Philonthus succicola, Thomson, 1860

Taphylininae, Iochini, Aorothiini, Thiini, Latyprosopini, Taphylinini, Mblyopinina, Nisolinina, Yptiomina, Hilonthina & Herman, Lee H., 2001, Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2441 (265), pp. 2435-3021 : 2971-2978

succicola ]; characters).

— Scheerpeltz, 1961d: 128 ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola ]; Italy).

— Lohse, 1964: 177 ( Philonthus ; subgenus Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola ]; characters; central Europe).

— Smetana, 1964d: 74 ( Philonthus ; subgenus Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola ]; Czechoslovakia).

— Horion, 1965: 129 ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola ]; all Europe).

— Pototskaia, 1966: 146, 158 ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola ]; larval characters).

— Pototskaia, 1967: 63 ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola ]; larval characters).

— Coiffait, 1967a: 392 ( Philonthus ; characters).

— Scheerpeltz, 1968a: 67 ( Philonthus ; synonym of chalceus ).

— Raianu, 1968a: 462 ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola ]; Romania).

— Szujecki, 1968a: 727 ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola ]; Poland).

— Szujecki, 1969a: 255 ( Philonthus ; notes; Poland).

— Jarrige, 1971: 494 ( Philonthus ; Iran).

— Bacchus and Hammond, 1972: 63 ( Philonthus ; lion dung; Britain).

— Coiffait, 1974a: 198 ( Philonthus ; Siberia).

— Coiffait, 1974: 222 ( Philonthus ; characters; Europe; Scandinavia; Britain; Mongolia).

— Coiffait, 1974: 223 ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola ]; synonym of succicola ).

— Shilov, 1975a: 375 [= 1975b: 98] ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola ]; Komi region of Russia).

— Legner and Moore, 1977: 176 ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications of succicola ]; found under bark).

— Pace and Zanetti, 1977: 289 ( Philonthus ; Italy).

— Pope, 1977: 29 ( Philonthus ; Britain).

— Muona, 1979: 14 ( Philonthus ; Finland; Russia; Sweden; Norway; Denmark; Baltic states).

— Tóth, 1984: 48 ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications

of succicola ]; characters; Hungary).

— Segers, 1986: 45 ( Philonthus ; Belgium).

— Muona and Viramo, 1986: 13 ( Philonthus ; Finland).

— Drugmand, 1987b: 14, 47 ( Philonthus ; habitat; distribution in Belgium).

— Lucht, 1987: 105 ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifications

of succicola ]; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Lohse and Lucht, 1989: 163 ( Philonthus ; notes).

— Lohse and Lucht, 1989: 163 ( Philonthus ; cited as chalceus Stephens [Note: Misidentification; based on work by Tottenham (1937a: 190; 1939d: 232) most uses of chalceus were misidentifi-

cations of succicola ]; synonym of succicola ).

sumatrensis Cameron, 1933d: 388 ( Philonthus ; Type locality: Sumatra: Fort de Kock ).

- DISTRIBUTION: Indonesia.

sumbaensis Scheerpeltz, 1957c: 234 , 324 ( Philonthus ; Type locality: Sumba: Mau Marru).

- DISTRIBUTION: Indonesia.

superbus Arrow, 1915 , see: Leucitus .

supernus Herman, 2001: 48 ( Philonthus ; replacement name for excelsus Bernhauer ).

- DISTRIBUTION: Peru.

excelsus Bernhauer, 1941c: 287 [= 1951: 277] ( Philonthus ; [preoccupied]; Type locality:

Süd-Peru: Ocaña, 3100 m; Oberh. Palco, 3240 m; Oberh. Totora, 4000 m).

— Herman, 2001: 48 ( Philonthus ; synonym of supernus ).

suspectus Erichson, 1840: 465 ( Philonthus ; Type locality: Caracas Columbiae).

— Fauvel, 1891: 115 ( Philonthus ; Venezuela).

— Bernhauer and Schubert, 1914: 356 ( Philonthus ; catalog; Uruguay).

