014, 1♂ 2♀ ( IBSP 207482-208723); Cave N8_0018 (GEM-1819) (6°10’07”S 50°09’26”W), 02-29/IV/2015, 4♀ ( IBSP 207646- 207648; 1♂ 1♀ ( IBSP 208505); Cave N8_0019 (GEM-1820) (6°10’10”S 50°09’25”W), 9♀ ( IBSP 207649-207652); 16/VII-06/VIII/2014, 1♀ ( IBSP 207404); 1♂ 6♀ ( IBSP 208506-208508); Cave N8_0020 (GEM-1821), (6°10’08”S 50°09’25”W), 3♂ 16♀ ( IBSP 208509-208510; IBSP 207405;- IBSP 208858-208860); Cave N8_0022 (GEM-1823) (6°10’05”S 50°09’28”W), 16/VII-06/VIII/2014, 2♀ ( IBSP 207406); 02-29/IV/2015, 2♂ 4♀ ( IBSP 207653-207654); 2♀ ( IBSP 208511); Cave N8_0023 (GEM-1824) (6°10’06”S 50°09’29”W), 3♂ 13♀ ( IBSP 207655-207660); 04/ IX-06/X/2014, 2♂ 6♀ ( IBSP 207483-207487); 02-29/IV/2015, 1♂ ( IBSP 207675); Cave N8_0024 (GEM-1825) (6°10’24”S 50°09’09”W), 16/VII-06/VIII/2014, 1♂ 3♀ ( IBSP 207407); 02-29/IV/2015, 1♂ 4♀ ( IBSP 207661- 207664); 16/VII-06/VIII/2014, 10♀ ( IBSP 208512-208515); Cave N8_0025 (GEM-1826) (6°10’29”S 50°09’04”W), 1♂ 3♀ ( IBSP 208516-208518); 02-29/IV/2015, 1♂ ( IBSP 207665); Cave N8_0026 (GEM-1827) (6°10’15”S 50°09’29”W), 1♀ ( IBSP 207666); 2♀ ( IBSP 207408-207409), Cave N8_0027 (GEM-1828) (6°10’32”S 50°09’28”W), 1♂ ( IBSP 207667); 3♀ ( IBSP 208519-208520); Cave N8_0028 (GEM-1829) (6°10’32”S 50°09’28”W), 1♀ ( IBSP 208569); 02-29/IV/2015, 1♀ ( IBSP 208755); Cave N8_0032 (GEM-2048) (6°09’53”S 50°09’30”W), 1♀ ( IBSP 207668); Cave N8_0033 (GEM-2049) (6°09’58”S 50°09’14”W), 1♀ ( IBSP 207669); 2♂ 4♀ ( IBSP 207670-207672); 16/VII-06/VIII/2014, 1♀ ( IBSP 208521); Cave N8_0038 (GEM-2055) (6°10’24”S 50°08’49”W), 3♂ 11♀ ( IBSP 208522-208527); 02-29/IV/2015, 1♀ ( IBSP 207673); 2♀ ( IBSP 207674), all collected by Equipe Carste et al.; Flona de Carajás, Cave?, no data, W.P. Munguia col., 1♀ ( IBSP 165891); Cave N1-0074 (6°1′16,53″S 50°16′49,27″W), 11/VI-02/VII/2014, 2♂ 2♀ 2 imm. ( IBSP 265416); Cave N1-0149 (6°2′30,98″S 50°16′28,09″W), 1♂ 1♀ 1 imm. ( IBSP 265417-265417); Cave N1-0204 (6°2′41,80″S 50°16′36,31″W), 24/II-13/ III/2015, 1♂ 1♀ 1 imm. ( IBSP 265415); Cave N1_0237 (6°1’15’’S 50°16’25’’W), 11/VI-02/VII/2014, 1♂ 1♀ 1 imm. ( IBSP 186183); Cave N1_0238 (6°1’15’’S 50°16’25’’W), 1♂ 4♀ ( IBSP 186184-186186); Cave N1_0246 (6°0’31’’S 50°18’6’’W), 1♀ ( IBSP 186187); Cave N1_0222 (6°0’35’’S 50°17’28’’W), 04/IX-06/X/2014, 3♀ 6 imm. ( IBSP 186188); Cave N1_0240 (6°1’19’’S 50°16’26’’W), 1♀ ( IBSP 186189); Cave N1_0056 (GEM-1254) (6°1’11’’S 50°16’44’’W), 07-28/I/2015, 1♀ 1 imm. ( IBSP 186192); all collected by Equipe Carste et al.; Cave N1_0228 (6°01’09”S 50°17’01”W), 2♂ 1♀ ( IBSP 209677-209679), all collected in 06/II/2020 by BioEspeleo Consultoria Ambiental; Cave N2_0001 (GEM-1930) (6°03’16’’S 50°14’23’’W), 26/IX-17/X/2012, 1♂ 1♀ ( IBSP 178316-178317); Cave N2_0005 (GEM-1989) (6°03’08’’S 50°14’30’’W), 1♂ ( IBSP 178318); 03-17/IV/2013, 6♀ ( IBSP 178335-178336); Cave N2_0008 (GEM-1992) (6°03’13’’S 50°14’21’’W), 1♀ ( IBSP 178319); 2♂ 1♀ ( IBSP 178337); Cave N2_0012 (GEM-1996) (6°03’08’’S 50°14’30’’W), 26/IX-17/X/2012, 1♂ 1♀ ( IBSP 178320-178321; 1♀ ( IBSP 178338); Cave N2_0013 (GEM-1997) (6°03’09’’S 50°14’32’’W), 1♂ ( IBSP 178322); 2♂ 1♀ ( IBSP 178339); Cave N2_0016 (GEM-2001) (6°03’08’’S 50°14’33’’W), 2♀ ( IBSP 178323-178324); Cave N2_0019 (GEM-2004) (6°03’06”S 50°14’38”W), 1♀ ( IBSP 178340); Cave N2_0021 (GEM-2006) (6°03’07’’S 50°14’34’’W), 1♀ ( IBSP 178341); 03-17/IV/2013, 1♂ 1♀ ( IBSP 178343-178344); Cave N2_0022 (GEM-2007) (6°02’59’’S 50°15’01’’W), 1♂ 2♀ ( IBSP 178326-178327); Cave N2_0026 (GEM-2061) (6°03’16’’S 50°14’23’’W), 6♂ 20♀ ( IBSP 178328-178332; IBSP 178345-178351); all collected by Equipe Carste et al.; Cave N3_0002 (GEM-1871) (6°01’43’’S 50°12’02’’W), 05-17/III/2013, 1♂ 3♀ ( IBSP 178355-178356); Cave N3_0003 (GEM-1872) (6°01’44’’S 50°12’03’’W), 26/IX-17/X/2012, 1♂ 1♀ ( IBSP 178291-178292; 1♂ 2♀ ( IBSP 178352-178354;; 1♀ ( IBSP 178369); Cave N3_0004 (GEM-1873) (6°01’45’’S 50°12’02’’W), 3♀ ( IBSP 178293-178295); Cave N3_0010 (GEM-1879) (6°01’45’’S 50°12’03’’W), 4♀ ( IBSP 178296); 05-17/III/2013, 1♂ 3♀ ( IBSP 178357); Cave N3_0015 (GEM-1886) (6°02’06’’S 50°12’27’’W), 03-17/IV/2013, 1♀ ( IBSP 178358); 26/IX-17/X/2012, 1♂ ( IBSP 178297); Cave N3_ 0017 (GEM-1888) (6°02’00’’S 50°12’48’’W), 05-17/III/2013, 1♂ 4♀ ( IBSP 178359-178360); Cave N3_0018 (GEM-1906)(6°01’42’’S 50°12’01’’W), 1♂ ( IBSP 178298); Cave N3_0019 (GEM-1907) (6°01’42’’S 50°12’01’’W), 1♀ ( IBSP 178361); 05-17/III/2013, 1♀ ( IBSP 178362); Cave N3_0020 (GEM-1908) (6°02’04’’S 50°12’33’’W), 2♂ 