Eudigraphis Silvestri, 1948

Karasawa, Shigenori, Kawano, Keisuke, Fukaya, Shin-ichi & Tsurusaki, Nobuo, 2020, Upgrading of Three Subspecies of Eudigraphis takakuwai to the Species Rank (Diplopoda: Penicillata: Polyxenida: Polyxenidae), Species Diversity 25, pp. 89-102 : 93

Genus Eudigraphis Silvestri, 1948

Eudigraphis Silvestri, 1948: 217 [type species: Eudigraphis ja- ponica Silvestri, 1948 (= Eudigraphis takakuwai )]; Nguy- en Duy-Jacquemin and Geoffroy 2003: 299; Short and Vahtera 2017: 2456.

Diagnosis. Head with 8 ocelli each side. Body with 10 segments, 9 pleural projections, a telson, and 13 pairs of legs in adults. Differs from Monographis Attems, 1907 , in 6th antennal segments having just 3 bacilliform sensilla, respectively, and by having caudal trichomes with more numerous (range: 2–6) barbed hooks arranged in a line. Differs from Unixenus in the presence of a single usually continuous row of trichomes along the posterior edge of the tergites.

Included species. Six species: E. takakuwai , E. nigricans , E. kinutensis , E. sinensis Ishii and Liang, 1990 , E. taiwanensis Ishii, 1990 , and E. xishuangbanna Ishii and Yin, 2000 .

Distribution. Japan, Taiwan, and China.

Remarks. Miyosi (1959) synonymized Eudigraphis with Monographis by expanding definition of the latter. However, Eudigraphis has been treated as a distinct genus different from Monographis since the revision of Monographis by Nguyen Duy-Jacquemin and Condé (1967) (also see Shinohara and Ishii 1984; Ishii 1988).

Japanese species of Eudigraphis seem to show close morphological similarity to Unixenus Jones, 1944 ( Short and Huynh 2013; Huynh and Veenstra 2018a), especially in position and the number of bacilliform sensilla on distal segments of the antennae (cf., fig. 6 in Huynh and Veenestra 2018a) and in morphology of the gnathochilarium (cf., fig. 4F in Huynh and Veenestra 2018a). However, Japanese Eudigraphis differs from Unixenus in morphology of biarticuated setae on the second legs and show similarity to Monographis in this character (cf., fig. 12 in Huynh and Veenestra 2018a). Separation of Eudigraphis from Unixenus seems to be supported by a molecular phylogeny provided by Short and Vahtera (2017). However, further detailed analyses seem to be needed.