- DISTRIBUTION: Mexico, Guatemala, Venezuela, Brazil, Uruguay.

quadraticollis Sharp, 1885: 409 ( Philonthus ; Type locality: Mexico, Cordova , Jalapa;

Guatemala, San Geronimo; South America, Rio de Janeiro).

— Bernhauer and Schubert, 1914: 357 ( Philonthus ; synonym of suspectus ).

suspiciosus Cameron, 1932: 129 ( Philonthus ; Type locality: Burma: Ruby Mines ;

Tenasserim).

— Scheerpeltz, 1933: 1363 ( Philonthus ; catalog).

- DISTRIBUTION: Myanmar.

suspiritus Last, 1987: 28 ( Philonthus ; Type locality: Irian Jaya: New Guinea: S. Highlands,

Dimifa, SE of Mt. Giluwe, 2200 m).

- DISTRIBUTION: New Guinea.

sutteri Scheerpeltz, 1957c: 234 , 321 ( Philonthus ; Type locality: Sumba: Baing).

- DISTRIBUTION: Indonesia.

suturalis Marsham, 1802 , see: discoideus Gravenhorst, 1802 .

suturalis Stephens, 1832 , see: discoideus Gravenhorst, 1802 .

suturalis Nordmann, 1837 , see: transbaicalia Hochhuth, 1851 .

suturamonus Tottenham, 1939 [Note: Nomen nudum], see: sanguinolentus Gravenhorst,

1802.

suturorubineous Pajni and Kohli, 1977: 517 ( Philonthus ; Type locality: India: Chandigarh,

Labour Colony).

- DISTRIBUTION: India.

svanetiensis Coiffait, 1974 , see: frigidus Märkel and Kiesenwetter, 1848 .

swynnertoni Tottenham, 1953 , see: altivagans Fauvel, 1907 .

sylvicola Cameron, 1949a: 320 ( Philonthus ; Type locality: Côte d’Ivoire: Mt. Tonkoui ,

900–1200 m).

- DISTRIBUTION: Ivory Coast.

synonymicus Roubal, 1915 , see: Bisnius .

syriacus Gridelli, 1924 , see: Rabigus .

syrovatkai Roubal, 1913 [Note: Nomen nudum], see: Bisnius .

tabius Tottenham, 1955: 176 ( Philonthus ; Type locality: Cape Colony: Queenstown ).

- DISTRIBUTION: South Africa.

tachiniformis Say, 1830: 37 [= 1834: 450] ( Staphylinus ; Type locality: Mexico).

— Erichson, 1840: 930 ( Philonthus ; characters; Mexico).

— Sharp, 1885: 425 ( Philonthus ; notes; Mexico).

— Bernhauer and Schubert, 1914: 357 ( Philonthus ; catalog).

- DISTRIBUTION: Mexico.

tarsalis Smetana, 1963d: 412 ( Philonthus ; subgenus Onychophilonthus ; Type locality: Ost- Asien: UdSSR, Primorskij kraj, Küsten-Provinz, Naturschutzgebiet, “Kedrovaja pad”). — Coiffait, 1967a: 439 ( Philonthus ; subgenus Onychophilonthus ; characters; eastern Asia).

— Coiffait, 1974: 306 ( Philonthus ; subgenus Onychophilonthus ; characters; East Asia).

— Hayashi, 1994: 124 ( Philonthus ; subgenus Onychophilonthus ; characters; Japan).

- DISTRIBUTION: Russia, Japan.

taterae Scheerpeltz, 1951b: 27 ( Philonthus ; Type locality: South Africa: Vredefort ,

Orange Free State; Zuurbekom, Transvaal, Roodepoort district).

— Tottenham, 1962: 217, 220 ( Philonthus ; characters).

- DISTRIBUTION: South Africa.

taunggyiensis Scheerpeltz, 1965: 218 ( Philonthus ; Type locality: S. Shan-States, Burma,

1500 m, Taunggyi).