17♀ ( IBSP 178299-178301; IBSP 178363-178368); Cave N3_0021 (GEM-1909) (6°02’37’’S 50°11’58’’W), 26/ IX-17/X/2012, 1♀ ( IBSP 178302); 02-23/VIII/2013, 1♀ ( IBSP 178571); Cave N3_0023 (GEM-1911) (6°02’35’’S 50°13’10’’W), 3♀ ( IBSP 178410- IBSP 178411); Cave N3_0026 (GEM-1914) (6°02’39’’S 50°13’09’’W), 05-17/ III/2013, 1♀ ( IBSP 178370); Cave N3_0029 (GEM-1918) (6°02’40’’S 50°13’12’’W), 26/IX-17/X/2012, 1♀ ( IBSP 178303); Cave N3_0031 (GEM-1920) (6°02’37’’S 50°13’09’’W), 1♀ ( IBSP 178304); 03-17/IV/2013, 2♂ 1♀ ( IBSP 178388-178390); Cave N3_0036 (GEM-1925) (6°02’46’’S 50°13’13’’W), 26/IX-17/X/2012, 4♀ ( IBSP 178305); Cave N3_0037 (GEM-1926) (6°02’45’’S 50°13’14’’W), 1♂ 4♀ ( IBSP 178306-178309); 05-17/III/2013, 8♀ ( IBSP 178371-178374); Cave N3_0040 (GEM-1929) (6°02’11’’S 50°13’02’’W), 1♀ ( IBSP 178375); Cave N3_0041 (GEM-1981) (6°02’22’’S 50°13’14’’W), 26/IX-17/X/2012, 1♂ 7♀ ( IBSP 178311-178312); 3♂ 3♀ ( IBSP 178376- 178377); Cave N3_0042 (GEM-1982) (6°02’22’’S 50°13’15’’W), 1♂ ( IBSP 178313); Cave N3_0043 (GEM-1983) (6°02’12’’S 50°13’04’’W), 5♀ ( IBSP 178314-178315); Cave N3_0047 (GEM-2011) (6°02’27’’S 50°13’40’’W), 03-17/IV/2013, 4♂ 7♀ ( IBSP 178391- IBSP 178394); Cave N3_0049 (GEM-2013) (6°02’25”S 50°13’36”W), 5♀ ( IBSP 178395-178397); Cave N3_0047 (GEM-2011) (6°02’27’’S 50°13’40’’W), 02-23/VIII/2013, 5♂ 1♀ ( IBSP 178412); Cave N3_0047 (GEM-2011) (6°02’27’’S 50°13’40’’W), 1♂ 4♀ ( IBSP 178413-178415); Cave N3_0049 (GEM-2013) (6°02’25”S 50°13’36”W), 2♀ ( IBSP 178416); Cave N3_0050 (GEM-2014) (6°02’24”S 50°13’39”W), 3♀ ( IBSP 178417; IBSP 178398-178399); Cave N3_0051 (GEM-2015) (6°02’37’’S 50°13’31’’W), 1♂ 1♀ ( IBSP 178418); Cave N3_0052 (GEM-2016) (6°02’25’’S 50°13’42’’W), 1♀ ( IBSP 178572); 1♀ ( IBSP 178573); Cave N3_0054 (GEM-2018) (6°02’26’’S 50°13’42’’W), 1♀ ( IBSP 178419); Cave N3_0054 (GEM-2018) (6°02’26’’S 50°13’42’’W), 03-17/IV/2013, 1♀ ( IBSP 178400); Cave N3_0058 (GEM-2024) (6°02’29’’S 50°13’22’’W), 1♀ ( IBSP 178378); Cave N3_0063 (GEM-2029) (6°02’31’’S 50°13’35’’W), 3♀ IBSP 178401-178402); 02-23/ VIII/2013, 2♀ ( IBSP 178420); 1♀ ( IBSP 178421); Cave N3_0066 (GEM-2034) (6°02’31’’S 50°13’34’’W), 1♀ ( IBSP 178422); Cave N3_0071 (GEM-2039) (6°02’36’’S 50°13’50’’W), 03-17/IV/2013, 1♀ ( IBSP 178403); Cave N3_0072 (GEM-2040) (6°02’36’’S 50°13’50’’W), 1♀ ( IBSP 178404); 1♀ ( IBSP 178423); Cave N3_0073 (GEM-2041) (6°02’36’’S 50°13’46’’W), 1♂ 3♀ ( IBSP 178405); 2♂ 5♀ ( IBSP 178425-178427); 1♀ ( IBSP 178938); Cave N3_0074 (GEM-2042) (6°02’35’’S 50°13’49’’W), 05-17/III/2013, 3♂ 20♀ ( IBSP 178379-178387; IBSP 178428- 178434); Cave N3_0077 (GEM-2046) (6°02’35’’S 50°13’49’’W), 1♀ ( IBSP 178435); Cave N3_0078 (GEM-2047) (6°02’36’’S 50°13’43’’W), 1♀ ( IBSP 178436); 1♀ ( IBSP 178406); all collected by Equipe Carste et al.; Cave N4E_0002 (GEM-1004) (6°02’25’’S 50°09’39’’W), 20/IV-04/V/2010, 1♀ ( IBSP 176272); Cave N4E_0004 (GEM-985)(6°02’26’’S 50°09’39’’W), 1♂ 1♀ ( IBSP 176273-176274); Cave N4E_0007 (GEM-981) (6°02’21’’S 50°09’36’’W), 1♀ ( IBSP 176275); Cave N4E_0008 (GEM-980) (6°02’21’’S 50°09’36’’W), 07-12/X/2008, 1♂ 4♀ ( MPEG 37252, ex IBSP 176280); 1♂ 2♀ ( ISLA, ex IBSP 176279); 3♂ 14♀ ( IBSP 174960-174961; 176278; IBSP 176281-176285); Cave N4E_0009 (GEM-983) (6°02’21’’S 50°09’36’’W), 1♀ ( IBSP 176286); Cave N4E_0010 (GEM-979) (6°02’20’’S 50°09’38’’W), 3♂ ( IBSP 174972); 3♀ ( IBSP 176287-176288); Cave N4E_0011 (GEM-978) (6°02’20’’S 50°09’38’’W), 8♀ ( IBSP 176289-176291); Cave N4WS_0012 (GEM-1147) (6°04’00’’S 50°11’46’’W), 1♀ ( IBSP 176429); Cave N4E_0013 (GEM-976) (6°02’18’’S 50°09’38’’W), 5♀ ( IBSP 174980; IBSP 176293- 176295); Cave N4E_0014 (GEM-991) (6°02’17’’S 50°09’37’’W), 4♂ 15♀ ( IBSP 174990; IBSP 176296-176303); Cave N4E_0015 (GEM-997) (6°02’10’’S 50°09’35’’W), 2♂ ( IBSP 176276); Cave N4E_0020 (GEM-1015) (6°02’02’’S 50°09’35’’W), 1♂ ( IBSP 176304); Cave N4E_0021 (GEM-1014) (6°02’02’’S 50°09’37’’W), 2♂ ( IBSP 176305-176306); Cave N4E_0022 (GEM-1100) (6°02’02’’S 50°10’04’’W), 07-12/X/2008, 3♂ 15♀ ( IBSP 175014; - IBSP 175018; IBSP 176307-176314); Cave N4E_0023 (GEM-1102) (6°02’01’’S 50°10’07’’W), 1♂ 7♀ ( IBSP 176315-176317); Cave N4E_0026 (GEM-1121) (6°02’14’’S 50°10’03’’W), 07-12/ X/2008, 1♂ ( IBSP 175029); 08-12/II/2007, 2♀ ( IBSP 175031); Cave N4E_0026 (GEM-1121) (6°02’14’’S 50°10’03’’W), 18/VIII-03/IX/2009, 4♂ 17♀ ( IBSP 176205-176216); Cave N4E_0027 (GEM-1120) (6°02’14”S 50°10’03”W), 10-17/IV/2016, 1♀ ( IBSP 193575); 1♂ 1♀ ( IBSP 193576); Cave N4E_0028 (GEM-1119) (6°02’14”S 50°10’02”W), 30/I/2018, 1♂ 2♀ ( IBSP 198020); 18/VIII-03/IX/2009, 1♂ 14♀ ( IBSP 176217-176218); Cave N4E_ 0031 (GEM-1003) (6°02’24’’S 50°09’39’’W), 1♀ ( IBSP 176220); Cave N4E_0033 (GEM-989) (6°02’25’’S 50°09’36’’W), 07-12/X/2008, 1♂ 1♀ ( IBSP 175061); 15-22/IX/2009, 1♂ 4♀ ( IBSP 176221-176223); Cave N4E_ 0039 (GEM-1108) (6°01’58’’S 50°09’39’’W), 24-30/VII/2009, 1♀ ( IBSP 176224); 19/II-04/III/2010, 2♀ ( IBSP 176239-176240); Cave N4E_0041 (GEM-1110) (6°01’59’’S 50°09’42’’W), 24-30/VII/2009, 1♀ ( IBSP 176225); Cave N4E_0044 (GEM-1113) (6°01’55’’S 50°09’50’’W), 2♀ ( IBSP 176226); 19/II-04/III/2010, 1♀ ( IBSP 176230); Cave N4E_0045 (GEM-987) (6°02’25’’S 50°09’40’’W), 3♂ 3♀ ( IBSP 176231-176233); Cave N4E_0051 (GEM-1008) (6°02’22’’S 50°09’38’’W), 24-30/VII/2009, 1♀ ( IBSP 176227); Cave N4E_0055 (GEM-1105) (6°01’55’’S 50°09’59’’W), 19/II-04/III/2010, 1♂ 1♀ ( IBSP 176234); Cave N4E_0057 (GEM-1112) (6°01’55’’S 50°09’49’’W), 7♀ ( IBSP 176228-176229; IBSP 176235-176238-); ((); Cave N4E_0062 (GEM-1520) (6°02’01’’S 50°09’12’’W), 2♀ ( IBSP 76241-176242)); 1♀ ( IBSP 176194); 1♂ ( IBSP 176196); Cave N4E_0070 (GEM-1536) (6°01’56’’S 50°09’10’’W), 3♂ 3♀ ( IBSP 176243; IBSP 176245-176246); Cave N4E_0072 (GEM-1541) (6°01’56’’S 50°09’13’’W), 1♂ 13♀ ( IBSP 176247- 176254; IBSP 176197-176198); Cave N4E_0073 (GEM-1542) (6°01’57’’S 50°09’14’’W), 19/II-04/III/2010, 1♂ 10♀ ( IBSP 176255; IBSP 176261; IBSP 176200-176203);Cave N4E_0074 (GEM-1543) (6°01’59’’S 50°09’21’’W), 19/II-04/III/2010, 1♀ ( IBSP 176260); Cave N4E_0076 (GEM-1545) (6°01’59’’S 50°09’03’’W), 1♀ ( IBSP 176262); Cave N4E_0077 (GEM-1546) (6°01’58’’S 50°09’03’’W), 2♀ ( IBSP 176263; IBSP 176193); Cave N4E_0088 (GEM-1561) (6°01’54’’S 50°09’03’’W), 1♀ ( IBSP 176264); Cave N4E_ 0089 (GEM-1562) (6°01’59’’S 50°09’06’’W), 2♂ 5♀ ( IBSP 176266-176267); 24-30/VII/2009, 1♂ 4♀ ( IBSP 176195); Cave N4E_0092 (GEM-1586) (6°02’22’’S 50°09’31’’W), 1♀ ( IBSP 176204); Cave N4E_0093 (GEM-1587) (6°02’22’’S 50°09’31’’W), 1♀ ( IBSP 176199); 19/II-04/III/2010, 1♂ 1♀ ( IBSP 176269); all collected by R. Andrade et. al.; Cave N4WS_0008 (GEM-1171) (6°05’22’’S 50°11’41’’W), 07-12/X/2008, 5♂ 8♀ ( IBSP 174802- 174803-); Cave N4WS_0011 (GEM-1145) (6°04’09’’S 50°11’38’’W), 20/IV-04/V/2010, 2♂ 3♀ ( IBSP 176430- 176431); Cave N4WS_0015 (GEM-1127) (6°03’59’’S 50°11’22’’W), 12♂ 55♀ ( IBSP 176432-176448; IBSP 174790; IBSP 174809; IBSP 174814; IBSP 174929); Cave N4WS_0017 (GEM-1146) (6°03’54’’S 50°11’44’’W), 20/IV-04/V/2010, 1♀ ( IBSP 176449), all collected by R. Andrade & I. Cizauskas et al. col.; Cave N4WS_0001 (GEM-1122) (6°03’46’’S 50°11’32’’W), 2♀ ( IBSP 178139); Cave N4WS_0002 (GEM-1152) (6°04’22’’S 50°11’40’’W), 2♂ 7♀ ( IBSP 178140); 18/XI-01/XII/2010, 2♀ ( IBSP 178114); Cave N4WS_0006 (GEM-1166) (6°04’36’’S 50°11’36’’W), 01-09/VI/2011, 11 ( IBSP 178141-178143; 178106); Cave N4WS_0007 (GEM-1170) (6°05’22’’S 50°11’41’’W), 1♀ ( IBSP 178144); 18/XI-01/XII/2010, 4♂ 4♀ ( IBSP 178115-178117); Cave N4WS_ 0008 (GEM-1171) (6°05’22’’S 50°11’41’’W), 10-19/V/2011, 11♂ 21♀ ( IBSP 178145- IBSP 178151; ( IBSP 178107- IBSP 178111); Cave N4WS_0009 (GEM-1172) (6°05’22’’S 50°11’41’’W), 10-19/V/2011, 4♂ 12♀ ( IBSP 178152- 178154; IBSP 178130-178134); Cave N4WS_0010 (GEM-1173) (6°05’23’’S 50°11’42’’W), 10-19/V/2011, 4♀ ( IBSP 178155-178156; IBSP 178137); Cave N4WS_0016 (GEM-1165) (6°04’35’’S 50°11’37’’W), 01-09/VI/2011, 1♂ 5♀ ( IBSP 178157; IBSP 178118); Cave N4WS_0021 (GEM-1130) (6°03’59’’S 50°11’24’’W), 1♀ ( IBSP 178158); Cave N4WS_0022 (6°03’47’’S 50°11’32’’W), 1♀ ( IBSP 178138); Cave N4WS_0032 (GEM-1138) (6°04’03’’S 50°11’32’’W), 1♀ ( IBSP 178112); Cave N4WS_0035 (GEM-1141) (6°05’06’’S 50°10’55’’W), 10-19/ V/2011, 1♀ ( IBSP 178159); Cave N4WS_0037 (GEM-1143) (6°06’43’’S 50°10’54’’W), 1♀ ( IBSP 178160); Cave N4WS_0039 (GEM-1148) (6°03’58’’S 50°11’36’’W), 1♀ ( IBSP 178161); 18/XI-01/XII/2010, 1♀ ( IBSP 178119); Cave N4WS_0041 (GEM-1150) (6°04’23’’S 50°11’40’’W), 1♀ ( IBSP 178120); Cave N4WS_0047 (GEM-1161) (6°04’31’’S 50°11’39’’W), 1♀ ( IBSP 178113); Cave N4WS_0049 (GEM-1167) (6°04’37’’S 50°11’37’’W), 01-09/ VI/2011, 1♀ ( IBSP 178162); Cave N4WS_0055 (GEM-1834) (6°04’50’’S 50°11’45’’W), 4♂ 10♀ ( IBSP 178163- 178164; IBSP 178121-178125); Cave N4WS_0058 (GEM-1837) (6°04’24’’S 50°11’39’’W), 10-19/V/2011, 2♀ ( IBSP 178165-178166); Cave N4WS_0063 (GEM-1842) (6°04’22’’S 50°11’45’’W), 18/XI-01/XII/2010, 2♀ ( IBSP 178126); Cave N4WS_0064 (GEM-1843) (6°04’52’’S 