Dichotomius protectus (Harold, 1867)

Rosa, Cecilia Lozano De La, Cupello, Mario & Vaz-De-Mello, Fernando Z., 2024, The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution, European Journal of Taxonomy 959 (1), pp. 1-272 : 105-106

Dichotomius protectus (Harold, 1867)

Fig. 42A

Pinotus protectus Harold, 1867d: 98 (original description). Type locality: Colombia. Name-bearing type: syntypes (MNHN), examined by FZVM.

Pinotus protectus tridentatus Luederwaldt, 1923: 10 (original description). Type locality: Ecuador and Venezuela. Name-bearing type: syntypes (MZSP), not examined.

Pinotus protectus – Harold 1869b: 130 (redescription, cited for Colombia); 1869d: 1010 (catalogue, cited for Colombia). — Candèze 1891: 330 (list). — Heyne & Taschenberg 1908: 62 (list, cited for Colombia). — Gillet 1911b: 61 (catalogue). — Campos 1921: 56 (cited for Ecuador). — Luederwaldt 1929: 34 (key). — Blackwelder 1944: 207 (list). — Gacharná 1951: 222 (list for Colombia).

Dichotomius (Dichotomius) protectus – Roze 1955: 44 (cited, checklist for Venezuela). — Blanco 1988: 40, 41 (catalogue). — Havranek 1989: 61, 62 (list, comments). — Escobar 2000: 208 (checklist for Colombia). — Medina et al. 2001: 138 (checklist for Colombia). — Hamel-Leigue et al. 2006: 15 (list, cited for Bolivia). — Pulido-Herrera et al. 2007: 307 (cited for Andean region of Colombia). — Orozco & Pérez 2008: 38–39 (list for Colombia). — Sarmiento-Garcés & Amat-García 2009: 290, 295 (diagnosis, key); 2014: 51, 64 (key, diagnosis). — Carvajal et al. 2011: 320 (list for Ecuador). — Krajcik 2012: 92 (list). — Ferrer-Paris et al. 2013: 109 (list). — Ratcliffe et al. 2015: 197 (checklist for Peru). — Chamorro et al. 2018: 94 (list for Ecuador); 2019: 119 (catalogue). — Giraldo et al. 2018: 100 (guide). — Pardo-Díaz et al. 2019: 5, 7, 11 (comments). — Nieto et al. 2020: 136 (report).

Material examined

VENEZUELA ‒ Aragua • 1 spec.; Cayagua ; 50 m a.s.l.; 6 Jul. 2004; D. García leg.; CEMT • 2 specs; Parque Nacional Henri Pittier, Estación Biológica Rancho Grande ; 1500 m a.s.l.; 21‒25 Feb. 1971; S. Peck leg.; human dung trap; forest; CEMT • 2 specs; road Tiara–Caracas ; 2000 m a.s.l.; 22‒25 Feb. 1971; S. Peck leg.; carrion; forest; CEMT . ‒ Distrito Capital • 1 spec.; Caracas ; Dec. 1949; CEMT . ‒ Mérida • 2 specs; Monte Zerpa, Merida ; 2200 m a.s.l.; Jul. 1995; A. de Ascençao leg.; dung; CEMT • 1 spec.; Zea ; Nov. 2003; G. Arriágada leg.; CEMT . ‒ Miranda • 2 specs; Altos de Pipe, Instituto Venezolano de Investigaciones Científicas (“ Campus IVIC ”); 10°23′51″ N, 66°58′15″ W; 1500 m a.s.l.; Jul. 2009; F.Z. Vaz-de-Mello leg.; CEMT . ‒ Táchira • 1 spec.; near Pregonero, Futura Presa Las Cuevas ; 600 m a.s.l.; 17 Jan. 1989; D. Havranek leg.; human faeces trap; CEMT • 2 specs; near Pregonero, presa/ La Honda ( La Idea ); 1100 m a.s.l.; 16 Aug. 1989; D. Havranek leg.; mini-cup trap C(T7-6); CEMT • 1 spec.; Vía Chorro del Indio ; Aug. 1987; J. Blanco leg.; CEMT • 1 spec.; Vía Chorro del Indio ; Aug. 1987; J. Blanco leg.; CEMT .

Distribution

Colombia, Venezuela, Ecuador, Peru, and Bolivia.

Subregions of Venezuela

Andes mountains and Central Coast Mountain Range.

Literature records

Gillet 1911b: 61 ( Venezuela). — Luederwaldt 1929: 34 ( Venezuela). — Blackwelder 1944: 207 ( Venezuela). — Roze 1955: 44 ( Venezuela). — Blanco 1988: 40, 41 ( Venezuela: Táchira). — Havranek 1989: 61 ( Venezuela: Táchira state). — Ferrer-Paris et al. 2013: 109 ( Venezuela: Mérida: La Azulita-Jají; Miranda: Altos de Pipe). — Chamorro et al. 2019: 119 ( Venezuela).

Figures

Fig. 42. Species distribution. A. Dichotomius protectus (Harold, 1867). B. Dichotomius robustus (Luederwaldt, 1935). Green diamond = CEMT collection data; black circle = literature data.