- DISTRIBUTION: Myanmar.

tauricus Nordmann, 1837 , see: Quedius .

tectipennis Coiffait, 1967 [Note: Nomen nudum], see: velatipennis Solsky, 1870 .

tectus Last, 1968 , see: Hesperus .

temperei Coiffait, 1987 , see: politus Linné, 1758 .

temporalis Mulsant and Rey, 1853: 61 [= 1853a: 74] ( Philonthus ; Type locality: Suisse).

— Fairmaire and Laboulbène, 1856: 516 ( Philonthus ; characters; France).

— Kraatz, 1855: 23 ( Xantholinus ; notes).

— Kraatz, 1857d: 573, note ( Philonthus ; characters).

— Rye, 1864a: 55 ( Philonthus ; characters; notes; Britain).

— Fauvel, 1874: 218 [= 1874a: 444] ( Philonthus ; characters; France; Germany; Switzerland; Tirol; Italy).

— Redtenbacher, 1874: 216 ( Philonthus ; characters).

— Mulsant and Rey, 1876: 379 [= 1877: 235] ( Philonthus ; characters; France).

— Ganglbauer, 1895: 445 ( Philonthus ; characters; Alps; Jura; Rheinland).

— Reitter, 1909: 126 ( Philonthus ; characters; Germany).

— Bernhauer and Schubert, 1914: 357 ( Philonthus ; catalog).

— Portevin, 1929: 359 ( Philonthus ; characters; France).

— Scheerpeltz, 1933: 1363 ( Philonthus ; catalog).

— Wüsthoff, 1934: 60 ( Philonthus ; illustration of aedeagus).

— Wüsthoff, 1937a: 347 ( Philonthus ; illustration of aedeagus).

— Smetana, 1958: 168 ( Philonthus ; subgenus Philonthus ; characters; ecology; Czechoslovakia). — Smetana, 1959: 148 ( Philonthus ; characters).

— Lohse, 1964: 178 ( Philonthus ; subgenus Philonthus ; characters; central Europe).

— Horion, 1965: 138 ( Philonthus ; Holland; Switzerland; France; Italy; Bosnia; Romania; Poland; Austria; Germany).

— Coiffait, 1967a: 395 ( Philonthus ; characters; Europe).

— Coiffait, 1974: 228 ( Philonthus ; characters; Europe).

— Tóth, 1984: 51 ( Philonthus ; characters; Hungary).

— Drugmand, 1987a: 311 ( Philonthus ; Belgium).

— Lucht, 1987: 105 ( Philonthus ; checklist; Germany; Austria; Switzerland; France; Benelux).

— Pilon and Zanetti, 1991: 63 ( Philonthus ; Italy).

— Smetana, 1993b: 48 ( Philonthus ; subgenus Philonthus ; Czechoslovakia?).

— Ciceroni and Zanetti, 1995: 29 ( Philonthus ; Italy).

— Schillhammer, 2000: 137 ( Philonthus ; characters; West Palaearctic).

- DISTRIBUTION: Europe.

punctiventris Kraatz, 1857d: 578 ( Philonthus ; [preoccupied]; Type locality: Baiern).

— Redtenbacher, 1857: 985 ( Philonthus ; characters; Austria).

— Fauvel, 1869: 489 ( Philonthus ; synonym of temporalis ).

— Rye, 1870a: 82 ( Philonthus ; notes).

— Fauvel, 1874: 218 [= 1874a: 444] ( Philonthus ; synonym of temporalis ).

— Redtenbacher, 1874: 216 ( Philonthus ; characters).

— Ganglbauer, 1895: 445 ( Philonthus ; synonym of temporalis ).

— Bernhauer and Schubert, 1914: 357 ( Philonthus ; catalog).

— Scheerpeltz, 1933: 1363 ( Philonthus ; synonym of temporalis ).

— Smetana, 1958: 168 ( Philonthus ; synonym of temporalis ).

— Coiffait, 1974: 228 ( Philonthus ; synonym of temporalis ).

