Graellsia Boissier (1841: 379)

Esmailbegi, Shokouh, Lysak, Martin A., Rahiminejad, Mohammad Reza, Mirtadzadini, Mansour, Mummenhoff, Klaus & Al-Shehbaz, Ihsan A., 2017, A taxonomic Revision of the genus Graellsia (Brassicaceae, tribe Thlaspideae), Phytotaxa 313 (1), pp. 105-116 : 106

Graellsia Boissier (1841: 379) . Type species: G. saxifragifolia (DC.) Boiss.

Syn.: Physalidium Fenzl in Tchihatcheff (1860: 327). Type species: P. stylosum (Boiss. & Hoh.) Fenzl

Description:—Herbs perennial, with woody caudex covered with persistent petiole bases of previous seasons. Trichomes absent. Stems erect to ascending, simple or branched distally, leafy. Basal leaves long petiolate, rosulate, simple, entire, obtusely toothed, or palmately lobed; cauline leaves petiolate, not auriculate at base, entire, dentate, or palmately lobed. Racemes many flowered, ebracteate, corymbose, elongated in fruit; rachis straight; fruiting pedicels erect to divaricate, persistent. Sepals ovate, free, deciduous, spreading, equal, base of lateral pair not saccate; petals white, erect at base with flaring blade, ascending, or spreading, longer than sepals; blade obovate to suborbicular, apex rounded; claw short, slightly differentiated from blade, shorter than sepals, glabrous, unappendaged, entire; stamens 6, slightly exserted, erect, tetradynamous; filaments wingless, unappendaged, glabrous, free; anthers ovate to oblong, not apiculate; nectar glands confluent, subtending bases of all stamens; median nectaries present; lateral nectaries annular; ovules 4−16 per ovary; placentation parietal. Fruit dehiscent or indehiscent, capsular silicles or rarely siliques, oblong, obovate, or elliptic, strongly latiseptate or angustiseptate, inflated or not, unsegmented; valves papery, obscurely veined or with a prominent midvien, glabrous, not keeled, smooth, wingless, unappendaged; gynophore absent; replum rounded, visible; septum complete, perforated, reduced to a rim, or lacking, membranous, veinless; style to 2 mm long, slender, persistent; stigma capitate, entire or slightly 2-lobed, unappendaged. Seeds uniseriate or biseriate, wingless or rarely distally winged, oblong, slightly flattened; seed coat smooth, not mucilaginous when wetted; cotyledons accumbent.

Nine species: Afghanistan, Iran, Iraq, Pakistan, Tajikistan, Turkey, and Turkmenistan.