50°11’43’’W), 1♂ 1♀ ( IBSP 178135-178136); Cave N4WS_ 0065 (GEM-1844) (6°05’53’’S 50°11’26’’W), 1♀ ( IBSP 178127); 1♀ ( IBSP 178168); Cave N4WS_0067 (GEM-1846) (6°04’22’’S 50°11’30’’W), 1♀ ( IBSP 178128); Cave N4WS_0070 (GEM-1849) (6°05’47’’S 50°11’41’’W), 10-19/V/2011, 1♀ ( IBSP 178169); Cave N4WS_0072 (GEM-1851) (6°04’25’’S 50°11’23’’W), 2♀ ( IBSP 178170); Cave N4WS_0073 (GEM-1852) (6°04’25’’S 50°11’37’’W), 2♀ ( IBSP 178171); 1♀ ( IBSP 178129); Cave N4WS_ 0075 (GEM-1859) (6°04’27’’S 50°11’22’’W), 1♀ ( IBSP 178172); all collected by F.P. Franco & C.A.R. Souza et al.; Cave N4E_0073 (GEM-1542) (6°01’57”S 50°09’14”W), 07/II/2020, 1♀ ( IBSP 209682); 07/II/2020, 1♂ 2♀ ( IBSP 209683-209685); Cave N4WS_0069, 04/XI/2019, 1♂ ( IBSP 209846); all collected by BioEspeleo Consultoria Ambiental; Cave N4WS-0006 (6°4′36,13″S 50°11′36,45″W), 21-30/VII/2018, 1♂ 3♀ ( IBSP 260643); Cave N4WS-0007 (6°5′22,08″S 50°11′41,09″W), 22-31/X/2018, 1♂ 2♀ ( IBSP 260671); Cave N4WS-0009 (6°5′22,12″S 50°11′41,45″W), 1♂ 4♀ ( IBSP 260672-260673); Cave N4WS-0010 (6°5′22,64″S 50°11′42,23″W), 7♀ ( IBSP 260677-260679); Cave N4WS-0018/48 (6°4′34,60″S 50°11′37,75″W), 21-30/VII/2018, 1♂ 3♀ ( IBSP 260639- 260640); Cave N4E-0021 (6°2′1,99″S 50°9′36,55″W), 1♀ ( IBSP 260647); Cave N4WS-0049 (6°4′37,56″S 50°11′37,03″W), 1♀ ( IBSP 260637); Cave N4WS-0055 (6°4′49,98″S 50°11′44,75″W), 2♀ ( IBSP 260650); Cave N4WS-0064 (6°4′51,74″S 50°11′43,09″W), 1♀ ( IBSP 260645), all collected by Brandt Meio Ambiente et al; Cave N5S_0004 (GEM-1054) (6°06’20’’S 50°08’02’’W), 14-23/X/2009, 1♀ ( IBSP 176799); Cave N5S_0007 (GEM-1044) (6°06’20’’S 50°07’59’’W), 4♀ ( IBSP 176793-176797); Cave N5S_0010 (GEM-1040) (6°06’20’’S 50°07’53’’W), 3♀ ( IBSP 176804-176805); Cave N5S_0011 (GEM-1036) (6°06’18’’S 50°07’47’’W), 1♂ 5♀ ( IBSP 176806-176809; ( IBSP 174531); ( IBSP 174933); Cave N5S_0012 (GEM-1034) (6°06’11’’S 50°07’31’’W), 14-23/ X/2009, 1♂ 14♀ ( IBSP 176810-176816); Cave N5S_0014 (GEM-1051) (6°06’19’’S 50°08’01’’W), 2♀ ( IBSP 176802-176803); Cave N5S_0017 (GEM-1029) (6°05’15’’S 50°07’11’’W), 25/VIII-03/IX/2009, 4♀ ( IBSP 176782- 176783); Cave N5S_0018 (GEM-1024) (6°05’11’’S 50°07’39’’W), 2♀ ( IBSP 176790); Cave N5S_0020 (GEM-1020) (6°05’15’’S 50°07’35’’W), 6♀ ( IBSP 176778-176780); Cave N5S_0021 (GEM-1018) (6°05’15’’S 50°07’34’’W), 1♀ ( IBSP 176781); Cave N5S_0032 (GEM-1033) (6°05’34’’S 50°07’08’’W), 14-16/XII/2010, 3♀ ( IBSP 178174); Cave N5S_0073 (GEM-1088) (6°06’01’’S 50°08’05’’W), 1♀ ( IBSP 176818); Cave N5S_0074 (GEM-1090) (6°06’02’’S 50°08’05’’W), 14♀ ( IBSP 176819-176820; IBSP 176784-176788); Cave N5S_0075 (GEM-1090) (6°06’02’’S 50°08’01’’W), 1♀ ( IBSP 176777); Cave N5S_0079 (GEM-1094) (6°06’08’’S 50°08’13’’W), 14/III-04/IV/2010, 1♀ ( IBSP 176821); all collected by R. Andrade & I. Cizauskas et al.; Cave N5S_0072 (GEM-1087) (6°06’03”S 50°08’06”W), 11/I/2018, 1♀ ( IBSP 198016); Cave N5S_0073 (GEM-1088) (6°06’01”S 50°08’05”W), 1♀ ( IBSP 198017); Cave N5S_0075 (GEM-1090) (6°06’02”S 50°08’01”W), 29/I/2018, 1♀ ( IBSP 198019); all collected by R. Andrade et. al.; Cave N5S_0074 (GEM-1090) (6°06’03”S 50°08’06”W), 04/II/2020,BioEspeleo ConsultoriaAmbiental col., 1♀ ( IBSP 209680);Cave N5E_0005(6°05’09”S 50°07’46”W), 1♀ ( IBSP 209674); Cave N5E_0005 (6°05’09” S 50°07’46”W), 1♂ ( IBSP 209675); all collected in 06/XI/2019 by BioEspeleo Consultoria Ambiental; Cave N5PSE-04 ( PESE _0004 ou GEM-1864) (6°06’19’’S 50°09’06’’W), 10- 19/V/2011, I. Cizauskas et al. col., 1♀ ( IBSP 178173); Cave N5W_0008 (6°04’54’’S 50°08’04’’W), 02-23/ VIII/2013, Equipe Carste col., 1♀ ( IBSP 178437); Cave N5SM2_0057 (GEM-1692) (6°07’47”S 50°08’05”W), 01/VII/2015, 2♀ ( IBSP 188668); Cave N5SM2_0075 (GEM-1743) (6°07’32”S 50°07’55”W), 02/VII/2015, 8♀ ( IBSP 188669-188670); all collected by X. Prous & T. Pietrobon; Cave N5SM2_0093 (GEM-1731), 24/IX/2010, Equipe Carste et al. col., 3♂ 9♀ 9 imm. ( IBSP 260669); Cave PPOR_0002 (GEM-2433) (6°03’23’’S 50°04’48’’W), 26/IX-17/X/2012, 3♀ ( IBSP 178333); 26/IX-17/X/2012, Equipe Carste col., 1♂ 1♀ ( IBSP 178334); 03-17/IV/2013, 2♀ ( IBSP 178407); 1♀ ( IBSP 178409); all collected by Equipe Carste col.; Cave GEM-1564 (6°19′4,41″S 50°5′48,20″W), 2♀ ( IBSP 174509); Cave GEM-1570 (6°19’37.37”S 50°6’48.93”W), 3♂ 7♀ ( IBSP 174511); 1♂ 10♀ ( IBSP 174512), all collected in 17-24/X/2008 by R. Andrade et al. col.; Cave CAV-08, JAGUAR (6°29’52’’S 51°09’43’’W), 2♂ 3♀ ( IBSP 175966); 1♀ ( IBSP 175967), all collected in 08-15/III/2012 