Pupinella illustris (Mabille, 1887) Jirapatrasilp & Sutcharit & Panha, 2022

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupinella illustris (Mabille, 1887) comb. nov.

Figs 11I-L , 12D, E

Pupina illustris Mabille, 1887: 136, 137. Type locality: Tonkin. Fischer 1891: 107. Fischer and Dautzenberg 1904: 431.

Pupina tonkiniana Bavay & Dautzenberg, 1899: 54, 55, pl. 3, fig. 6, 6a (as Pupina tonkiana in the original description). Type locality: Entre Lang-Son [Lang Son Province, Vietnam] et That-Khé [That Ke, Lang Son Province, Vietnam]. Syn. nov.

Pupina (Tylotoechus) illustris - Kobelt 1902: 314, 315.

Pupina (Tylotoechus) tonkiniana - Kobelt 1902: 323, 324. Zilch 1957: 48.

Pupina tonkiniana - Fischer and Dautzenberg 1904: 432. Fischer-Piette 1950: 167. Do et al. 2015: 126, fig. 6b, Son La Province, Vietnam. Raheem et al. 2017: 5 (plate figure).

Type material examined.

Syntypes of Pupina illustris MNHN-IM-2000-35842 (9 shells; Figs 11I, J , 12D ) from Tonkin. Lectotype of Pupina tonkiniana MNHN-IM-2000-35838 (Fig. 11K ) from Lang-Son et That-Khé. Paralectotypes of Pupina tonkiniana SMF 109932/10 (10 shells; Figs 11L , 12E ) from Tonkin: That-khé. Paralectotypes of Pupina tonkiniana RBINS MT976/2 (14 shells) from Lang Son et That-khé.

Diagnosis.

Shell elongate fusiform; last whorl ca. 55-60% of shell height. Apertural lip expanded and slightly reflected; apertural lip when observed from lateral view almost straight. Parietal callus absent. Parietal tooth pointily sharp, located next to wide posterior canal. Anterior canal funnel-like. Umbilicus closed.

Differential diagnosis.

Pupinella illustris can be distinguished from all other species in mainland Southeast Asia by an elongate fusiform shell shape, an absence of parietal callus and a pointily sharp parietal tooth located next to a wide posterior canal.

Distribution.

Northern Vietnam ( Do et al. 2015; Raheem et al. 2017).

Remarks.

This taxon is allocated to the genus Pupinella due to the presence of a funnel-like anterior canal, which is the diagnostic character of this genus. In the original description of Pupina tonkiniana , two ways of spelling were shown: the spelling ' Pupina tonkiana ' in the description, and ' Pupina tonkiniana ' in the plate caption. Later, Kobelt (1902) acted as the First Reviser ( ICZN 1999: Art. 24.2.3) in selecting ' Pupina tonkiniana ' as the correct original spelling. As the original description did not explicitly state that the description of P. tonkiniana was based on a single specimen (nor could this be inferred), the designation of a holotype by Fischer-Piette (1950) in fact constitutes a lectotype designation ( ICZN 1999: Art. 74.6).

Upon examining the type specimens of both P. illustris and P. tonkiniana , the type series of P. tonkiniana agree well with all the syntypes of P. illustris in having an elongate fusiform shell shape, an expanded and slightly reflected apertural lip without a parietal callus, and a sharp, tooth-like, parietal tooth located next to a wide posterior canal. Thus, P. tonkiniana is regarded herein as a junior subjective synonym of P. illustris .

Figures

Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H).

Figure 12. A-C Pupinella mansuyi: A syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen B specimen CUMZ 12148 from Pha Chu Mount, Nan, and C specimen CUMZ 12149 from Pha Tub Cave, Nan. D, E Pupinella illustris D syntype of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin and E paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe F Pupinella sonlaensis, paratype ZRC. MOL. 9377. Photo: F. Trus, RBINS (A), P. Maestrati, MNHN (D), S. K. Tan, ZRC (F).