— Schillhammer, 2000: 137 ( Philonthus ; synonym of temporalis ).

temporalis Coiffait, 1977 , see: aeolus Herman, 2001 .

temporalisoides Drugmand, 1988 , see: addendus Sharp, 1867 .

temporasinus Schubert, 1911 , see: morio Boheman, 1848 .

temporatus Last, 1968 , see: Hesperus .

tenebrosus Boheman, 1858: 28 ( Philonthus ; Type locality: Buenos Ayres).

— Lynch, 1884: 149 ( Philonthus ; characters; Argentina).

— Bernhauer and Schubert, 1914: 357 ( Philonthus ; catalog).

— Scheerpeltz, 1933: 1363 ( Philonthus ; catalog).

- DISTRIBUTION: Argentina.

tenellicornis Scheerpeltz, 1978: 220 ( Philonthus ; subgenus Glaphyronthus ; Type locality: Turisca, Timor).

- DISTRIBUTION: Indonesia.

tenellus Wollaston, 1864 , see: Neobisnius .

tenuemicans Scheerpeltz, 1965: 207 ( Philonthus ; Type locality: N.E. Burma, Kambaiti ,

2000 m).

- DISTRIBUTION: Myanmar.

tenuepunctatus Scheerpeltz, 1965: 220 ( Philonthus ; Type locality: N.E. Burma, Kambaiti , 2000 m).

- DISTRIBUTION: Myanmar.

tenuicornis Mulsant and Rey, 1853: 58 [= 1853a: 71] ( Philonthus ; Type locality: Lyonnais , Bourgogne).

— Gyllenhal, 1810: 319 ( Staphylinus ; [Note: Misidentification: Bernhauer and Schubert, 1914: 331 cited carbonarius Gyllenhal as a valid species; Gyllenhal did not describe the species as new, he attributed it to Gravenhorst]; characters; Sweden).

— Fairmaire and Laboulbène, 1856: 516 ( Philonthus ; characters; France).

— Kraatz, 1855: 23 ( Xantholinus ; notes; France).

— Kraatz, 1857d: 574, note ( Philonthus ; characters).

— Rye, 1863: 82 (Philontus; [Note: Misidentification: Bernhauer and Schubert, 1914: 339 cited punctiventris Janson as a synonym of carbonarius Gyllenhal ; Janson did not describe the species that was cited in Rye’s article. Janson reported the species from Britain based on one specimen; Rye reported Janson’s finding as punctiventris Kraatz ]; notes; characters; Britain).

— Fauvel, 1874: 217 [= 1874a: 443] ( Philonthus ; synonym of carbonarius ).

— Redtenbacher, 1874: 216 ( Philonthus ; characters).

— Seidlitz, 1875: 273 ( Philonthus ; cited as carbonarius Gyllenhal ; characters; Baltic region).

— Mulsant and Rey, 1876: 381 [= 1877: 237] ( Philonthus ; characters; larval characters; France).

— J. Sahlberg, 1876: 14 ( Philonthus ; cited as carbonarius Gyllenhal ; Finland).

— Fowler, 1888: 263 ( Philonthus ; cited as carbonarius Gyllenhal ; characters; Britain).

— Ganglbauer, 1895: 445 ( Philonthus ; cited as carbonarius Gyllenhal and with tenuicornis as synonym; characters; Europe).

— Reitter, 1909: 126 ( Philonthus ; cited as carbonarius Gyllenhal ; characters; Germany).

— Bernhauer and Schubert, 1914: 331 ( Philonthus ; synonym of carbonarius Gyllenhal ; [Note: The carbonarius of Gyllenhal is an unavailable, misidentified species]).

— Johansen, 1914: 395 ( Philonthus ; [Note: Gyllenhal cited as author]; characters; Denmark).

— Cameron, 1932: 81 ( Philonthus ; synonym of carbonarius ).

— Scheerpeltz, 1933: 1335 ( Philonthus ; synonym of carbonarius ).

— Tottenham, 1939d: 231 ( Philonthus ; cited with carbonarius Gyllenhal as a junior synonym; [Note: Gyllenhal did not describe carbonarius ]).