by Equipe Carste col.; Canaã dos Carajás, Serra da Bocaina, Cave SB _0131 (GEM-1599) (6°21’15’’S 49°59’10’’W), 20-26/VI/2013, C. A. R. Souza & J. Mascarenhas et al., 1♀ ( IBSP 183537); Serra dos Carajás, Cave ST_0039 (6°18’39”S 50°05’56”W), 25/I/2016, BioEspeleo Consultoria Ambiental col., 1♂ 1♀ ( IBSP 192832); Cave S11C_0002 (6°22’54’’S 50°22’47’’W), 16/IV/2016, 2♀ ( IBSP 192038); Cave S11C_0008 (6°22’33”S 50°23’09”W), 29/VII/2015, 1♀ ( IBSP 192642); Cave S11C_0009 (6°22’34”S 50°23’10”W), 1♀ ( IBSP 192641); Cave S11C_0011 (6°22’42”S 50°22’48”W), 21/IV/2016, 1♂ 4♀ ( IBSP 192886); Cave S11C_0014 (6°24’10’’S 50°22’57’’W), 14/IV/2016, 1♀ ( IBSP 191685); 01/IX/2015, 1♂ 2♀ ( IBSP 191677); 1♂ 4♀ ( IBSP 192045); Cave S11C_0015 (6°24’10”S 50°22’57”W), 1♂ 1♀ ( IBSP 192037); Cave S11C_0016 (6°23’54”S 50°22’57”W), 20/VIII/2015, 1♂ ( IBSP 191689); Cave S11C_0023 (6°24’15”S 50°23’12”W), 16/III/2016, 2♀ ( IBSP 191683); Cave S11C_0029 (6°23’50’’S 50°23’22’’W), 29/VIII/2015, 2♀ ( IBSP 192057); Cave S11C_0038 (6°22’41”S 50°22’47”W), 03/VIII/2015, 1♀ ( IBSP 192647); Cave S11C_0046 (6°24’02”S 50°22’43”W), 02/IX/2015, 1♀ ( IBSP 192882); Cave S11C_0083 (6°24’05”S 50°22’50”W), 11/III/2016, 6♀ ( IBSP 192319); Cave S11C_0086 (6°23’58”S 50°22’47”W), 1♀ ( IBSP 192316); Cave S11C_0090 (6°24’05”S 50°22’48”W), 1♀ ( IBSP 192314); Cave S11C_0098 (6°24’10”S 50°23’08”W), 26/VIII/2015, 1♀ ( IBSP 192318); 14/IV/2016, 1♂ ( IBSP 192649); Cave S11C_0113 (6°22’56”S 50°23’36”W), 07/VIII/2015, 4♀ ( IBSP 192322); Cave S11C_0114 (6°22’57”S 50°23’36”W), 1♀ ( IBSP 192326); Cave S11C_0117 (6°23’00”S 50°23’40”W), 25/VIII/2015, 1♀ ( IBSP 191672); 1♀ ( IBSP 192315); 20/IV/2016, 1♂ 15♀ ( IBSP 192323); Cave S11C_0119 (6°23’02”S 50°23’45”W), 15/IV/2016, 1♀ ( IBSP 192656); Cave S11C_0120 (6°22’59”S 50°23’40”W), 11/IV/2016, 1♂ 1♀ ( IBSP 192327); 10/VIII/2015, 1♂ 2♀ ( IBSP 193200); Cave S11C_ 0122 (6°22’55”S 50°23’36”W), 07/VIII/2015, 2♀ ( IBSP 192655); Cave S11C_0123 (6°22’56”S 50°23’36”W), 14/III/2016, 2♀ ( IBSP 192317); 1♀ ( IBSP 192648); 07/VIII/2015, 1♀ ( IBSP 192822); Cave S11C_0124 (6°22’58”S 50°23’40”W), 08/VIII/2015, 4♀ ( IBSP 192308); 11/IV/2016, 2♀ ( IBSP 192325); Cave S11C_0126 (6°22’58”S 50°23’40”W), 08/VIII/2015, 2♂ 4♀ ( IBSP 192321; IBSP 192324; IBSP 192651); Cave S11C_0129 (6°22’58”S 50°23’39”W), 2♀ ( IBSP 192638); Cave S11C_0134 (6°22’52”S 50°23’35”W), 05/VIII/2015, 2♀ ( IBSP 192313); 14/III/2016, 3♂ 5♀ ( IBSP 192538); Cave S11C_0142 (6°22’59”S 50°23’43”W), 15/IV/2016, 1♀ ( IBSP 192653); Cave S11C_0151 (6°21’59”S 50°23’21”W), 12/IV/2016, 2♀ ( IBSP 192311); 3♀ ( IBSP 192534); Cave S11C_0153 (6°22’03”S 50°23’21”W), 01/VIII/2015, 5♀ ( IBSP 191671); 15/III/2016, 1♀ ( IBSP 192310); 6♀ ( IBSP 192532); 01/VIII/2015, 5♀ ( IBSP 192637); Cave S11C_0156 (6°22’06”S 50°23’21”W), 03/VIII/2015, 1♀ ( IBSP 192068); Cave S11C_0165 (6°24’31”S 50°24’05”W), 07/IV/2016, 1♂ 3♀ ( IBSP 192900); Cave S11C_0169 (6°22’47”S 50°22’44”W), 09/IX/2015, 4♀ ( IBSP 192535); Cave S11C_0170 (6°21’56”S 50°22’58”W), 28/VII/2015, 8♀ ( IBSP 192052); 05/IV/2016, 3♀ ( IBSP 192053); 1♂ 2♀ ( IBSP 192062); 28/VII/2015, 2♀ ( IBSP 192635); Cave S11C_ 0181 (6°24’32”S 50°23’45”W), 07/IV/2016, 5♀ ( IBSP 192054); Cave S11C_0184 (6°24’35”S 50°23’49”W), 03/ IX/2015, 1♀ ( IBSP 192644); Cave S11C_0189 (6°21’33”S 50°23’04”W), 19/III/2016, 1♀ ( IBSP 192639); Cave S11C_0190 (6°21’34”S 50°23’05”W), 1♀ ( IBSP 192047); Cave S11C_0191 (6°21’58”S 50°22’56”W), 1♀ ( IBSP 192070); Cave S11C_0192 (6°21’36”S 50°23’08”W), 5♀ ( IBSP 192058-192060); Cave S11C_0201 (6°22’01”S 50°23’07”W), 27/VII/2015, 4♀ ( IBSP 192056); 12/IV/2016, 1♀ ( IBSP 192063); 2♀ ( IBSP 192065); 1♀ ( IBSP 192640); Cave S11C_0202 (6°21’59”S 50°22’55”W), 05/IV/2016, 2♀ ( IBSP 192051); 1♂ 1♀ ( IBSP 192061); 3♀ ( IBSP 192093); Cave S11C_0204 (6°21’45”S 50°23’02”W), 12/IV/2016, 1♀ ( IBSP 192049); Cave S11C_0206 (6°21’43”S 50°23’12”W), 25/VII/2015, 1♀ ( IBSP 192059); 1♀ ( IBSP 192064); 18/III/2016, 1♀ ( IBSP 192067); Cave S11C_0214 (6°21’47”S 50°23’03”W), 31/VII/2015, 1♀ ( IBSP 191679); 2♂ 4♀ ( IBSP 192042); all collected by BioEspeleo Consultoria Ambiental; Cave SB_0026 (GEM-1423) (6°18’45”S 49°59’55”W), 17/I-02/II/2012, 1♀ ( IBSP 196547); Cave SB_0066 (GEM-1482) (6°18’44”S 49°52’57”W), 05-15/III/2012, 1♀ ( IBSP 196570), all collected by F. Pellegatti-Franco et. al.; Floresta Nacional de Canaã dos Carajás, in caves, 28♂ 10♀ 12 imm. ( IBSP 115502-115508);, all collected in 28/XI-03/X/2007 by R. Andrade et. al.; 23/VIII-02/IX/2007, 1♂ 1 imm. ( IBSP 97783); Cave N1-72 (Tio Edy) (6°1′13,28″S 50°17′17,51″W), 28-IX- 03/X/2007, 8♂ 2♀ 6 imm. ( IBSP 97873); Cave N1-18 (Fome Zero) (6°2′1,39″S 50°16′17,23″W), 1♂ 1imm. ( IBSP 97876); 28-IX- 03/X/2007, 6♂ 4 imm. ( IBSP 97881); Gruta N1-170 (Pereira) (6°1′23,56″S 50°17′58,09″W), 3♂ 3 imm. ( IBSP 97681); Gruta N1-173 (Escondida) (6°1′27,20″S 50°17′55,42″W), 4♂ 4 imm. ( IBSP 97676); Gruta N4_E07 (6°2′21,10″S 50°9′35,74″W), 20/X-01/XI/2006, 1♂ 1 imm. ( IBSP 97845); Gruta N4_E08 (6°2′20,84″S 50°9′35,74″W), 5♂ 2♀ 3 imm. ( IBSP 97861); Gruta N4_E11 (6°2′19,87″S 50°9′38,11″W), 1♂ 1♀ ( IBSP 97863); 3♂ 3 imm. ( IBSP 97849); Gruta N4_ E13 (6°2′17,65″S 50°9′37,69″W), 2♂ 1♀ 1 imm. ( IBSP 97828); Gruta N4_E22 (6°2′1,87″S 50°10′4,00″W), 10♂ 2♀ 8 imm. ( IBSP 97842); Gruta N4_ E22 (6°2′1,87″S 50°10′4,00″W), 2♂ 1♀ 1 imm. ( IBSP 97853); Gruta N4_E28 (6°2′13,82″S 50°10′1,71″W), 08-12/II/2007, 1♂ 1♀ ( IBSP 97771); Gruta N4_E61 (6°2′20,75″S 50°10′2,96″W), 1♂ 1 imm. ( IBSP 97766); Gruta S11_D40 (6°24′38,43″S 50°19′29,24″W), 23/VIII-02/IX/2007, 1♂ 1 imm. ( IBSP 97775); Gruta S11_D55 (6°24′22,81″S 50°19′12,37″W), 1♂ 1 imm. ( IBSP 97792); FLONA de Carajás, Cave S11D_0001 (650) (6°23’54’’S 50°21’25’’W), 13-30/I/2010, 24♀ ( IBSP 172307-172315); 01-14/VII/2010, 7♀ ( IBSP 172393-172395); Cave S11D_0005 (647) (6°24’03’’S 50°20’60’’W), 13-30/I/2010, 1♀ ( IBSP 172338); Cave S11D_0006 (648) (6°24’04’’S 50°21’01’’W), 19-22/II/2010, 2♀ ( IBSP 172343); 1♀ ( IBSP 172344); 01-14/ VII/2010, 2♀ ( IBSP 172396-172397); Cave S11_0007 (S11-07) (6°27’20’’S 50°14’30’’W), 24/II-04/III/2010, 2♀ ( IBSP 172366); Cave S11D_0010 (651) (6°23’54’’S 50°21’25’’W), 13-30/I/2010, 2♀ ( IBSP 172297); Cave S11D_ 0011 (652) (6°23’51’’S 50°21’30’’W), 1♀ ( IBSP 172331); Cave S11D_0012 (653) (6°23’46’’S 50°21’34’’W), 25♀ ( IBSP 172298-172306); 01-14/VII/2010, 13♀ ( IBSP 172398-172400); Cave S11D_0013 (654) (6°24’06’’S 50°21’08’’W), 18♀ ( IBSP 172326-172327; IBSP 172401-172402); Cave S11D_0014 (655) (6°24’06’’S 50°21’10’’W), 8♀ ( IBSP 172403-172404; IBSP 172318; 172319; IBSP 172363); Cave S11D_0017 (682) (6°23’56’’S 50°21’23’’W), 5♀ ( IBSP 172295; IBSP 172405-172406); Cave S11_0018 (S11-18) (6°26’08’’S 50°17’44’’W), 1♀ ( IBSP 172365); Cave S11D_0021 (662) (6°24’43’’S 50°21’35’’W), 19-22/II/2010, 1♀ ( IBSP 172351); Cave S11D_ 0022 (663) (6°24’46’’S 50°21’33’’W), 3♀ ( IBSP 172352-172353); Cave S11D_0023 (664) (6°24’46’’S 50°21’32’’W), 4♀ ( IBSP 172349-172350); 24/II-04/III/2010, R. Andrade & I. Cizauskas et al. col., 2♀ ( IBSP 172361); 03-19/VIII/2010, 1♀ ( IBSP 172437); Cave S11D_0024 (2152) (6°24’46’’S 50°21’15’’W), 19-22/II/2010, 2♂ 13♀ ( IBSP 172345-172346; IBSP 1723470; IBSP 172348; IBSP 172407-172408); Cave S11_0025 (S11-25) (6°25’18’’S 50°18’19’’W), 24/II-04/III/2010, 1♂ ( IBSP 172362); Cave S11D_0026 (667) (6°24’49’’S 50°21’17’’W), 2♀ ( IBSP 172354); Cave S11_0027 (S11-27) (6°24’24’’S 50°14’56’’W), 1♀ ( IBSP 172364); 1♀ ( IBSP 172438); Cave S11D_0028 (669) (6°24’40’’S 50°21’06’’W), 19-22/II/2010, 1♀ ( IBSP 172355); Cave S11D_0029 (670) (6°24’40’’S 50°20’44’’W), 1♂ ( IBSP 172357); Cave S11D_0029 (670) (6°24’40’’S 50°20’44’’W), 1♀ ( IBSP 172358); 03-19/VIII/2010, 1♂ 6♀ ( IBSP 172409-172411); Cave S11D_0033 (675) (6°24’40’’S 50°20’37’’W), 1♀ ( IBSP 172329); Cave S11D_0037 (680) (6°24’46’’S 50°21’31’’W), 9♀ ( IBSP 172359-172360; IBSP 172415- 172416); Cave S11D_0038 (683) (6°23’49’’S 50°20’26’’W), 5♀ ( IBSP 172325; IBSP 172417); Cave S11D_0039 (39) (6°23’46’’S 50°20’27’’W), 23/VIII-02/IX/2007, 12♀ ( IBSP 174474; IBSP 172340-172342; IBSP 172418- 172421); Cave S11D_0040 (685) (6°24’38’’S 50°19’29’’W), 13-30/I/2010, 13♀ ( IBSP 172333-172334; IBSP 172422-172423); Cave S11D_0043 (689) (6°24’48’’S 50°19’17’’W), 11♀ ( IBSP 172335-172337; IBSP 174648; IBSP 172424-172425); Cave S11D_0044 (690) (6°25’01’’S 50°18’55’’W), 13-30/I/2010, 1♂ ( IBSP 172317); Cave S11D_0046 (692) (6°24’53’’S 50°18’59’’W), 1♀ ( IBSP 172332); 03-19/VIII/2010, 4♀ ( IBSP 172426); Cave S11D_0047 (693) (6°24’38’’S 50°18’55’’W), 1♀ ( IBSP 172339); 01-14/VII/2010, 1♀ ( IBSP 172427); Cave S11D_ 0050 (696) (6°24’25’’S 50°19’14’’W), 1♀ ( IBSP 172316); Cave S11D_0055 (701) (6°24’23’’S 50°19’12’’W), 1♀ ( IBSP 172320); 4♀ ( IBSP 172428); Cave S11D_0064 (710) (6°23’31’’S 50°18’48’’W), 3♀ ( IBSP 172321-172322); Cave S11D_0066 (712) (6°23’34’’S 50°18’53’’W), 1♂ ( IBSP 172324); Cave S11D_0069 (715) (6°23’34’’S 50°19’07’’W), 4♀ ( IBSP 