— Smetana, 1958: 166 ( Philonthus ; subgenus Philonthus ; cited with carbonarius Gyllenhal as

the senior name and tenuicornis as a junior synonym; [Note: carbonarius was described by Gravenhorst; the carbonarius Gyllenhal referred to by Gyllenhal has been associated with tenuicornis ]; characters; ecology; Czechoslovakia).

— Coiffait, 1967a: 395 ( Philonthus ; characters; Europe).

— Jarrige, 1971: 495 ( Philonthus ; Iran).

— Coiffait, 1974: 228 ( Philonthus ; characters; Europe; Scandinavia; Britain).

— Pace and Zanetti, 1977: 289 ( Philonthus ; Italy).

— Pope, 1977: 29 ( Philonthus ; Britain).

— Raichev, 1979: 82 ( Philonthus ; Bulgaria).

— Hinton, 1981: 661 ( Philonthus ; characters of egg).

— Frank, 1982: 43 ( Philonthus ; list of parasites).

— Lohse and Lucht, 1989: 165 ( Philonthus ; note).

— Pilon and Zanetti, 1991: 62 ( Philonthus ; Italy).

— Welch, 1993: 228 ( Philonthus ; ovariole number and ovary structure).

— Smetana, 1993b: 48 ( Philonthus ; subgenus Philonthus ; Czech Republic; Slovakia).

— Ciceroni and Zanetti, 1995: 29 ( Philonthus ; Italy).

— Smetana, 1995e: 123 ( Philonthus ; characters; introduced species; British Columbia).

— Luff, 1996: 30 ( Philonthus ; collecting notes; Isle of Man).

— Schillhammer, 2000: 130 ( Philonthus ; characters; West Palaearctic region as far east as Caucasus and Iran; introduced in Nearctic).

- DISTRIBUTION: Europe, Iran, Canada.

tenuimarginatus Scheerpeltz, 1960c: 112 ( Philonthus ; Type locality: Bolivien: Cochabamba, 2600 m).

- DISTRIBUTION: Bolivia.

tenuissimus Tottenham, 1949 , see: Gabronthus .

tergomaculatus Lecoq, 1997: 282 ( Philonthus ; Type locality: Madagascar Est , Ambodivoangy).

- DISTRIBUTION: Madagascar.

terminalis LeConte, 1863 , see: Neobisnius .

terminipennis Tottenham, 1939d: 233 ( Philonthus ; replacement name for apicipennis ).

- DISTRIBUTION: India, Nepal.

apicipennis Cameron, 1932: 117 ( Philonthus ; [preoccupied]; Type locality: Dehra Dun. Siwaliks : Mussoorie. Haldwani).

— Scheerpeltz, 1933: 1332 ( Philonthus ; catalog).

— Tottenham, 1939d: 233 ( Philonthus ; synonym of terminipennis ).

— Coiffait, 1984e: 374 ( Philonthus ; cited as valid species; Nepal).

testaceipennis Erichson, 1840: 435 ( Philonthus ; Type locality: Mexico).

— Solsky, 1868a: 127 ( Philonthus ; Mexico).

— Sharp, 1885: 400 ( Philonthus ; characters; Mexico).

— Bernhauer and Schubert, 1914: 357 ( Philonthus ; catalog).

- DISTRIBUTION: Mexico.

testaceipennis Roubal, 1948 [Note: Nomen nudum], see: Bisnius .

testaceocinctus Bernhauer, 1915d: 140 ( Philonthus ; Type locality: Deutsch-Ostafrika: Unjanjembe ).

— Scheerpeltz, 1933: 1364 ( Philonthus ; catalog).

— Tottenham, 1962: 139, 215, 217, 218 ( Philonthus ; characters; Uganda).

- DISTRIBUTION: Tanzania, Uganda.

testudo Smetana, 1995e: 263 ( Philonthus ; Type locality: Georgia: Tift Co.: Tifton. Also

cited from Florida, Mississippi).

- DISTRIBUTION: USA.