172323); 4♀ ( IBSP 172429); Cave S11D_0077 (723) (6°23’33’’S 50°18’59’’W), 1♂ 2♀ ( IBSP 172279); Cave S11D_0079 (725) (6°23’33’’S 50°18’56’’W), 4♀ ( IBSP 172290-172291); 01-14/VII/2010, 1♀ ( IBSP 172379); Cave S11D_0081 (727) (6°23’34’’S 50°18’53’’W), 5♀ ( IBSP 172380-172381; IBSP 172284- 172285); Cave S11D_0086 (732) (6°23’47’’S 50°19’24’’W), 1♀ ( IBSP 172292); Cave S11D_0087 (733) (6°23’46’’S 50°19’23’’W), 1♀ ( IBSP 172382); Cave S11D_0088 (734) (6°23’45’’S 50°19’22’’W), 1♀ ( IBSP 172383); 4♀ ( IBSP 172286-172288); Cave S11D_0089 (735) (6°23’45’’S 50°19’20’’W), 2♀ ( IBSP 172289); Cave S11D_0093 (739) (6°23’41’’S 50°19’18’’W), 3♀ ( IBSP 172384; IBSP 172280-172385); Cave S11D_0094 (740) (6°23’40’’S 50°19’17’’W), 2♀ ( IBSP 172386-172387); Cave S11D_0095 (741) (6°23’37’’S 50°19’26’’W), 2♀ ( IBSP 172388); 1♀ ( IBSP 172278); Cave S11D_0096 (742) (6°23’37’’S 50°19’27’’W), 2♂ 5♀ ( IBSP 172389-172391; IBSP 172293-172294); Cave S11D_0099 (745) (6°23’44’’S 50°20’27’’W), 2♀ ( IBSP 172392); Cave S11D_0101 (747) (6°23’22’’S 50°21’48’’W), 2♂ 5♀ ( IBSP 172281-172283); 1♂ 4♀ ( IBSP 172412-172414); Cave S11D_0104 (6°23’50’’S 50°21’59’’W), 30/VII-02/IX/2011, 1♀ ( IBSP 172440); 14-19/XII/2011, 1♀ ( IBSP 172441); Cave S11D_0112 (2153) (6°24’46’’S 50°21’14’’W), 1♂ ( IBSP 174073); Cave S11D_114 (TRL-02) (6°25’15’’S 50°18’53’’W), 2♀ ( IBSP 172442); Cave S11D_116 (TRL-04) (6°25’18’’S 50°18’59’’W), 3♀ ( IBSP 172439); 1♀ ( IBSP 172443); all collected by R. Andrade & I. Cizauskas et al.; Cave S11B_0009 (6°21′18,16″S 50°23′22,62″W), 10/XI/2018, 1♀ ( IBSP 265397); Cave S11B_0076B (6°21′30,95″S 50°23′44,67″W), 2018, 1♀ ( IBSP 265411); Cave S11B_0177 (6°20′21,12″S 50°24′55,52″W), 22/III/2019, 2♀ ( IBSP 265398-265399); 24/IX/2018, 1♀ ( IBSP 265413), Cave S11B_0178 (6°20′54,46″S 50°24′49,88″W), 1♂ 5♀ ( IBSP 265400-265404); Cave S11B_0186 (6°20′23,04″S 50°24′56,00″W), 2♀ ( IBSP 265409); Cave S11B_0187 (6°20′22,94″S 50°24′56,00″W), 13/IX/2018, 1♀ ( IBSP 265405); Cave S11B_0190 (6°20′28,86″S 50°24′49,13″W), 11/I/2019, 1♂ ( IBSP 265406); 15/IX/2018, 1♀ ( IBSP 265410); Cave S11B_0193 (6°20′27,56″S 50°24′50,72″W), 19/XI/2018, 1♀ ( IBSP 265407); 23/VI/2018, 1♀ ( IBSP 265412); Cave S11B_0212 (6°20′4,69″S 50°25′39,83″W), 18/I/2019, 1♀ ( IBSP 265408), all collected by Ativo Ambiental; Cave CAV_0001 (CAV-01) (6°24’42’’S 50°20’05’’W), 22-31/V/2010, 2♀ ( IBSP 172367); Cave CAV_0003 ( CAV _03) (6°24’41’’S 50°20’05’’W), 1♀ ( IBSP 172368); Cave CAV_0006 ( CAV _06) (6°24’40’’S 50°19’57’’W), 2♀ ( IBSP 172369); 2♀ ( IBSP 172430); Cave CAV_0012 (CAV-12) (6°24’38’’S 50°19’38’’W), 1♂ ( IBSP 172370); Cave CAV_0018 (CAV-18) (6°24’23’’S 50°22’09’’W), 1♀ ( IBSP 172371); 3♀ ( IBSP 172431); Cave CAV_0019 (CAV-19) (6°24’21’’S 50°22’08’’W), 1♀ ( IBSP 172432); Cave CAV_0021 (CAV-21) (6°24’20’’S 50°22’10’’W), 1♀ ( IBSP 172433); 22-31/V/2010, 1♀ ( IBSP 172372); Cave CAV_0024 (CAV-24) (6°24’20’’S 50°21’57’’W), 22-31/V/2010, 1♂ ( IBSP 172373); Cave CAV_0034 (CAV-34) (6°24’09’’S 50°22’56’’W), 3♂ 4♀ ( IBSP 172374); 2♀ ( IBSP 172375); 1♂ ( IBSP 172376); 5♀ ( IBSP 172434); Cave CAV_0035 (CAV-35) (6°24’23’’S 50°23’06’’W), 2♀ ( IBSP 172377); 2♀ ( IBSP 172435); Cave CAV_0041 (CAV-41) (6°24’56’’S 50°21’50’’W), 1♀ ( IBSP 172378); 1♀ ( IBSP 172436); all collected by R. Andrade & I. Cizauskas et al.; Curionópolis, Cave CRIS-008 (6°27’07’’S 49°40’42’’W), 29/VII-06/VIII/2008, 1♀ ( IBSP 174567); Cave CRIS-013 (6°26’50’’S 49°40’56’’W), 1♀ ( IBSP 174582); Cave CRIS-014 (6°26’49’’S 49°40’57’’W), 2♂ 3♀ ( IBSP 174584); Cave CRIS-025 (6°27’34’’S 49°42’38’’W), 6♀ ( IBSP 174614); Cave CRIS-034 (6°27’34’’S 49°42’21’’W), 4♀ ( IBSP 174638); Cave CRIS-036 (6°27’32’’S 49°42’16’’W), 1♂ 1♀ ( IBSP 174642); all collected in 29/VII-06/VIII/2008 by R. Andrade et al.; Cave CRIS-017 (6°26’21’’S 49°40’54’’W), 2♀ ( IBSP 175945-175946); all collected in 23/II/2011 by R. Andrade & V. Felice; Cave SL_0004 (05°57’49’’S 49°38’58’’W), 03/VII/2010, R. Zampaulo col., 1♀ ( IBSP 189970); Cave SL_0008 (5°57’28’’S 49°38’12’’W), 17-24/X/2008, R. Andrade et al. col., 1♂ ( IBSP 174761); Cave SL_0025 (05°58’21’’S 49°38’41’’W), 08/VI/2010, R. Zampaulo col., 1♂ ( IBSP 190031); Cave SL 27, 06/I/2011, Equipe Carste col., 1♀ ( IBSP 196700); Cave SL_0031 (5°58′19,05″S 49°38′34,89″W); 30/IX/2020, R.A. Zampaulo & T. Pietrobon col., 1♀ ( IBSP 265414); Cave SL_0061 (05°58’46’’S 49°37’19’’W), 10/VI/2010, R. Zampaulo col., 1♂ ( IBSP 190032; photo genitalia); Cave SL_0076 (05°58’01’’S 49°38’32’’W), 29/VI/2010, R. Zampaulo col., 1♂ ( IBSP 190029); Cave SL_0082 (5°57’28’’S 49°38’12’’W), 17-24/X/2008, R. Andrade et al. col., 1♀ ( IBSP 174762); Cave SL_0137 (6°00’04”S 49°36’57”W), 03/IX/2014, 1♀ ( IBSP 186555); Cave SL_0142 (06°17’34’’S 49°36’58”W), 1-19/III/2016, 1♀ ( IBSP 191327); Cave SL_0183 (6°01’15”S 49°37’31”W), 11/IX/2014, 2♀ ( IBSP 186553-186554); all collected by Equipe Spelayon.

Etymology. The specific name is a popular expression from Pará originated from the Tupi language for ended or exhausted. Noun in apposition.

Diagnosis. Speocera babau sp. nov. differs by the male having SCI in the embolus narrow and elongated, dagger shape and projecting forward slender and with large base conductor in the palp ( Figs 7E – F View FIGURE 7 ; 9A – C View FIGURE 9 )—while in S. pinima sp. nov. it is pentagonal and the small conductor, fig. 1E, and very elongated and curved at tip, with shorter conductor in S. piquira sp. nov., fig. 12E. The female of Speocera babau sp. nov. resemble those of S. pinima sp. nov. by its long afferent spermathecae and elongated lateral plates (10D – E) but differs by basally projected spermathecae ( Fig. 10E – F View FIGURE 10 ), curved distal afferent spermathecae ( Fig. 10E – F View FIGURE 10 ) and enlarged and oval terminal depression ( Fig. 10D View FIGURE 10 ) in the genitalia.

Description. Male (Holotype). Total length 1.1. Carapace length 0.5, ovoid; narrowing gradually anteriorly; yellow, flat pars cephalic, fovea absent ( Fig. 7A View FIGURE 7 ). Clypeus length 0.02, with a pair of long bristles ( Fig. 8A View FIGURE 8 ). Eyes: PME oval; ALE and PLE rounded, ranging from opalescent or white ( Fig. 8A – B View FIGURE 8 ). Chelicerae light yellow; promargin with six teeth, attached to a very long lamina ( Fig. 8C View FIGURE 8 ); retromargin without two very small teeth. Sternum yellow. Endites yellow, with large serrula with more than 15 denticles, distal macrosetae paired and crosier-like, and many multifid macrosetae present ( Fig. 8D View FIGURE 8 ). Labium yellow, sulcated distally ( Fig. 8D View FIGURE 8 ). Legs cream; formula 4123; total length: I 1.8; II 1.6; III 1.4; IV 2.0. Male palp: palpal femur length 0.02; palpal tibia enlarged basally; cymbium conical distally, bearing subdistal pair of modified setae and a projected and bifid tarsal organ ( Fig. 8F View FIGURE 8 ); bulb spherical ( Figs 7B – D View FIGURE 7 ; 9A View FIGURE 9 ); conductor longer than SCI, enlarged at base ( Figs 9A – C View FIGURE 9 ); embolus with small and conical SCII, very short SCIII and voluminous SCIV ( Figs 7E – F View FIGURE 7 ; 9A – F View FIGURE 9 ). Abdomen length 0.6, oval; uniformly gray. Three epiandrous spigots, with large, conic and striated base ( Fig. 8E View FIGURE 8 ). Colulus triangular with six bristles.

Female (Paratype, IBSP 207354). Total length: 1.05; carapace length: 0.45; Carapace as in male, light yellow pattern ( Fig. 10A View FIGURE 10 ). Pedipalp without claw, with conical tip and subdistal bifid tarsal organ ( Fig. 11D View FIGURE 11 ). Clypeus: 0.02 diameter, with setae as in male. Eyes, chelicerae, sternum, endites and labium ( Fig. 11A – C View FIGURE 11 ) as in male. Legs as in male; formula 4123, total length: I 1.6; II 1.5; III 1.2; IV 1.8. Abdomen length 0.5, yellow ( Fig. 10A View FIGURE 10 ). Colulus triangular with six bristles. Genitalia externally with long and distally curved lateral plates, large ventral plate and terminal depression sulcated ( Figs 10B – D View FIGURE 10 ; 11E – F View FIGURE 11 ); internally with together and paired spermathecae, and afferent spermathecae curved distally ( Fig. 10E – F View FIGURE 10 ).

Variation. 10 ♂♂: total length 0.9 – 1.1; carapace 0.4 – 0.6; femur I 0.8 – 0.9; 10 ♀♀: total length 0.9 – 1.12; carapace 0.4 – 0.55; femur I 0.75 – 0.85.

Distribution. Widely distributed in iron caves the Carajás mountains, municipalities of Canaã dos Carajás, Curionópolis, Eldorado dos Carajás, Parauapebas and São Félix do Xingu, state of Pará, Brazil ( Fig. 19B, D View FIGURE 19 ) in a range of approximately 170 km.

Natural history. Speocera babau sp. nov. was commonly collected in the soil, generally found on floors, along roots and under blocks inside caves. Like other representatives of the genus and with other Ochyroceratidae , they have maternal behavior and carry the egg sac in the chelicera until hatching of the eggs ( Fig. 18C–D View FIGURE 18 ).